Abronia macrocarpa is a rare species of flowering plant known by the common name large-fruited sand verbena. It is endemic to eastern Texas

Texas

Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

, where it is known from three counties. It inhabits harsh, open sand dunes

Dune

In physical geography, a dune is a hill of sand built by wind. Dunes occur in different forms and sizes, formed by interaction with the wind. Most kinds of dunes are longer on the windward side where the sand is pushed up the dune and have a shorter "slip face" in the lee of the wind...

 on savanna

Savanna

A savanna, or savannah, is a grassland ecosystem characterized by the trees being sufficiently small or widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to support an unbroken herbaceous layer consisting primarily of C4 grasses.Some...

s, growing in deep, poor soils. It was first collected in 1968 and described as a new species in 1972. It is a federally listed endangered species

Endangered species

An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

 of the United States.

This is a perennial herb with a hairy, glandular stem growing up to half a meter tall. The glandular oval leaf blades are up to 5 centimeters long by 3.5 wide and are borne on relatively long petioles

Petiole (botany)

In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...

. The inflorescence

Inflorescence

An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Strictly, it is the part of the shoot of seed plants where flowers are formed and which is accordingly modified...

 is a cluster of up to 75 magenta or light purple flowers each up to 3 centimeters long. The tubular, strongly fragrant flowers open at dusk and are pollinated

Pollination

Pollination is the process by which pollen is transferred in plants, thereby enabling fertilisation and sexual reproduction. Pollen grains transport the male gametes to where the female gamete are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself...

 by moth

Moth

A moth is an insect closely related to the butterfly, both being of the order Lepidoptera. Moths form the majority of this order; there are thought to be 150,000 to 250,000 different species of moth , with thousands of species yet to be described...

s. The winged fruit is up to 1.5 centimeters long. It is dispersed

Biological dispersal

Biological dispersal refers to species movement away from an existing population or away from the parent organism. Through simply moving from one habitat patch to another, the dispersal of an individual has consequences not only for individual fitness, but also for population dynamics, population...

 by wind.

Threats to this endangered species include habitat loss as its range is consumed for development and oil exploration

Oil well

An oil well is a general term for any boring through the earth's surface that is designed to find and acquire petroleum oil hydrocarbons. Usually some natural gas is produced along with the oil. A well that is designed to produce mainly or only gas may be termed a gas well.-History:The earliest...

. The habitat is also damaged by off-road vehicle

Off-road vehicle

An off-road vehicle is considered to be any type of vehicle which is capable of driving on and off paved or gravel surface. It is generally characterized by having large tires with deep, open treads, a flexible suspension, or even caterpillar tracks...

s, people on foot and on horseback

Equestrianism

Equestrianism more often known as riding, horseback riding or horse riding refers to the skill of riding, driving, or vaulting with horses...

, fire suppression activity, and the invasion of non-native species

Introduced species

An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...

 such as bermudagrass

Cynodon dactylon

Cynodon dactylon , also known as Dūrvā Grass, Bermuda Grass, Dubo, Dog's Tooth Grass, Bahama Grass, Devil's Grass, Couch Grass, Indian Doab, Arugampul, Grama, and Scutch Grass, is a grass native to north and east Africa, Asia and Australia and southern Europe...

 and weeping lovegrass (Eragrostis curvula).

The total remaining number is estimated at several thousand individual plants in nine populations.

External links

• USDA Plants Profile
• Kennedy, K. and J. Poole. (1992). Large-fruited Sand Verbena Recovery Plan. USFWS.