Anoplosaurus
Encyclopedia
Anoplosaurus is an extinct genus
of nodosaurid dinosaur
, from the late Albian
-age Lower Cretaceous Cambridge Greensand
of Cambridgeshire
, England
. It has been classified with the armored dinosaurs
and ornithopod
s, but current thought has been in agreement with the "armored dinosaur" interpretation.
, back
, and sacrum
, parts of the pectoral girdle, humerus
fragments, part of the left femur
, left tibia
, foot
bones, ribs, and other fragments. He regarded it as possibly juvenile
, due to its small size. Other workers began to see it as an armored
dinosaur, until 1923 when Franz Nopcsa suggested that some of the remains belonged to a camptosaur
, and other remains, which he removed from the genus, belonged to Acanthopholis
. This suggestion was followed (with modifications as iguanodontian taxonomy changed over the years), Suberbiola and Barrett reexamined the material. They wrote that the material all belonged to a primitive
nodosaurid, the lack of armor possibly due to the young age of the animal at death. Reviews since then have followed this interpretation of the genus as an armored dinosaur.
A second species, A. major, was named by Seeley in 1879 for a neck vertebra and three partial caudal
vertebrae he removed from Acanthopholis stereocercus, from the same formation as the type species
. This species now appears to be chimeric
, the neck vertebra coming from an ankylosaur, the caudals from an indeterminate iguanodont.
al, low-slung herbivore
, with armour on its body for protection.
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of nodosaurid dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...
, from the late Albian
Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous epoch/series. Its approximate time range is 112.0 ± 1.0 Ma to 99.6 ± 0.9 Ma...
-age Lower Cretaceous Cambridge Greensand
Cambridge Greensand
The Cambridge Greensand is a geological formation in England whose strata date back to the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.-Vertebrate paleofauna:...
of Cambridgeshire
Cambridgeshire
Cambridgeshire is a county in England, bordering Lincolnshire to the north, Norfolk to the northeast, Suffolk to the east, Essex and Hertfordshire to the south, and Bedfordshire and Northamptonshire to the west...
, England
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...
. It has been classified with the armored dinosaurs
Ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are first known to have appeared in the early Jurassic Period of...
and ornithopod
Ornithopod
Ornithopods or members of the clade Ornithopoda are a group of ornithischian dinosaurs that started out as small, bipedal running grazers, and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world, and dominated the North American...
s, but current thought has been in agreement with the "armored dinosaur" interpretation.
History
Harry Govier Seeley named this genus in 1879 for a partial postcranial skeleton, composed of dentary fragments, numerous vertebra from the neckNeck
The neck is the part of the body, on many terrestrial or secondarily aquatic vertebrates, that distinguishes the head from the torso or trunk. The adjective signifying "of the neck" is cervical .-Boner anatomy: The cervical spine:The cervical portion of the human spine comprises seven boney...
, back
Dorsum (biology)
In anatomy, the dorsum is the upper side of animals that typically run, fly, or swim in a horizontal position, and the back side of animals that walk upright. In vertebrates the dorsum contains the backbone. The term dorsal refers to anatomical structures that are either situated toward or grow...
, and sacrum
Sacrum
In vertebrate anatomy the sacrum is a large, triangular bone at the base of the spine and at the upper and back part of the pelvic cavity, where it is inserted like a wedge between the two hip bones. Its upper part connects with the last lumbar vertebra, and bottom part with the coccyx...
, parts of the pectoral girdle, humerus
Humerus
The humerus is a long bone in the arm or forelimb that runs from the shoulder to the elbow....
fragments, part of the left femur
Femur
The femur , or thigh bone, is the most proximal bone of the leg in tetrapod vertebrates capable of walking or jumping, such as most land mammals, birds, many reptiles such as lizards, and amphibians such as frogs. In vertebrates with four legs such as dogs and horses, the femur is found only in...
, left tibia
Tibia
The tibia , shinbone, or shankbone is the larger and stronger of the two bones in the leg below the knee in vertebrates , and connects the knee with the ankle bones....
, foot
Foot
The foot is an anatomical structure found in many vertebrates. It is the terminal portion of a limb which bears weight and allows locomotion. In many animals with feet, the foot is a separate organ at the terminal part of the leg made up of one or more segments or bones, generally including claws...
bones, ribs, and other fragments. He regarded it as possibly juvenile
Juvenile (organism)
A juvenile is an individual organism that has not yet reached its adult form, sexual maturity or size. Juveniles sometimes look very different from the adult form, particularly in terms of their colour...
, due to its small size. Other workers began to see it as an armored
Armour (zoology)
Armour in animals is external or superficial protection against attack by predators, formed as part of the body , usually through the hardening of body tissues, outgrowths or secretions. It has therefore mostly developed in 'prey' species...
dinosaur, until 1923 when Franz Nopcsa suggested that some of the remains belonged to a camptosaur
Camptosaurus
Camptosaurus is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America. The name means 'flexible lizard', ....
, and other remains, which he removed from the genus, belonged to Acanthopholis
Acanthopholis
Acanthopholis is a genus of ankylosaurian dinosaur in the family Nodosauridae that lived during the Early Cretaceous Period of England....
. This suggestion was followed (with modifications as iguanodontian taxonomy changed over the years), Suberbiola and Barrett reexamined the material. They wrote that the material all belonged to a primitive
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...
nodosaurid, the lack of armor possibly due to the young age of the animal at death. Reviews since then have followed this interpretation of the genus as an armored dinosaur.
A second species, A. major, was named by Seeley in 1879 for a neck vertebra and three partial caudal
Tail
The tail is the section at the rear end of an animal's body; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, reptiles, and birds...
vertebrae he removed from Acanthopholis stereocercus, from the same formation as the type species
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
. This species now appears to be chimeric
Chimera (paleontology)
In paleontology, a chimera is a fossil which was reconstructed with elements coming from more than a single species of animal. A now classic example of chimera is Protoavis.-List of paleontological chimeras:*Brontosaurus*Lametasaurus...
, the neck vertebra coming from an ankylosaur, the caudals from an indeterminate iguanodont.
Palaeobiology
As a possible nodosaurid, Anoplosaurus would have been a quadrupedQuadruped
Quadrupedalism is a form of land animal locomotion using four limbs or legs. An animal or machine that usually moves in a quadrupedal manner is known as a quadruped, meaning "four feet"...
al, low-slung herbivore
Herbivore
Herbivores are organisms that are anatomically and physiologically adapted to eat plant-based foods. Herbivory is a form of consumption in which an organism principally eats autotrophs such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in...
, with armour on its body for protection.