The Black-capped Donacobius (Donacobius atricapilla) is a conspicuous, vocal South American bird

Bird

Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

. It is found in tropical swamps and wetlands in Argentina

Argentina

Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

, Bolivia

Bolivia

Bolivia officially known as Plurinational State of Bolivia , is a landlocked country in central South America. It is the poorest country in South America...

, Brazil

Brazil

Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

, Colombia

Colombia

Colombia, officially the Republic of Colombia , is a unitary constitutional republic comprising thirty-two departments. The country is located in northwestern South America, bordered to the east by Venezuela and Brazil; to the south by Ecuador and Peru; to the north by the Caribbean Sea; to the...

, Ecuador

Ecuador

Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

, French Guiana

French Guiana

French Guiana is an overseas region of France, consisting of a single overseas department located on the northern Atlantic coast of South America. It has borders with two nations, Brazil to the east and south, and Suriname to the west...

, Guyana

Guyana

Guyana , officially the Co-operative Republic of Guyana, previously the colony of British Guiana, is a sovereign state on the northern coast of South America that is culturally part of the Anglophone Caribbean. Guyana was a former colony of the Dutch and of the British...

, Paraguay

Paraguay

Paraguay , officially the Republic of Paraguay , is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to the east and northeast, and Bolivia to the northwest. Paraguay lies on both banks of the Paraguay River, which runs through the center of the...

, Peru

Peru

Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

, Suriname

Suriname

Suriname , officially the Republic of Suriname , is a country in northern South America. It borders French Guiana to the east, Guyana to the west, Brazil to the south, and on the north by the Atlantic Ocean. Suriname was a former colony of the British and of the Dutch, and was previously known as...

, and Venezuela

Venezuela

Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

; also Panama

Panama

Panama , officially the Republic of Panama , is the southernmost country of Central America. Situated on the isthmus connecting North and South America, it is bordered by Costa Rica to the northwest, Colombia to the southeast, the Caribbean Sea to the north and the Pacific Ocean to the south. The...

 of Central America

Central America

Central America is the central geographic region of the Americas. It is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. When considered part of the unified continental model, it is considered a subcontinent...

.

Taxonomy

The Black-capped Donacobius is the only member of the genus Donacobius. Its familial placement is not established, and ornithologists disagree as to its closest relations. In the 19th century, it was placed in the Turdidae, and in the 20th century, moved to the Mimidae. It had various English names, including the "Black-capped Mockingthrush". In the 1980s and 1990s, suggestions that it was a type of wren (Troglodytidae) were accepted by the South American Classification Committee (SACC), the American Ornithologists Union (AOU) and most other authorities. More recently, listing organizations and authors follow Van Remsen and Keith Barker

Keith Barker

Keith Barker is an English first-class cricketer who plays for Warwickshire. He is an all-rounder....

's conclusion that it is not a wren either, but instead most closely related to an Old World

Old World

The Old World consists of those parts of the world known to classical antiquity and the European Middle Ages. It is used in the context of, and contrast with, the "New World" ....

 (probably African) lineage. A current proposal to the SACC would create a monotypic family, Donacobiidae, for this species, but this is not universally accepted as some authorities insist it may prove to be a member of an existing Old World family, presumably the Megaluridae

Megaluridae

Locustellidae is a newly recognized family of small insectivorous songbirds , formerly placed in the Old World warbler "wastebin" family. It contains the grass-warblers, grassbirds, and the Bradypterus "bush-warblers". These birds occur mainly in Eurasia, Africa, and the Australian region...

 which seem to be its closest living relatives. These are long-tailed "warbler

Warbler

There are a number of Passeriformes called "warblers". They are not particularly closely related, but share some characteristics, such as being fairly small, vocal and insectivorous....

s" from around the Indian Ocean

Indian Ocean

The Indian Ocean is the third largest of the world's oceanic divisions, covering approximately 20% of the water on the Earth's surface. It is bounded on the north by the Indian Subcontinent and Arabian Peninsula ; on the west by eastern Africa; on the east by Indochina, the Sunda Islands, and...

 region, many of which despite their cryptic

Crypsis

In ecology, crypsis is the ability of an organism to avoid observation or detection by other organisms. It may be either a predation strategy or an antipredator adaptation, and methods include camouflage, nocturnality, subterranean lifestyle, transparency, and mimicry...

 coloration, smaller size and more solitary habits, are similar to the Donacobius in stance, habitat and some aspects of their behavior (such as the nests); these "warblers" also have a similarly voice, but being higher-pitched they sound less grating and more like locust

Locust

Locusts are the swarming phase of short-horned grasshoppers of the family Acrididae. These are species that can breed rapidly under suitable conditions and subsequently become gregarious and migratory...

s (namely Locustella).

Habitat, including Amazon Basin

Black-capped Donacobiuses are common in a wide range of Amazonian wetlands, including oxbow lake

Oxbow lake

An oxbow lake is a U-shaped body of water formed when a wide meander from the main stem of a river is cut off to create a lake. This landform is called an oxbow lake for the distinctive curved shape, named after part of a yoke for oxen. In Australia, an oxbow lake is called a billabong, derived...

s, riparian zones, and other areas with tall dense aquatic or semi-aquatic vegetation. A third of the species range is outside the Amazon Basin

Amazon Basin

The Amazon Basin is the part of South America drained by the Amazon River and its tributaries that drains an area of about , or roughly 40 percent of South America. The basin is located in the countries of Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, and Venezuela...

, from Panama, northern Colombia, and western Venezuela, the Orinoco River system of Venezuela, to southeast coastal and inland Brazil, and neighboring countries southward, Paraguay, and extreme northern Argentina.

Behavior

Mating for life, pairs of Black-capped Donacobiuses can be seen frequently and throughout the day atop thickets of dense lakeside or streamside vegetation. They often will engage in antiphonic dueting. Adult offspring will remain with their parents and help raise siblings from subsequent nesting periods in a system of cooperative breeding

Cooperative breeding

Cooperative breeding is a social system in which individuals contribute care to offspring that are not their own at the expense of their own reproduction . When reproduction is monopolized by one or few of the adult group members and most adults do not reproduce, but help rear the breeder’s...

.

External links

• Black-capped Donacobius videos on the Internet Bird Collection
• Photo; Article sunbirdtours(Venezuelan species)
• Photo-Medium Res; Article chandra.as.utexas.edu
• Black-capped Donacobius photo gallery VIREO Photo-High Res