Ardisia elliptica is an evergreen tree, also known as the shoebutton, native to the west coast of India

India

India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

, Sri Lanka

Sri Lanka

Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka is a country off the southern coast of the Indian subcontinent. Known until 1972 as Ceylon , Sri Lanka is an island surrounded by the Indian Ocean, the Gulf of Mannar and the Palk Strait, and lies in the vicinity of India and the...

, Indochina

Indochina

The Indochinese peninsula, is a region in Southeast Asia. It lies roughly southwest of China, and east of India. The name has its origins in the French, Indochine, as a combination of the names of "China" and "India", and was adopted when French colonizers in Vietnam began expanding their territory...

, Malaysia, Indonesia

Indonesia

Indonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...

 and New Guinea

New Guinea

New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

. It a prolific reproducer which has made it a successful invasive species in other locations.

A. elliptica is a tropical understory shrub that can reach heights of up 5 meters. Undamaged plants in forest habitats are characterized by a single stem, producing short, perpendicular branches. Leaves are ellipic to elliptic-obovate, entire, leathery and alternate. Umbellate inflorescences develop in leaf axils of branch leaves. Petals are light pink. Fruits are drupes that first turn red as they mature and then deep purple / black. Pulp staining fingers a deep purple. Seeds are approximately spherical with a diameter of about 5 mm.

Given ideal conditions, individuals can reach reproductive maturity in 2–4 years in the field and 1–2 years in a shade house. Large adults in bright forested sites have been measured producing up to 400 fruits. However, adults can also successfully set fruit

Fruit

In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 under shady conditions. Seed

Seed

A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s do not have any long-term dormancy (i.e., greater than 6 months), however, seedlings and juveniles can survive under very shady conditions for many years. Given enough light, juveniles rapidly develop into reproductive adults. Its fruit is readily consumed by both avian and mammalian frugivores, rapid spread across a landscape is possible.

The species has become invasive in Hawai‘i, Southern Florida

Florida

Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

, Okinawa and Jamaica

Jamaica

Jamaica is an island nation of the Greater Antilles, in length, up to in width and 10,990 square kilometres in area. It is situated in the Caribbean Sea, about south of Cuba, and west of Hispaniola, the island harbouring the nation-states Haiti and the Dominican Republic...

.

Phytoconstituents

Rapanone, bauerenol, amyrin, syringic acid

Syringic acid

Syringic acid is a naturally occurring O-methylated trihydroxybenzoic acid, a type of chemical compound. It can be found in the açaí palm and its oil , in Ardisia elliptica...

, isorhamnetin

Isorhamnetin

Isorhamnetin is an O-methylated flavonol, a type of chemical compound. It can be found in Tagetes lucida, a psychedelic plant from Mexico and Central America.-Metabolism:...

 and quercetin

Quercetin

Quercetin , a flavonol, is a plant-derived flavonoid found in fruits, vegetables, leaves and grains. It also may be used as an ingredient in supplements, beverages or foods.-Occurrence:...

, bergenin

Bergenin

Bergenin alias Cuscutin is trihydroxybenzoic acid glycoside. It is the C-glycoside of 4-O-methyl gallic acid. It possesses an O-demethylated derivative called norbergenin. These are chemical compounds and drugs of Ayurveda, commonly known as Paashaanbhed...

, 5-(Z-Heptadec-4'-enyl)resorcinol, 5-pentadecylresorcinol.

EthnoPharmacology

In Malaysia, a decoction of leaves is said to assuage retrosternal pains, and a paste made from the leaves is used to treat herpes and measles. In Thai traditional medicine, the fruits are used to cure diarrhoea with fever. In Southeast Asia leaves are used to treat scabies, and fruit for intestinal worms.