1999 in paleontology
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Insects

Name Novelty Status Authors Age Unit Location Notes Images

Neoephemera antiqua
Neoephemera antiqua
Neoephemera antiqua is an extinct species of mayfly in the family Neoephemeridae that is known from early Eocene, Ypresian stage, lake deposits near the small community of Republic in Ferry County, Washington, USA.-History and classification:...


sp. nov

valid
  • Sinitchenkova

Ypresian
Ypresian
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between and , is preceded by the Thanetian age and is followed by the Eocene Lutetian age....


Klondike Mountain Formation
Klondike Mountain Formation
The Klondike Mountain Formation is an early Eocene geologic formation that records the sedimentation in the southern most of a string of highland subtropical/Temperate lakes in Washington state and British Columbia. The formation is best known for exceptionally well preserved plant and insect fossils...


 United States

Oldest neoephemerid mayfly species described

Pachycondyla rebekkae
Ypresiomyrma
Ypresiomyrma is an extinct genus of bulldog ants in the subfamily Myrmeciinae which contains three described species. One species is known from the Isle of Fur in Denmark and two are from British Columbia, Canada.-History and classification:...


sp. nov

valid
  • Rust & Andersen

Ypresian
Ypresian
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between and , is preceded by the Thanetian age and is followed by the Eocene Lutetian age....


Fur Formation
Fur Formation
Essentially the Fur Formation is defined as a clayey diatomite with a lage number of volcanic ash layers.The Fur Formation is a marine geological formation of Ypresian age which crops out in the Limfjord region of Denmark from Silstrup via Mors and Fur to Ertebølle, and can be seen in many cliffs...


 Denmark

moved by Archibald, Cover, & Moreau to Ypresiomyrma
Ypresiomyrma
Ypresiomyrma is an extinct genus of bulldog ants in the subfamily Myrmeciinae which contains three described species. One species is known from the Isle of Fur in Denmark and two are from British Columbia, Canada.-History and classification:...



Newly named taxa

Name Status Authors Age Unit Location Notes Images
Nardovelifer
Nardovelifer
Nardovelifer altipinnis is the oldest known lamprid fish. It dates from the Campanian epoch of Nardò, Italy....

Valid
  • C. Sorbini
  • L. Sorbini
Campanian Nardo  Italy

Newly named dinosaurs

Data courtesy of George Olshevsky
George Olshevsky
George Olshevsky is a freelance editor, writer, publisher, amateur paleontologist, and mathematician living in San Diego, California.Olshevsky maintains the comprehensive online Dinosaur Genera List...

's dinosaur genera list and Dr. Jeremy Montague's dinosaur genus database.
Name Status Authors Notes Images
Achillobator
Achillobator
Achillobator is a genus of dromaeosaurid theropod dinosaur from the late Cretaceous Period of what is now Mongolia, about 90 million years ago. It was probably an active bipedal predator, hunting with the large sickle-shaped claw on the second toe of each hind foot...

Valid taxon
  • Altangerel Perle
    Altangerel Perle
    Altangerel Perle is a Mongolian palaeontologist.He is employed at the National University of Mongolia. He has described species such as Goyocephale lattimorei, Achillobator giganticus and Erlikosaurus andrewsi. He has been honored by Polish palaeontologist Halszka Osmólska, who named the species...

  • Mark Norell
  • Clark

  • A large dromaeosaurid.

    Agustinia
    Agustinia
    Agustinia is a genus of sauropod dinosaur from the Early Cretaceous Period of South America. Like all known sauropods, it was quadrupedal and herbivorous. Although some sauropods are known to have body armor, Agustinias armor was unique even among sauropods...

    Valid taxon
    • Jose Bonaparte
      José Bonaparte
      José Fernando Bonaparte, Ph.D. , is an Argentine paleontologist who discovered a plethora of South American dinosaurs and mentored a new generation of Argentine paleontologists like Rodolfo Coria...


    A 50 foot (15 meters) long sauropod with bony spikes running down its back.
    "Airakoraptor
    Airakoraptor
    "Airakoraptor" was a genus of theropod dinosaur from the Late Cretaceous Period. It lived in what is now Mongolia and was first described by Perle, Norell & Clark, in 1999. It is considered a nomen nudum because it has not been fully described...

    "
    Nomen nudum
    Nomen nudum
    The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...

    • Altangerel Perle
      Altangerel Perle
      Altangerel Perle is a Mongolian palaeontologist.He is employed at the National University of Mongolia. He has described species such as Goyocephale lattimorei, Achillobator giganticus and Erlikosaurus andrewsi. He has been honored by Polish palaeontologist Halszka Osmólska, who named the species...

    • Mark Norell
  • Clark

  • A possible dromaeosaurid that has yet to be fully described.
    Animantarx
    Animantarx
    Animantarx is a genus of nodosaurid ankylosaurian dinosaur from the Upper Cretaceous of western North America. Like other nodosaurs, it would have been a slow-moving quadrupedal herbivore covered in heavy armor scutes, but without a tail club...

    Valid taxon
    • Kenneth Carpenter
      Kenneth Carpenter
      Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life...

    • James Kirkland
  • Burge
  • Bird

  • A 10 foot (3 meters) long nodosaur whose name means "living citadel."
    "Archaeoraptor
    Archaeoraptor
    "Archaeoraptor" is the generic name informally assigned in 1999 to a fossil from China in an article published in National Geographic magazine. The magazine claimed that the fossil was a "missing link" between birds and terrestrial theropod dinosaurs. Even prior to this publication there had been...

    "

    Hoax

    Stephen Czerkas vide:
    C. P. Sloan vide:
    • Olson

    Fraudulent chimera of Yanornis
    Yanornis
    Yanornis is an extinct genus of omnivorous Early Cretaceous birds, thought to be closely related to the common ancestor of all modern birds. One species, Yanornis martini,...

    and Microraptor
    Microraptor
    Microraptor is a genus of small, four-winged dromaeosaurid dinosaurs. Numerous well-preserved fossil specimens have been recovered from Liaoning, China...

    .
    Atlasaurus
    Atlasaurus
    Atlasaurus was a moderately large genus of sauropod dinosaur from Middle Jurassic beds in North Africa.-Discovery and species:Atlasaurus was discovered by Monbaron, Russell & Taquet in 1999...

    Valid taxon
    • Monbaron
    • Dale A. Russell
  • Philippe Taquet
    Philippe Taquet
    Philippe Taquet is a French paleontologist who specializes in dinosaur systematics of finds primarily in northern Africa.He is a member of the French Academy of Sciences since November 30, 2004...


  • A somewhat primitive sauropod about 15 m (50 ft) long.
    Beipiaosaurus
    Beipiaosaurus
    Beipiaosaurus is a genus of therizinosauroid theropod dinosaur from the Early Cretaceous of China.The exact classification of therizinosaurs had in the past been hotly debated, since their prosauropod-like teeth and body structure indicate that they were generally herbivorous, unlike typical...

    Valid taxon
    • Xu Xing
    • Tang Z.
  • Wang X

  • A therizinosaur 2.2 meters (7.3 ft) in length. It is one of the largest known feathered dinosaurs.
    "Bilbeyhallorum" Nomen nudum
    Nomen nudum
    The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...

    • Burge
    • Bird
  • McClelland
  • Cicconetti

  • Formally named Cedarpelta
    Cedarpelta
    Cedarpelta is an extinct genus of basal ankylosaurid ankylosaur, based on material recovered from the Lower Cretaceous of North America. The skull lacks extensive cranial ornamentation, a trait which has been interpreted as plesiomorphic for ankylosaurs....

    in 2001
    2001 in paleontology
    -Newly named insects:-Newly named bony fishes:-Newly named amphibians:-Newly named turtles:-Ichthyosaurs:-Newly named basal lepidosauromorphs:-Newly named plesiosaurs:-Newly named sphenodonts:-Newly named basal archosauromorphs:...

    .
    Cedarosaurus
    Cedarosaurus
    Cedarosaurus was a nasal-crested macronarian dinosaur genus from the Early Cretaceous Period . It was a sauropod which lived in what is now Utah...

    Valid taxon
    • Tidwell
    • Kenneth Carpenter
      Kenneth Carpenter
      Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life...

  • Brooks

  • A brachiosaurid.
    Chaoyangsaurus
    Chaoyangsaurus
    Chaoyangsaurus was a marginocephalian dinosaur from the Late Jurassic of China...

    Valid taxon
    • Zhao X.
    • Cheng Z.
  • Xu Xing

  • An extremely primitive ceratopsian with many synonyomous nomena nuda in the scientific literature.
    Dinheirosaurus
    Dinheirosaurus
    Dinheirosaurus is a genus of diplodocid sauropod dinosaur. It has been found in the upper section of the Late Jurassic strata of the Camadas de Alcobaça Formation located in central-western Portugal at Porto Dinheiro, Lourinhã...

    Valid taxon
    • Jose Bonaparte
      José Bonaparte
      José Fernando Bonaparte, Ph.D. , is an Argentine paleontologist who discovered a plethora of South American dinosaurs and mentored a new generation of Argentine paleontologists like Rodolfo Coria...

  • Mateus

  • A diplodocid.
    "Eucoelophysis
    Eucoelophysis
    Eucoelophysis is a genus of dinosauriform from the Late Triassic period Chinle Formation of New Mexico...

    "

    Misidentification.
    • Sullivan
  • S. G. Lucas,

  • A misidentified non-dinosaurian dinosauriform.
    "Eugongbusaurus
    Eugongbusaurus
    "Eugongbusaurus" is the informal name given to a genus of dinosaur that lived about 160 to 155 million years ago, in the Late Jurassic. It was either a hypsilophodont, or a less-derived ornithischian...

    "
    Nomen nudum
    Nomen nudum
    The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...

    • Knoll

    A hypsilophodont that hasn't yet been formally named.
    Gondwanatitan
    Gondwanatitan
    Gondwanatitan was a titanosaurian sauropod dinosaur. Gondwanatitan was found in Brazil, at the time part of the southern supercontinent Gondwana , in the late Cretaceous Period . Like some other sauropods, Gondwanatitan was tall and ate tough shoots and leaves off of the tops of trees. G...

    Valid taxon
    • Kellner
  • Azevedo

  • A titanosaur.
    Guaibasaurus
    Guaibasaurus
    Guaibasaurus is an extinct genus of basal dinosaur known from the Triassic of Rio Grande do Sul, southern Brazil.-Discovery:Guaibasaurus was originally named on the basis of the holotype, MCN PV2355, a well-preserved partial postcranial skeleton and the paratype, MCN PV2356, an...

    Valid taxon
    • Jose Bonaparte
      José Bonaparte
      José Fernando Bonaparte, Ph.D. , is an Argentine paleontologist who discovered a plethora of South American dinosaurs and mentored a new generation of Argentine paleontologists like Rodolfo Coria...

    • Ferigolo
  • Ribeiro

  • A basal saurischian.
    Jobaria
    Jobaria
    Jobaria was a sauropod dinosaur discovered in the Sahara Desert in 1997. It was named after "Jobar", a creature of local legends, and is thought to have been about 18 metres long. It was found in the Tiourarén Formation, originally thought to represent the Hauterivian to Barremian stages of the...

    Valid taxon
    • Paul Sereno
      Paul Sereno
      Paul Callistus Sereno is an American paleontologist from the University of Chicago who discovered several new dinosaur species on several continents. He has conducted excavations at sites as varied as Inner Mongolia, Argentina, Morocco, and Niger...

    • Beck
    • Dutheil
    • Larsson
    • Lyon
    • Moussa
  • Sadleir
  • Sidor
    Christian Sidor
    Christian A. Sidor is an American biologist and paleontologist, Associate Professor of the Department of Biology, University of Washington, Seattle, WA, curator of vertebrate paleontology in the Burke Museum of Natural History and Culture, member of the editorial board of the Journal of Vertebrate...

  • Varricchio
  • G. P. Wilson
  • Jeffrey A. Wilson
    Jeffrey A. Wilson
    Jeffrey A. Wilson also known as "JAW "is a professor of geological sciences and assistant curator at the Museum of Paleontology at the University of Michigan....

    .

  • A sauropod named after the mythological "Jobar" and thought to have been about 18 metres long.
    Lessemsaurus
    Lessemsaurus
    Lessemsaurus is an extinct genus of sauropodomorph dinosaur named for the writer of popular science books Don Lessem. The type species, L. sauropoides, was formally described by José Fernando Bonaparte in 1999...

    Valid taxon
    • Jose Bonaparte
      José Bonaparte
      José Fernando Bonaparte, Ph.D. , is an Argentine paleontologist who discovered a plethora of South American dinosaurs and mentored a new generation of Argentine paleontologists like Rodolfo Coria...


    A primitive sauropodomorph in the family Melanorosauridae.
    Lirainosaurus
    Lirainosaurus
    Lirainosaurus is the name given to a genus of dinosaur from the Late Cretaceous. It was a titanosaurid sauropod which lived in what is now Spain. The type species, Lirainosaurus astibiae, was described by Sanz, Powell, Le Loeuff, Martinez, and Pereda-Suberbiola in 1999.-References:J. L. Sanz, J. E....

    Valid taxon
    • Sanz
    • Jaime Eduardo Powell
    • Le Loeuff
  • Martínez
  • Pereda-Suberbiola

  • A titanosaurid.
    Lurdusaurus
    Lurdusaurus
    Lurdusaurus is a genus of large ornithopod dinosaur which lived in the Aptian stage of the Early Cretaceous, sometime between 121 to 112 million years ago.-Discovery and naming:...

    Valid taxon
    • Philippe Taquet
      Philippe Taquet
      Philippe Taquet is a French paleontologist who specializes in dinosaur systematics of finds primarily in northern Africa.He is a member of the French Academy of Sciences since November 30, 2004...

  • Dale A. Russell
  • "Newtonsaurus
    Newtonsaurus
    "Newtonsaurus" is the informal name given to an as yet undescribed genus of dinosaur from the Late Triassic. It was probably a ceratosaur, which lived in what is now the United Kingdom. The type species is "N...

    "
    Nomen nudum
    Nomen nudum
    The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...


    Welles vide:
    • Welles
  • Pickering
  • Nigersaurus
    Nigersaurus
    Nigersaurus is a genus of diplodocoid sauropod dinosaur from the middle Cretaceous period, about 119 to 99 million years ago during the Aptian or Albian age. This dinosaur was described by Paul Sereno and colleagues in 1999...

    Valid taxon
  • Paul Sereno
    Paul Sereno
    Paul Callistus Sereno is an American paleontologist from the University of Chicago who discovered several new dinosaur species on several continents. He has conducted excavations at sites as varied as Inner Mongolia, Argentina, Morocco, and Niger...

  • Beck
  • Dutheil
  • Larsson
  • Lyon
  • Moussa
  • Sadleir
  • Sidor
  • Varricchio
  • G. P. Wilson
  • Jeffrey A. Wilson
    Jeffrey A. Wilson
    Jeffrey A. Wilson also known as "JAW "is a professor of geological sciences and assistant curator at the Museum of Paleontology at the University of Michigan....

    .
  • Nodocephalosaurus
    Nodocephalosaurus
    Nodocephalosaurus is a genus of herbivorous ankylosaurine ankylosaurid dinosaur from Upper Cretaceous deposits of San Juan Basin, New Mexico. The holotype was recovered from the Late Campanian De-na-zin Member of the Kirtland Formation and consists of an incomplete skull...

    Valid taxon
  • Sullivan
  • Proyandusaurus Valid taxon
  • Knoll
  • Qantassaurus
    Qantassaurus
    Qantassaurus is a genus of two-legged, plant-eating ornithischian dinosaur that lived in Australia about 115 million years ago, when the continent was still partly south of the Antarctic Circle...

    Valid taxon
  • Thomas H. Rich
  • Patricia Vickers-Rich.
  • Santanaraptor
    Santanaraptor
    Santanaraptor is a genus of theropod dinosaur that lived in South America during the Albian or Aptian stage of the Early Cretaceous, about 108 million years ago. Known from a partial juvenile specimen recovered with mineralized soft tissue, which was unearthed in 1996 from the Santana Formation of...

    Valid taxon
  • Kellner
  • Saturnalia
    Saturnalia (dinosaur)
    Saturnalia is an extinct genus of basal sauropodomorph dinosaur known from the Triassic of Rio Grande do Sul, southern Brazil.-Discovery:Saturnalia was originally named on the basis of three partial skeletons...

    Valid taxon
  • Langer
  • Abdala
  • Richter
  • Benton
  • Sinornithosaurus
    Sinornithosaurus
    Sinornithosaurus is a genus of feathered dromaeosaurid dinosaur from the early Cretaceous Period of the Yixian Formation in what is now China. It was the fifth non–avian feathered dinosaur genus discovered by 1999...

    Valid taxon
  • Xu Xing
  • Wang X.
  • Wu X.
  • Tangvayosaurus
    Tangvayosaurus
    Tangvayosaurus is a genus of sauropod dinosaur from the Aptian-Albian age Lower Cretaceous Grès Supérior Formation of Savannakhet, Laos...

    Valid taxon
  • Allain
  • Philippe Taquet
    Philippe Taquet
    Philippe Taquet is a French paleontologist who specializes in dinosaur systematics of finds primarily in northern Africa.He is a member of the French Academy of Sciences since November 30, 2004...

  • Battail, Dejax
  • Richir
  • Véran
  • Limon-Duparcmeur
  • Vacant
  • Mateus
  • Sayarath
  • Khenthavong
  • Phouyavong
  • Tehuelchesaurus
    Tehuelchesaurus
    Tehuelchesaurus is a genus of dinosaur. It is named in honor of the Tehuelche people native to the Argentinian province of Chubut, where it was first found....

    Valid taxon
  • Thomas H. Rich
  • Patricia Vickers-Rich
  • Gimenez
  • Cúneo
  • Puerta
  • Vacca Teyuwasu Kischlat

  • New taxa

    s="wikitable sortable" align="center" width="100%">
    Name Status Authors Notes

    Callawayasaurus
    Callawayasaurus
    Callawayasaurus is a genus of plesiosaur from the family Elasmosauridae. When the first Callawayasaurus fossil was first discovered by Samuel Paul Welles in 1962, he described it as Alzadasaurus colombiensis before it was moved into its current genus by Kenneth Carpenter in...


    Valid

    Carpenter

    Valid
    er="0" style= height:"100%" align="right" style="background:transparent;">

    New taxa

    s="wikitable sortable" align="center" width="100%">
    Name Status Authors Notes

    Dendrorhynchoides
    Dendrorhynchoides
    Dendrorhynchoides was a genus of anurognathid pterosaur from the Early Cretaceous Yixian Formation, near Beipiao, Chaoyang, Liaoning, China...


    Valid

    Ji S.-A.
    Ji Q.
    Padian

    Siroccopteryx

    Valid

    Mader
    Kellner

    Tendaguripterus
    Tendaguripterus
    Tendaguripterus was a genus of dsungaripteroid pterodactyloid pterosaur from the Kimmeridgian-Tithonian-age Upper Jurassic Middle Saurian Beds of Tendaguru, Mtwara Region, Tanzania....


    Valid

    Unwin
    Heinrich
    er="0" style= height:"100%" align="right" style="background:transparent;">

    Non-mammalian

    Name Status Authors Discovery year Age Unit Location Notes Images

    '

    Valid

    '

    Valid
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