The Squinting Bush Brown (Bicyclus anynana) is a butterfly

Butterfly

A butterfly is a mainly day-flying insect of the order Lepidoptera, which includes the butterflies and moths. Like other holometabolous insects, the butterfly's life cycle consists of four parts: egg, larva, pupa and adult. Most species are diurnal. Butterflies have large, often brightly coloured...

 of the Nymphalidae

Nymphalidae

The Nymphalidae is a family of about 5,000 species of butterflies which are distributed throughout most of the world. These are usually medium sized to large butterflies. Most species have a reduced pair of forelegs and many hold their colourful wings flat when resting. They are also called...

 family. It is found in Zimbabwe

Zimbabwe

Zimbabwe is a landlocked country located in the southern part of the African continent, between the Zambezi and Limpopo rivers. It is bordered by South Africa to the south, Botswana to the southwest, Zambia and a tip of Namibia to the northwest and Mozambique to the east. Zimbabwe has three...

, Mozambique

Mozambique

Mozambique, officially the Republic of Mozambique , is a country in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west and Swaziland and South Africa to the southwest...

, KwaZulu-Natal

KwaZulu-Natal

KwaZulu-Natal is a province of South Africa. Prior to 1994, the territory now known as KwaZulu-Natal was made up of the province of Natal and the homeland of KwaZulu....

, from Swaziland

Swaziland

Swaziland, officially the Kingdom of Swaziland , and sometimes called Ngwane or Swatini, is a landlocked country in Southern Africa, bordered to the north, south and west by South Africa, and to the east by Mozambique...

 to Ethiopia

Ethiopia

Ethiopia , officially known as the Federal Democratic Republic of Ethiopia, is a country located in the Horn of Africa. It is the second-most populous nation in Africa, with over 82 million inhabitants, and the tenth-largest by area, occupying 1,100,000 km2...

, southern Somalia

Somalia

Somalia , officially the Somali Republic and formerly known as the Somali Democratic Republic under Socialist rule, is a country located in the Horn of Africa. Since the outbreak of the Somali Civil War in 1991 there has been no central government control over most of the country's territory...

, Kenya

Kenya

Kenya , officially known as the Republic of Kenya, is a country in East Africa that lies on the equator, with the Indian Ocean to its south-east...

, Uganda

Uganda

Uganda , officially the Republic of Uganda, is a landlocked country in East Africa. Uganda is also known as the "Pearl of Africa". It is bordered on the east by Kenya, on the north by South Sudan, on the west by the Democratic Republic of the Congo, on the southwest by Rwanda, and on the south by...

, eastern Zaire

Zaire

The Republic of Zaire was the name of the present Democratic Republic of the Congo between 27 October 1971 and 17 May 1997. The name of Zaire derives from the , itself an adaptation of the Kongo word nzere or nzadi, or "the river that swallows all rivers".-Self-proclaimed Father of the Nation:In...

, the Comoros

Comoros

The Comoros , officially the Union of the Comoros is an archipelago island nation in the Indian Ocean, located off the eastern coast of Africa, on the northern end of the Mozambique Channel, between northeastern Mozambique and northwestern Madagascar...

 and Socotra

Socotra

Socotra , also spelt Soqotra, is a small archipelago of four islands in the Indian Ocean. The largest island, also called Socotra, is about 95% of the landmass of the archipelago. It lies some east of the Horn of Africa and south of the Arabian Peninsula. The island is very isolated and through...

.
The wingspan

Wingspan

The wingspan of an airplane or a bird, is the distance from one wingtip to the other wingtip. For example, the Boeing 777 has a wingspan of about ; and a Wandering Albatross caught in 1965 had a wingspan of , the official record for a living bird.The term wingspan, more technically extent, is...

 is 35–40 mm for males and 49-45 for females. There are two extended generations per year. The wet season form is on wing in spring and summer and the dry season form in autumn and winter

The larvae feed on various Poaceae

Poaceae

The Poaceae is a large and nearly ubiquitous family of flowering plants. Members of this family are commonly called grasses, although the term "grass" is also applied to plants that are not in the Poaceae lineage, including the rushes and sedges...

 species. Larvae have been reared on Ehrharta erecta

Ehrharta erecta

Ehrharta erecta is a species of grass also known as panic veldtgrass. The species is native to Southern Africa and Yemen. It is a documented invasive species in the United States, New Zealand, Australia, southern Europe, and China....

.

Research

Researchers at Yale University

Yale University

Yale University is a private, Ivy League university located in New Haven, Connecticut, United States. Founded in 1701 in the Colony of Connecticut, the university is the third-oldest institution of higher education in the United States...

 have discovered that the temperature at which the larvae are raised has an effect on the imago

Imago

In biology, the imago is the last stage of development of an insect, after the last ecdysis of an incomplete metamorphosis, or after emergence from the pupa where the metamorphosis is complete...

's sexual behaviour. Females raised in a cooler environment are more likely to actively court male butterflies. In a warmer environment, the male butterflies traditionally take on the active role in courtship

Subspecies

• Bicyclus anynana anynana (Kenya to Tanzania, Zambia, Malawi, Mozambique, Rhodesia, Botswana, South Africa, Comoro Islands)
• Bicyclus anynana centralis Condamin, 1968 (Uganda, southern Zaire, eastern Zaire, northern Angola)
• Bicyclus anynana socotrana (Butler, 1881) (Socotra Island)