Black Tupelo
Encyclopedia
Nyssa sylvatica, commonly known as black tupelo, tupelo, or black gum, is a medium-sized deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 native to eastern North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 from New England
New England
New England is a region in the northeastern corner of the United States consisting of the six states of Maine, New Hampshire, Vermont, Massachusetts, Rhode Island, and Connecticut...

 and southern Ontario
Ontario
Ontario is a province of Canada, located in east-central Canada. It is Canada's most populous province and second largest in total area. It is home to the nation's most populous city, Toronto, and the nation's capital, Ottawa....

 south to central Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

 and eastern Texas
Texas
Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

, as well as Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

.

Names

Nyssa sylvaticas genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 name (Nyssa) refers to a Greek
Ancient Greece
Ancient Greece is a civilization belonging to a period of Greek history that lasted from the Archaic period of the 8th to 6th centuries BC to the end of antiquity. Immediately following this period was the beginning of the Early Middle Ages and the Byzantine era. Included in Ancient Greece is the...

 water nymph
Naiad
In Greek mythology, the Naiads or Naiades were a type of nymph who presided over fountains, wells, springs, streams, and brooks....

; the species epithet sylvatica refers to its woodland habitat.

The species' common name tupelo is of Native American
Native Americans in the United States
Native Americans in the United States are the indigenous peoples in North America within the boundaries of the present-day continental United States, parts of Alaska, and the island state of Hawaii. They are composed of numerous, distinct tribes, states, and ethnic groups, many of which survive as...

 origin, coming from the Creek
Creek language
The Creek language, also known as Muskogee or Muscogee , is a Muskogean language spoken by Muscogee and Seminole people primarily in the U.S. states of Oklahoma and Florida....

 words ito ‘tree’ and opilwa ‘swamp’; it was in use by the mid-18th century

While these trees are often known as simply "tupelo", the fuller name black tupelo helps distinguish it from the other species of the tupelo
Tupelo
The tupelo , black gum, or pepperidge tree, genus Nyssa , is a small genus of about 9 to 11 species of trees with alternate, simple leaves...

 genus (Nyssa), some of which have overlapping ranges, such as water tupelo (N. aquatica) and swamp tupelo (N. biflora). The name "tupelo" is used primarily in the American South
Southern United States
The Southern United States—commonly referred to as the American South, Dixie, or simply the South—constitutes a large distinctive area in the southeastern and south-central United States...

; northward and in Appalachia
Appalachian Mountains
The Appalachian Mountains #Whether the stressed vowel is or ,#Whether the "ch" is pronounced as a fricative or an affricate , and#Whether the final vowel is the monophthong or the diphthong .), often called the Appalachians, are a system of mountains in eastern North America. The Appalachians...

, the tree is more commonly called the black gum or the sour gum, although no part of the plant is particularly gummy. Both of these names contrast it with a different tree species with a broadly overlapping range, the sweet gum (Liquidambar styraciflua), which does produce an aromatic resin. Another common name used occasionally is the Northeast
Northeastern United States
The Northeastern United States is a region of the United States as defined by the United States Census Bureau.-Composition:The region comprises nine states: the New England states of Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island and Vermont; and the Mid-Atlantic states of New...

 is pepperidge.

On Martha's Vineyard
Martha's Vineyard
Martha's Vineyard is an island located south of Cape Cod in Massachusetts, known for being an affluent summer colony....

, in Massachusetts
Massachusetts
The Commonwealth of Massachusetts is a state in the New England region of the northeastern United States of America. It is bordered by Rhode Island and Connecticut to the south, New York to the west, and Vermont and New Hampshire to the north; at its east lies the Atlantic Ocean. As of the 2010...

, this species is called "beetlebung", perhaps for its use in making the mallet known as a beetle, used for hammering bung
Bung
A bung is truncated cylindrical or conical closure to seal a container, such as a bottle, tube or barrel. Unlike a lid which encloses a container from the outside without displacing the inner volume, a bung is partially inserted inside the container to act as a seal...

s (stoppers) into barrels.

Description

Nyssa sylvatica grows to 20–25 m (65.6–82 ft) tall, rarely to 35 metres (114.8 ft), with a trunk diameter of 50–100 cm (19.7–39.4 in), rarely up to 170 centimetres (66.9 in). These trees typically have a straight trunk with the branches extending outward at right angles. The bark
Bark
Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines and shrubs. Bark refers to all the tissues outside of the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner...

 is dark gray and flaky when young, but it becomes furrowed with age, resembling alligator
Alligator
An alligator is a crocodilian in the genus Alligator of the family Alligatoridae. There are two extant alligator species: the American alligator and the Chinese alligator ....

 hide on very old stems. The twigs of this tree are reddish-brown, usually hidden by a greyish skin. The pith is chambered with greenish partitions.

The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 of this species are variable in size and shape. They can be oval, elliptical, or obovate, and 5–12 cm (2–5 in) long. They have lustrous upper surfaces, with entire, often wavy margins. The foliage turns purple in autumn, eventually becoming an intense bright scarlet.

The flowers
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

 are very small, in greenish-white in clusters at the top of a long stalk. The fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is a black-blue, ovoid stone fruit, about 10 mm long with a thin, oily, bitter-to-sour tasting flesh. There are from one to three such fruit together on a long slender stalk.

Additional characteristics include:
  • Bark: Light reddish brown, deeply furrowed and scaly. Branchlets at first pale green to orange, sometimes smooth, often downy, later dark brown.
  • Wood: Pale yellow, sapwood white; heavy, strong, very tough, hard to split, not durable in contact with the soil. Used for turnery. Sp. gr., 0.6353; weight of cu. ft., 39.59.
  • Winter buds: Dark red, obtuse, one-fourth of an inch long. Inner scales enlarge with the growing shoot, becoming red before they fall.
  • Leaves: Alternate, often crowded at the end of the lateral branches, simple, linear, oblong to oval, two to five inches (127 mm) long, one-half to three inches (76 mm) broad, wedge-shaped or rounded at base, entire, with margin slightly thickened, acute or acuminate. They come out of the bud conduplicate, coated beneath with rusty tomentum, when full grown are thick, dark green, very shining above, pale and often hairy beneath. Feather-veined, midrib and primary veins prominent beneath. In autumn they turn bright scarlet, or yellow and scarlet. Petioles one-quarter to one-half an inch long, slender or stout, terete or margined, often red.
  • Flowers: May, June, when leaves are half grown. Polygamodiœcious, yellowish green, borne on slender downy peduncles. Staminate in many-flowered heads; pistillate in two to several flowered clusters.
  • Calyx: Cup-shaped, five-toothed.
  • Corolla: Petals five, imbricate in bud, yellow green, ovate, thick, slightly spreading, inserted on the margin of the conspicuous disk.
  • Stamens: Five to twelve. In staminate flowers exserted, in pistillate short, often wanting.
  • Pistil: Ovary inferior, one to two-celled; style stout, exserted, reflexed above the middle. Entirely wanting in sterile flower. Ovules, one in each cell.
  • Fruit: Fleshy drupe, one to three from each flower cluster. Ovoid, two-thirds of an inch long, dark blue, acid. Stone more or less ridged. October.

Distribution

Nyssa sylvatica grows in various uplands and in alluvial stream bottoms from southwestern Maine
Maine
Maine is a state in the New England region of the northeastern United States, bordered by the Atlantic Ocean to the east and south, New Hampshire to the west, and the Canadian provinces of Quebec to the northwest and New Brunswick to the northeast. Maine is both the northernmost and easternmost...

 and New York
New York
New York is a state in the Northeastern region of the United States. It is the nation's third most populous state. New York is bordered by New Jersey and Pennsylvania to the south, and by Connecticut, Massachusetts and Vermont to the east...

, to extreme southern Ontario
Ontario
Ontario is a province of Canada, located in east-central Canada. It is Canada's most populous province and second largest in total area. It is home to the nation's most populous city, Toronto, and the nation's capital, Ottawa....

, central Michigan
Michigan
Michigan is a U.S. state located in the Great Lakes Region of the United States of America. The name Michigan is the French form of the Ojibwa word mishigamaa, meaning "large water" or "large lake"....

, Illinois
Illinois
Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

, and central Missouri
Missouri
Missouri is a US state located in the Midwestern United States, bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska. With a 2010 population of 5,988,927, Missouri is the 18th most populous state in the nation and the fifth most populous in the Midwest. It...

, south to southern Florida, eastern Texas
Texas
Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

, and eastern Oklahoma
Oklahoma
Oklahoma is a state located in the South Central region of the United States of America. With an estimated 3,751,351 residents as of the 2010 census and a land area of 68,667 square miles , Oklahoma is the 28th most populous and 20th-largest state...

. It also occurs locally in central and southern Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

. Optimum development is made on lower slopes and terraces in the Southeastern United States
Southeastern United States
The Southeastern United States, colloquially referred to as the Southeast, is the eastern portion of the Southern United States. It is one of the most populous regions in the United States of America....

.

Ecology

Nyssa sylvatica is found in a variety of upland
Upland
Upland may refer to:*Highland , an area of higher land*Upland Place namesIn Germany:*Upland - mountain range in the state of Hesse.In Sweden:*Uppland, a province of Sweden In the United States:...

 and wetland
Wetland
A wetland is an area of land whose soil is saturated with water either permanently or seasonally. Wetlands are categorised by their characteristic vegetation, which is adapted to these unique soil conditions....

 habitat
Habitat
* Habitat , a place where a species lives and grows*Human habitat, a place where humans live, work or play** Space habitat, a space station intended as a permanent settlement...

s in its extensive range. Its flowers are an important source of honey
Honey
Honey is a sweet food made by bees using nectar from flowers. The variety produced by honey bees is the one most commonly referred to and is the type of honey collected by beekeepers and consumed by humans...

 and its fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

s are important to many bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 species. Hollow trunks provide nesting or denning opportunities for bee
Bee
Bees are flying insects closely related to wasps and ants, and are known for their role in pollination and for producing honey and beeswax. Bees are a monophyletic lineage within the superfamily Apoidea, presently classified by the unranked taxon name Anthophila...

s and various mammals. It is longest living non-clonal flowering plant in Eastern North America, capable of obtaining ages of over 650 years.

Habitats

Nyssa sylvatica is found in a wide range of climates, due to its extensive distribution. It commonly grows in both the creek bottoms of the southern coastal plains to altitudes of about 900 meters (3000 ft) in the Southern Appalachians
Appalachian Mountains
The Appalachian Mountains #Whether the stressed vowel is or ,#Whether the "ch" is pronounced as a fricative or an affricate , and#Whether the final vowel is the monophthong or the diphthong .), often called the Appalachians, are a system of mountains in eastern North America. The Appalachians...

. These trees grow best on well-drained, light-textured soils on the low ridges of second bottoms and on the high flats of silty alluvium. In the uplands it grows best on the loams and clay loams of lower slopes and coves.

The species occurs 35 different forest cover types. When found on drier upper slopes and ridges, it is seldom of log size or quality.

Wildlife

Nyssa sylvatica is an important food source for many migrating birds in the fall. Its early color change (foliar fruit flagging) is thought to attract birds to the available fruit, which ripen before many other fall fruits and berries. The fruit is quite marked, dark blue, in clusters of two or three. The sour fruits are eagerly sought by many kinds of birds, including: American Robin
American Robin
The American Robin or North American Robin is a migratory songbird of the thrush family. It is named after the European Robin because of its reddish-orange breast, though the two species are not closely related, with the European robin belonging to the flycatcher family...

, Swainson's Thrush
Swainson's Thrush
Swainson's Thrush , also called Olive-backed Thrush, is a medium-sized thrush. This species is 16–18 cm in length, and has the white-dark-white underwing pattern characteristic of Catharus thrushes...

, Gray-cheeked Thrush
Gray-cheeked Thrush
The Grey-cheeked Thrush, Catharus minimus, is a medium-sized thrush. This species is 15–17 cm in length, and has the white-dark-white underwing pattern characteristic of Catharus thrushes. It is a member of a close-knit group of migrant species together with the Veery and Bicknell's Thrush ;...

, Hermit Thrush
Hermit Thrush
The Hermit Thrush is a medium-sized North American thrush. It is not very closely related to the other North American migrant species of Catharus, but rather to the Mexican Russet Nightingale-thrush.-Description:...

, Wood Thrush
Wood Thrush
The Wood Thrush, Hylocichla mustelina, is a North American passerine bird. It is closely related to other thrushes such as the American Robin and is widely distributed across North America, wintering in Central America and southern Mexico...

, Northern Cardinal
Northern Cardinal
The Northern Cardinal or Redbird or Common Cardinal is a North American bird in the genus Cardinalis. It can be found in southern Canada, through the eastern United States from Maine to Texas and south through Mexico...

, Northern Mockingbird
Northern Mockingbird
The Northern Mockingbird, Mimus polyglottos, is the only mockingbird commonly found in North America. This species was first described by Linnaeus in his Systema naturae in 1758 as Turdus polyglottos....

, Blue Jay
Blue Jay
The Blue Jay is a passerine bird in the family Corvidae, native to North America. It is resident through most of eastern and central United States and southern Canada, although western populations may be migratory. It breeds in both deciduous and coniferous forests, and is common near and in...

, Red-bellied Woodpecker
Red-bellied Woodpecker
The Red-bellied Woodpecker, Melanerpes carolinus, is a medium-sized woodpecker of the Picidae family. It breeds in southern Canada and the northeastern United States, ranging as far south as Florida and as far west as Texas...

, Yellow-bellied Sapsucker
Yellow-bellied Sapsucker
The Yellow-bellied Sapsucker is a medium-sized woodpecker found in North America, Central America and the Caribbean.-Taxonomy:...

, Northern Flicker
Northern Flicker
The Northern Flicker is a medium-sized member of the woodpecker family. It is native to most of North America, parts of Central America, Cuba, the Cayman Islands, and is one of the few woodpecker species that migrate. There are over 100 common names for the Northern Flicker...

, Pileated Woodpecker
Pileated Woodpecker
The Pileated Woodpecker is a very large North American woodpecker, almost crow-sized, inhabiting deciduous forests in eastern North America, the Great Lakes, the boreal forests of Canada, and parts of the Pacific coast. It is also the largest woodpecker in America.Adults are long, and weigh...

, Eastern Phoebe
Eastern Phoebe
The Eastern Phoebe is a small passerine bird. This tyrant flycatcher breeds in eastern North America, although its normal range does not include the southeastern coastal USA....

, Brown Thrasher
Brown Thrasher
The Brown Thrasher , sometimes erroneously called the Brown Thrush, is a bird in the Mimidae family, a group that also includes the New World catbirds and mockingbirds.-Description:...

, Eastern Bluebird
Eastern Bluebird
The Eastern Bluebird, Sialia sialis, is a small thrush found in open woodlands, farmlands and orchards, and most recently can be spotted in suburban areas. It is the state bird of Missouri and New York....

, European Starling
European Starling
The Common Starling , also known as the European Starling or just Starling, is a passerine bird in the family Sturnidae.This species of starling is native to most of temperate Europe and western Asia...

, Scarlet Tanager
Scarlet Tanager
The Scarlet Tanager is a medium-sized American songbird. Formerly placed in the tanager family , it and other members of its genus are now classified in the cardinal family . The species's plumage and vocalizations are similar to other members of the cardinal family.-Description:Adults have pale...

, Gray Catbird
Gray Catbird
The Gray Catbird , also spelled Grey Catbird, is a medium-sized northern American perching bird of the mimid family. It is the only member of the "catbird" genus Dumetella...

, Cedar Waxwing
Cedar Waxwing
The Cedar Waxwing is a member of the family Bombycillidae or waxwing family of passerine birds. It breeds in open wooded areas in North America, principally southern Canada and the northern United States.-Description:...

, and American Crow
American Crow
The American Crow is a large passerine bird species of the family Corvidae. It is a common bird found throughout much of North America...

, all primarily eastern North American birds migrating or residing year-round within the tree's range.

The limbs of these trees often deteriorate early, and the decayed holes make excellent dens for squirrel
Squirrel
Squirrels belong to a large family of small or medium-sized rodents called the Sciuridae. The family includes tree squirrels, ground squirrels, chipmunks, marmots , flying squirrels, and prairie dogs. Squirrels are indigenous to the Americas, Eurasia, and Africa and have been introduced to Australia...

s, raccoon
Raccoon
Procyon is a genus of nocturnal mammals, comprising three species commonly known as raccoons, in the family Procyonidae. The most familiar species, the common raccoon , is often known simply as "the" raccoon, as the two other raccoon species in the genus are native only to the tropics and are...

s, Virginia Opossum
Virginia Opossum
The Virginia opossum , commonly known as the North American opossum or tlacuache in Mexico, is the only marsupial found in North America north of Mexico. A solitary and nocturnal animal about the size of a domestic cat, and thus the largest opossum, it is a successful opportunist...

s, as well as nesting sites for honeybees.

Uses

Cultivation

Nyssa sylvatica is cultivated as an ornamental tree in parks and large gardens, where it is often used as a specimen or shade tree. The tree is best when grown in sheltered but not crowded positions, developing a pyramidal shape in youth, and spreading with age. The stem rises to the summit of the tree in one tapering unbroken shaft, the branches come out at right angles to the trunk and either extend horizontally or droop a little, making a long-narrow, cone-like head.

The leaves are short-petioled and so have little individual motion, but the branches sway as a whole. The spray is fine and abundant and lies horizontally so that the foliage arrangement is not unlike that of the beech
Beech
Beech is a genus of ten species of deciduous trees in the family Fagaceae, native to temperate Europe, Asia and North America.-Habit:...

 (Fagus). Its often spectacular autumnal coloring, with intense reds to purples, is highly valued in landscape settings. It is the most fiery and brilliant of the 'brilliant group' that includes maple
Acer
Acer may refer to:* Acer , the genus of trees and shrubs commonly known as Maple, from the Latin word ācer meaning sharp* Acer Inc., a Taiwan-based international computer company...

, dogwood, sassafras
Sassafras
Sassafras is a genus of three extant and one extinct species of deciduous trees in the family Lauraceae, native to eastern North America and eastern Asia.-Overview:...

, and sweet gum, as well as various species of tupelo
Nyssa
Nyssa may refer to:* the genus name for the tupelo tree* Nyssa , a female name** Nyssa , a companion of the Fourth and Fifth Doctors in the long-running British science fiction series Doctor Who...

.

Honey production

Nyssa sylvatica is a major source of wild honey in many areas within its range. Hollow sections of black tupelo trunks were formerly used as bee gums
Beehive (beekeeping)
A beehive is an enclosed structure in which some honey bee species of the subgenus Apis live and raise their young. Natural beehives are naturally occurring structures occupied by honeybee colonies, while domesticated honeybees live in man-made beehives, often in an apiary. These man-made...

 by beekeeper
Beekeeper
A beekeeper is a person who keeps honey bees for the purposes of securing commodities such as honey, beeswax, pollen, royal jelly; pollinating fruits and vegetables; raising queens and bees for sale to other farmers; and/or for purposes satisfying natural scientific curiosity...

s.

Wood

The wood of Nyssa sylvatica is heavy, hard, cross-grained, and difficult to split, especially after drying. This resistance to splitting led to its use for making maul
Maul
A splitting maul also known as a block buster, or block splitter is a heavy, long-handled hammer used for splitting a piece of wood along its grain. One side of its head is like a sledgehammer, and the other side is like an axe.- Wedged mauls :...

s, pulley
Pulley
A pulley, also called a sheave or a drum, is a mechanism composed of a wheel on an axle or shaft that may have a groove between two flanges around its circumference. A rope, cable, belt, or chain usually runs over the wheel and inside the groove, if present...

s, wheel hub
Wheel
A wheel is a device that allows heavy objects to be moved easily through rotating on an axle through its center, facilitating movement or transportation while supporting a load, or performing labor in machines. Common examples found in transport applications. A wheel, together with an axle,...

s, agricultural roller
Roller (agricultural tool)
The roller is an agricultural tool used for flattening land or breaking up large clumps of soil, especially after ploughing. Typically, rollers are pulled by tractors or, prior to mechanisation, a team of animals such as horses or oxen....

s, bowl
Bowl (vessel)
A bowl is a common open-top container used in many cultures to serve food, and is also used for drinking and storing other items. They are typically small and shallow, although some, such as punch bowls and salad bowls, are larger and often intended to serve many people.Bowls have existed for...

s, and paving blocks. The wood is also used for pallets, rough floors, pulpwood, and firewood.

External links

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