Robert T. Paine, PhD

PHD

PHD may refer to:*Ph.D., a doctorate of philosophy*Ph.D. , a 1980s British group*PHD finger, a protein sequence*PHD Mountain Software, an outdoor clothing and equipment company*PhD Docbook renderer, an XML renderer...

, is a zoologist and professor emeritus of the University of Washington

University of Washington

University of Washington is a public research university, founded in 1861 in Seattle, Washington, United States. The UW is the largest university in the Northwest and the oldest public university on the West Coast. The university has three campuses, with its largest campus in the University...

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who coined the keystone species

Keystone species

A keystone species is a species that has a disproportionately large effect on its environment relative to its abundance. Such species play a critical role in maintaining the structure of an ecological community, affecting many other organisms in an ecosystem and helping to determine the types and...

concept in order to explain the relationship between Pisaster ochraceus, a species of starfish, and Mytilus californianus, a species of mussel

Mussel

The common name mussel is used for members of several families of clams or bivalvia mollusca, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which are often more or less rounded or oval.The...

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In his classic 1966 paper, Dr. Robert Paine described such a system in Makah Bay

Neah Bay, Washington

Neah Bay is a census-designated place on the Makah Indian reservation in Clallam County, Washington, United States. The population was 794 at the 2000 census.-Geography:Neah Bay is located at ....

 in Washington State.
This led to his 1969 paper where he proposed the keystone species concept.