The Chinese Hwamei or Melodious Laughingthrush (Leucodioptron canorum, formerly Garrulax canorus) is a passerine

Passerine

A passerine is a bird of the order Passeriformes, which includes more than half of all bird species. Sometimes known as perching birds or, less accurately, as songbirds, the passerines form one of the most diverse terrestrial vertebrate orders: with over 5,000 identified species, it has roughly...

 bird

Bird

Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 of eastern Asia

Asia

Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 in the Old World babbler

Old World babbler

The Old World babblers or timaliids are a large family of mostly Old World passerine birds. They are rather diverse in size and coloration, but are characterised by soft fluffy plumage. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian Subcontinent...

 family, Timaliidae. The name "Hwamei" comes from the Chinese

Chinese language

The Chinese language is a language or language family consisting of varieties which are mutually intelligible to varying degrees. Originally the indigenous languages spoken by the Han Chinese in China, it forms one of the branches of Sino-Tibetan family of languages...

 画眉 (huà-méi) and means "painted eyebrow" referring to the distinctive marking around the bird's eyes. The species is a popular cagebird because of its attractive song.

Taxonomy

It has two subspecies

Subspecies

Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

: L. c. canorum, native to mainland Asia, and L. c. owstoni of Hainan

Hainan

Hainan is the smallest province of the People's Republic of China . Although the province comprises some two hundred islands scattered among three archipelagos off the southern coast, of its land mass is Hainan Island , from which the province takes its name...

 Island. The Taiwan Hwamei

Taiwan Hwamei

The Taiwan Hwamei is a passerine bird in the Old World babbler family, Timaliidae. It is endemic to the island of Taiwan. It was formerly regarded as a subspecies of the Chinese Hwamei but has recently been split as a separate species...

 (Leucodioptron taewanum) was formerly considered to be a subspecies of the Chinese Hwamei but has recently been split as a separate species. Based on a study of the mitochondrial

Mitochondrial DNA

Mitochondrial DNA is the DNA located in organelles called mitochondria, structures within eukaryotic cells that convert the chemical energy from food into a form that cells can use, adenosine triphosphate...

 cytochrome b

Cytochrome b

Cytochrome b/b6 is the main subunit of transmembrane cytochrome bc1 and b6f complexes. In addition, it commonly refers to a region of mtDNA used for population genetics and phylogenetics.- Function :...

 gene

Gene

A gene is a molecular unit of heredity of a living organism. It is a name given to some stretches of DNA and RNA that code for a type of protein or for an RNA chain that has a function in the organism. Living beings depend on genes, as they specify all proteins and functional RNA chains...

, Li et al. (2006) suggested that the two species diverged about 1.5 million years ago with the two Chinese Hwamei subspecies diverging about 600,000 years ago. The two were formerly placed in the genus

Genus

In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Garrulax with the other laughingthrush

Laughingthrush

The Laughingthrushes are the genus Garrulax of the large Old World babbler family of passerine birds. They occur in tropical Asia, with the greatest number of species occurring in the Himalaya and southern China....

es but have recently been moved to a new genus Leucodioptron.

The Chinese Hwamei was introduced to Taiwan

Taiwan

Taiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...

 in large numbers in the 1980s and hybridization with the native Taiwan Hwamei is occurring which may threaten the genetic uniqueness of the latter form.

Description

It is 21 to 25 cm long with broad, rounded wings and a fan-shaped tail. The plumage

Plumage

Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

 is mostly reddish-brown with dark steaks on the crown, back and throat. There is a white ring around the eye which extends backwards as a white stripe. The bill

Beak

The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 and feet are yellowish. Birds on Hainan

Hainan

Hainan is the smallest province of the People's Republic of China . Although the province comprises some two hundred islands scattered among three archipelagos off the southern coast, of its land mass is Hainan Island , from which the province takes its name...

 Island (L. c. owstoni) are paler below and more olive-coloured above. The Taiwan Hwamei is greyer and more streaked and lacks the white markings on the head.

The song

Bird song

Bird vocalization includes both bird calls and bird songs. In non-technical use, bird songs are the bird sounds that are melodious to the human ear. In ornithology and birding, songs are distinguished by function from calls.-Definition:The distinction between songs and calls is based upon...

 is a loud, clear, varied whistling with regular repetition and imitations of other birds. The call is a rasping whistle or rattle.

Distribution and habitat

The nominate subspecies L. c. canorum occurs across south-eastern and central China

China

Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

 and in northern and central Vietnam

Vietnam

Vietnam – sometimes spelled Viet Nam , officially the Socialist Republic of Vietnam – is the easternmost country on the Indochina Peninsula in Southeast Asia. It is bordered by China to the north, Laos to the northwest, Cambodia to the southwest, and the South China Sea –...

 and Laos

Laos

Laos Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ Sathalanalat Paxathipatai Paxaxon Lao, officially the Lao People's Democratic Republic, is a landlocked country in Southeast Asia, bordered by Burma and China to the northwest, Vietnam to the east, Cambodia to the south and Thailand to the west...

. The race L. c. owstoni is found on Hainan.

L. c. canorum has been introduced

Introduced species

An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...

 to Taiwan, Singapore

Singapore

Singapore , officially the Republic of Singapore, is a Southeast Asian city-state off the southern tip of the Malay Peninsula, north of the equator. An island country made up of 63 islands, it is separated from Malaysia by the Straits of Johor to its north and from Indonesia's Riau Islands by the...

, Japan

Japan

Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

  and Hawaii

Hawaii

Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

. In the Hawaiian Islands it was introduced in the early 20th century and now occurs in both native forest and man-made habitats. It is common on Kauai

Kauai

Kauai or Kauai, known as Tauai in the ancient Kaua'i dialect, is geologically the oldest of the main Hawaiian Islands. With an area of , it is the fourth largest of the main islands in the Hawaiian archipelago, and the 21st largest island in the United States. Known also as the "Garden Isle",...

, Maui

Maui

The island of Maui is the second-largest of the Hawaiian Islands at and is the 17th largest island in the United States. Maui is part of the state of Hawaii and is the largest of Maui County's four islands, bigger than Lānai, Kahoolawe, and Molokai. In 2010, Maui had a population of 144,444,...

 and Hawaii Island but less so on Oahu

Oahu

Oahu or Oahu , known as "The Gathering Place", is the third largest of the Hawaiian Islands and most populous of the islands in the U.S. state of Hawaii. The state capital Honolulu is located on the southeast coast...

 and Molokai

Molokai

Molokai or Molokai is an island in the Hawaiian archipelago. It is 38 by 10 miles in size with a land area of , making it the fifth largest of the main Hawaiian Islands and the 27th largest island in the United States. It lies east of Oahu across the 25-mile wide Kaiwi Channel and north of...

.

The bird inhabits scrubland, open woodland

Woodland

Ecologically, a woodland is a low-density forest forming open habitats with plenty of sunlight and limited shade. Woodlands may support an understory of shrubs and herbaceous plants including grasses. Woodland may form a transition to shrubland under drier conditions or during early stages of...

, secondary forest

Secondary forest

A secondary forest is a forest or woodland area which has re-grown after a major disturbance such as fire, insect infestation, timber harvest or windthrow, until a long enough period has passed so that the effects of the disturbance are no longer evident...

, parks and gardens up to 1800 metres above sea level. It is common in much of its range and is not considered a threatened species

Threatened species

Threatened species are any speciesg animals, plants, fungi, etc.) which are vulnerable to endangerment in the near future.The World Conservation Union is the foremost authority on threatened species, and treats threatened species not as a single category, but as a group of three categories,...

.

Behaviour

It is a skulking bird which is often very difficult to see. It typically feeds on the ground among leaf litter, foraging for insects and fruit. It usually occurs in pairs or in small groups.

The breeding season lasts from May to July. A large cup-shaped nest

Bird nest

A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American Robin or Eurasian Blackbird, or the elaborately woven hanging nest of the...

 is built up to two metres above the ground in a tree or bush or amongst undergrowth. Two to five blue or blue-green eggs

Egg (biology)

An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

are laid.

External links

• Oriental Bird Images - Hwamei
• Hwamei videos on the Internet Bird Collection