African Bush Elephant
Encyclopedia
The African Bush Elephant or African Savanna Elephant (Loxodonta africana) is the larger of the two species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of African elephant. Both it and the African Forest Elephant
African Forest Elephant
The African Forest Elephant is a forest dwelling elephant of the Congo Basin. Formerly considered either a synonym or a subspecies of the African Savanna Elephant , a 2010 study established that the two are distinct species...

 have usually been classified as a single species, known simply as the African Elephant. Some authorities still consider the currently available evidence insufficient for splitting the African Elephant into two species. It is also known as the Bush Elephant.

Description

The African Elephant is the largest living terrestrial animal, normally reaching 6 to 7.3 m (19.7 to 24 ft) in length and 3.5 to 4 m (11.5 to 13.1 ft) in height at the head, and weighing between 6000 to 9000 kg (13,227.7 to 19,841.6 lb).

The largest on record, shot in Angola
Angola
Angola, officially the Republic of Angola , is a country in south-central Africa bordered by Namibia on the south, the Democratic Republic of the Congo on the north, and Zambia on the east; its west coast is on the Atlantic Ocean with Luanda as its capital city...

 in 1955, was a bull weighing 12274 kg (27,059.5 lb) and standing 4.2 metres (13.8 ft) high, the body of which is now mounted in the rotunda of the National Museum of Natural History
National Museum of Natural History
The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. Admission is free and the museum is open 364 days a year....

 in Washington, D.C.
Washington, D.C.
Washington, D.C., formally the District of Columbia and commonly referred to as Washington, "the District", or simply D.C., is the capital of the United States. On July 16, 1790, the United States Congress approved the creation of a permanent national capital as permitted by the U.S. Constitution....

. (The museum's website states that the specimen weighs only 8 tons.) The Bush Elephant normally moves at a rate of 6 km/h (4 mph), but it can reach a top speed of 40 km/h (25 mph) when scared or upset.

The animal is characterized by its large head; two large ears that cover its shoulders and radiate excess heat; a large and muscular trunk; two prominent tusk
Tusk
Tusks are elongated, continuously growing front teeth, usually but not always in pairs, that protrude well beyond the mouth of certain mammal species. They are most commonly canines, as with warthogs, wild boar, and walruses, or, in the case of elephants and narwhals, elongated incisors...

s, which are well-developed in both sexes, although more commonly in males; a short neck; a large, barrel-like body; four long and heavy legs; and a relatively short tail.

The animal is protected by a heavy but flexible layer of gray-brown skin, dotted with mostly undeveloped patches of hair and long, black hair at the tip of its tail. Its back feet have three toes that form a hoof, while the number of toes on the front feet have varied between four and five. The forehead is smoother and less convex than that of the Asian Elephant
Asian Elephant
The Asian or Asiatic elephant is the only living species of the genus Elephas and distributed in Southeast Asia from India in the west to Borneo in the east. Three subspecies are recognized — Elephas maximus maximus from Sri Lanka, the Indian elephant or E. m. indicus from mainland Asia, and E. m....

.

The trunk is the most characteristic feature of the African Bush Elephant. It is formed by the fusion and elongation of the nose and upper lip, forming a flexible and strong organ made purely of muscle.

Little scientific research has been carried out into elephants' cognitive
Animal cognition
Animal cognition is the title given to the study of the mental capacities of non-human animals. It has developed out of comparative psychology, but has also been strongly influenced by the approach of ethology, behavioral ecology, and evolutionary psychology...

 or perceptual
Perception
Perception is the process of attaining awareness or understanding of the environment by organizing and interpreting sensory information. All perception involves signals in the nervous system, which in turn result from physical stimulation of the sense organs...

 abilities. An exception is a recent report that African Bush Elephants are able to use seismic
Seismology
Seismology is the scientific study of earthquakes and the propagation of elastic waves through the Earth or through other planet-like bodies. The field also includes studies of earthquake effects, such as tsunamis as well as diverse seismic sources such as volcanic, tectonic, oceanic,...

 vibrations at infrasound
Infrasound
Infrasound is sound that is lower in frequency than 20 Hz or cycles per second, the "normal" limit of human hearing. Hearing becomes gradually less sensitive as frequency decreases, so for humans to perceive infrasound, the sound pressure must be sufficiently high...

 frequencies for communication.

Diet

African Bush Elephants are herbivorous. Their diet varies according to their habitat; elephants living in forests, partial deserts, and grasslands all eat different proportions of herbs and tree or shrubbery leaves. Elephants inhabiting the shores of Lake Kariba
Lake Kariba
Lake Kariba is the world's largest artificial lake and reservoir. It lies 1300 kilometers upstream from the Indian Ocean, along the border between Zambia and Zimbabwe...

 have been recorded eating underwater plant life. To break down the plants they consume, the African Bush Elephant has four large molar
Molar (tooth)
Molars are the rearmost and most complicated kind of tooth in most mammals. In many mammals they grind food; hence the Latin name mola, "millstone"....

s, two in each mandible of the jaw. Each of these molars is 10 cm wide and 30 cm long. Over time, these molars are worn away and new ones are grown to replace them as the elephant ages. Around the age of 15 their milk teeth
Deciduous teeth
Deciduous teeth, otherwise known as reborner teeth, baby teeth, temporary teeth and primary teeth, are the first set of teeth in the growth development of humans and many other mammals. In some Asian countries they are referred to as fall teeth as they will eventually fall out, while in almost all...

 are replaced by new ones that last until the age of 30, and then by another set which wear off past the age of 40, being replaced by the last set of teeth that last approximately until the age of 65–70. Not much later, the animal dies of starvation from not being able to feed correctly. There are known cases of over 80 year old specimens in captivity.

These animals typically ingest an average of 225 kg of vegetable matter daily, which is defecated without being fully digested. That, combined with the long distances that they can cover daily in search of more food, contributes notably to the dispersion of many plant seeds that germinate
Germination
Germination is the process in which a plant or fungus emerges from a seed or spore, respectively, and begins growth. The most common example of germination is the sprouting of a seedling from a seed of an angiosperm or gymnosperm. However the growth of a sporeling from a spore, for example the...

 in the middle of a nutrient-filled feces mound. Elephants rip apart all kind of plants, and knock down trees with the tusks if they are not able to reach the tree leaves. Elephants also drink great quantities of water, over 190 liter
Litre
pic|200px|right|thumb|One litre is equivalent to this cubeEach side is 10 cm1 litre water = 1 kilogram water The litre is a metric system unit of volume equal to 1 cubic decimetre , to 1,000 cubic centimetres , and to 1/1,000 cubic metre...

s per day.

In some national parks there is overpopulation
Overpopulation
Overpopulation is a condition where an organism's numbers exceed the carrying capacity of its habitat. The term often refers to the relationship between the human population and its environment, the Earth...

, so that managers of overpopulated parks often contact other parks with fewer specimens to transfer excess individuals.

Social behavior

The African Bush Elephant is an intelligent animal. Experiments with reasoning and learning show that they are one of the smartest mammals together with their Asian cousins. This is mostly due to their large brain
Brain
The brain is the center of the nervous system in all vertebrate and most invertebrate animals—only a few primitive invertebrates such as sponges, jellyfish, sea squirts and starfishes do not have one. It is located in the head, usually close to primary sensory apparatus such as vision, hearing,...

.

Herds are made up of related females and their young, directed by the eldest female, called the matriarch. Infrequently, an adult male goes with them, but those usually leave the pack when reaching adolescence to form bachelor herds with other elephants of the same age. Later they lead a solitary life, approaching the female herds only during the mating season. Nevertheless, elephants do not get too far from their families and recognize them when re-encountered. Sometimes, several female herds can blend for a period of time, reaching even hundreds of individuals.

The matriarch is the one who decides the route and shows to each other member of the herd all the water sources she knows, which the rest will memorize in the future. The relations among the members of the herd are very tight; when a female gives birth to a baby the rest go to acknowledge it touching her with the trunk; and when an old elephant dies the rest of the herd will stay by the corpse for a while. The famous elephant graveyard
Elephant's graveyard
An elephant graveyard is a place where, according to legend, older elephants instinctively direct themselves when they reach a certain age. They then die there alone, far from the group....

s are a myth, but it is true that these animals can recognize a carcass of its species when they find one during their trips, and even if it is a stranger, they form around it and sometimes they even touch its forehead with their trunk.

Mating happens when the female becomes receptive, an event that can occur anytime during the year. When she is ready, she starts emitting infrasounds that attract the males, sometimes from kilometers away. The adult males start arriving at the herd during the following days and begin fighting, causing some injuries and even broken tusks. The female shows her acceptance of the victor by rubbing her body against his. They mate, and then both go their own way. After 22 months of gestation (the longest among mammals), the female gives birth to a single 90 cm high calf which weighs more than 100 kg. The baby feeds on the mothers milk until the age of 5, but also eats solid food from as early as 6 months old. Just a few days after birth, the calf can follow the herd by foot.

Some African Bush Elephants will attack and kill rhinoceros
Rhinoceros
Rhinoceros , also known as rhino, is a group of five extant species of odd-toed ungulates in the family Rhinocerotidae. Two of these species are native to Africa and three to southern Asia....

es. This behavior, when it occurs, is mostly observed with younger adult male elephants who have come into musth
Musth
Musth or must is a periodic condition in bull elephants, characterized by highly aggressive behavior, accompanied by a large rise in reproductive hormones - testosterone levels in an elephant in musth can be as much as 60 times greater than in the same elephant at other times...

 prematurely.

Predators

The adult African Bush Elephant generally has no natural predators
Predation
In ecology, predation describes a biological interaction where a predator feeds on its prey . Predators may or may not kill their prey prior to feeding on them, but the act of predation always results in the death of its prey and the eventual absorption of the prey's tissue through consumption...

 due to its great size, but the calves (especially the newborn) are vulnerable to lion
Lion
The lion is one of the four big cats in the genus Panthera, and a member of the family Felidae. With some males exceeding 250 kg in weight, it is the second-largest living cat after the tiger...

 and crocodile
Nile crocodile
The Nile crocodile or Common crocodile is an African crocodile which is common in Somalia, Ethiopia, Uganda, Kenya, Egypt, Tanzania, Zambia, Zimbabwe, Gabon, South Africa, Malawi, Sudan, Botswana, and Cameroon...

 attacks, and (rarely) to leopard
Leopard
The leopard , Panthera pardus, is a member of the Felidae family and the smallest of the four "big cats" in the genus Panthera, the other three being the tiger, lion, and jaguar. The leopard was once distributed across eastern and southern Asia and Africa, from Siberia to South Africa, but its...

 and hyena
Hyena
Hyenas or Hyaenas are the animals of the family Hyaenidae of suborder feliforms of the Carnivora. It is the fourth smallest biological family in the Carnivora , and one of the smallest in the mammalia...

 attacks. There are prides of lions who prey on both infants and juveniles especially in the drought months. Lions in Chobe National Park in Botswana have been observed for some time taking both infants (23% of elephant kills) and juveniles. Predation, as well as drought, contribute significantly to infant mortality.

Humans are the elephant's major predator. Elephants have been hunted for meat, skin, bones, and tusks. Elephant trophy hunting
Trophy hunting
Trophy hunting is the selective hunting of wild game animals. Although parts of the slain animal may be kept as a hunting trophy or memorial , the carcass itself is sometimes used as food....

 increased in the 19th and 20th centuries, when tourism and plantations increasingly attracted sport hunters. In 1989, hunting of the African Bush Elephant for ivory trading
Ivory trade
The ivory trade is the commercial, often illegal trade in the ivory tusks of the hippopotamus, walrus, narwhal, mammoth, and most commonly, Asian and African elephants....

 was forbidden, after the elephant population fell from several million at the beginning of the 20th century to fewer than 700,000. Trophy hunting continues today. The population of African Bush Elephants was halved during the 1980s. Scientists then estimated that, if no protective measures were taken, the wild elephant would be extinct by 1995. The protection that the elephant now receives has been partially successful, but despite increasingly severe penalties imposed by governments against illegal hunting, poaching is still common. CITES still considers this species as threatened with extinction.

Species differences

A 2010 genetic study confirmed that the African Bush Elephant and the African Forest Elephant
African Forest Elephant
The African Forest Elephant is a forest dwelling elephant of the Congo Basin. Formerly considered either a synonym or a subspecies of the African Savanna Elephant , a 2010 study established that the two are distinct species...

 are distinct species. By sequencing DNA
DNA sequencing
DNA sequencing includes several methods and technologies that are used for determining the order of the nucleotide bases—adenine, guanine, cytosine, and thymine—in a molecule of DNA....

 of 375 nuclear gene
Nuclear gene
A Nuclear gene is a gene located in the cell nucleus of a eukaryote. The term is used to distinguish nuclear genes from the genes in the mitochondrion, and in case of plants, also the chloroplast, which host their own genetic system and can produce proteins from scratch...

s, scientists determined that the two species diverged around the same time as the Asian elephant
Asian Elephant
The Asian or Asiatic elephant is the only living species of the genus Elephas and distributed in Southeast Asia from India in the west to Borneo in the east. Three subspecies are recognized — Elephas maximus maximus from Sri Lanka, the Indian elephant or E. m. indicus from mainland Asia, and E. m....

 and the woolly mammoth
Woolly mammoth
The woolly mammoth , also called the tundra mammoth, is a species of mammoth. This animal is known from bones and frozen carcasses from northern North America and northern Eurasia with the best preserved carcasses in Siberia...

 and are as distinct from one another as those two species. , conservation organizations such as the United Nations Environment Programme
United Nations Environment Programme
The United Nations Environment Programme coordinates United Nations environmental activities, assisting developing countries in implementing environmentally sound policies and practices. It was founded as a result of the United Nations Conference on the Human Environment in June 1972 and has its...

's World Conservation Monitoring Centre
World Conservation Monitoring Centre
The United Nations Environment Programme's World Conservation Monitoring Centre is an executive agency of the United Nations Environment Programme, based in Cambridge in the United Kingdom. UNEP-WCMC has been part of UNEP since 2000, and has responsibility for biodiversity assessment and support...

 and the International Union for Conservation of Nature (IUCN) had not distinguished between the two species of African elephants for purposes of assessing their conservation status. , the IUCN Red List
IUCN Red List
The IUCN Red List of Threatened Species , founded in 1963, is the world's most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the world's main authority on the conservation status of species...

 classified African elephants as a whole as vulnerable species
Vulnerable species
On 30 January 2010, the IUCN Red List of Threatened Species identified 9694 Vulnerable species, subspecies and varieties, stocks and sub-populations.-References:...

 and the Central African elephant population (forest elephants) as Endangered.

Another possible species or subspecies formerly existed, but although formally described it has not been widely recognized by the scientific community. The North African Elephant
North African Elephant
The North African Elephant was a possible subspecies of the African Bush Elephant , or possibly a separate elephant species, that existed in North Africa until becoming extinct in Ancient Roman times. These were the famous war elephants used by Carthage in the Punic Wars, their conflict with the...

 (Loxodonta africana pharaohensis), also known as the Carthaginian Elephant or Atlas Elephant, was the animal famously used as a war elephant
War elephant
A war elephant was an elephant trained and guided by humans for combat. Their main use was to charge the enemy, trampling them and breaking their ranks. A division of war elephants is known as elephantry....

 by Carthage
Carthage
Carthage , implying it was a 'new Tyre') is a major urban centre that has existed for nearly 3,000 years on the Gulf of Tunis, developing from a Phoenician colony of the 1st millennium BC...

 in its long struggle against Rome
Rome
Rome is the capital of Italy and the country's largest and most populated city and comune, with over 2.7 million residents in . The city is located in the central-western portion of the Italian Peninsula, on the Tiber River within the Lazio region of Italy.Rome's history spans two and a half...

.

Conservation

While the species is designated as vulnerable
Vulnerable species
On 30 January 2010, the IUCN Red List of Threatened Species identified 9694 Vulnerable species, subspecies and varieties, stocks and sub-populations.-References:...

, conditions vary somewhat by region within Eastern
East Africa
East Africa or Eastern Africa is the easterly region of the African continent, variably defined by geography or geopolitics. In the UN scheme of geographic regions, 19 territories constitute Eastern Africa:...

 and Southern Africa
Southern Africa
Southern Africa is the southernmost region of the African continent, variably defined by geography or geopolitics. Within the region are numerous territories, including the Republic of South Africa ; nowadays, the simpler term South Africa is generally reserved for the country in English.-UN...

.

In 2006, an elephant slaughter
2006 Zakouma elephant slaughter
The 2006 Zakouma elephant slaughter refers to a series of poaching massacres of African elephants in the vicinity of Zakouma National Park in southeastern Chad. These killings were documented in aerial surveys conducted from May through August 2006 and total at least 100 animals...

 was documented in southeastern Chad
Chad
Chad , officially known as the Republic of Chad, is a landlocked country in Central Africa. It is bordered by Libya to the north, Sudan to the east, the Central African Republic to the south, Cameroon and Nigeria to the southwest, and Niger to the west...

 by aerial surveys. A series of poaching
Poaching
Poaching is the illegal taking of wild plants or animals contrary to local and international conservation and wildlife management laws. Violations of hunting laws and regulations are normally punishable by law and, collectively, such violations are known as poaching.It may be illegal and in...

 incidents, resulting in the killing of over 100 elephants, was carried out during the late spring and summer of 2006 in the vicinity of Zakouma National Park
Zakouma National Park
Zakouma National Park is a national park, located between Sarh and Am Timan, in the south of Chad. Created in 1963, it was Chad's first national park, and has an area of almost...

. This region has a decades-old history of poaching of elephants, which has caused the elephant population of the region, which exceeded 300,000 in 1970, to drop to approximately 10,000 today. The African Bush Elephant officially is protected by Chadian government, but the resources and manpower provided by the government (with some European Union
European Union
The European Union is an economic and political union of 27 independent member states which are located primarily in Europe. The EU traces its origins from the European Coal and Steel Community and the European Economic Community , formed by six countries in 1958...

 assistance) have proven insufficient to stop the poaching.

Human encroachment into or adjacent to natural areas where Bush Elephants occur has led to recent research into methods of safely driving groups of elephants away from humans, including the discovery that playback of the recorded sounds of angry honey bees
European honey bee
The Western honey bee or European honey bee is a species of honey bee. The genus Apis is Latin for "bee", and mellifera comes from Latin melli- "honey" and ferre "to bear"—hence the scientific name means "honey-bearing bee"...

are remarkably effective at prompting elephants to flee an area.

External links

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