Bulimulidae
Encyclopedia
Bulimulinae is a taxonomic subfamily of medium-sized to large, air-breathing, tropical and sub-tropical land snail
Land snail
A land snail is any of the many species of snail that live on land, as opposed to those that live in salt water and fresh water. Land snails are terrestrial gastropod mollusks that have shells, It is not always an easy matter to say which species are terrestrial, because some are more or less...

s, terrestrial
Terrestrial animal
Terrestrial animals are animals that live predominantly or entirely on land , as compared with aquatic animals, which live predominantly or entirely in the water , or amphibians, which rely on a combination of aquatic and terrestrial habitats...

 pulmonate gastropod molluscs
Mollusca
The Mollusca , common name molluscs or mollusksSpelled mollusks in the USA, see reasons given in Rosenberg's ; for the spelling mollusc see the reasons given by , is a large phylum of invertebrate animals. There are around 85,000 recognized extant species of molluscs. Mollusca is the largest...

 in the family Orthalicidae
Orthalicidae
Orthalicidae are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the informal group Sigmurethra of the clade Stylommatophora...

, according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
The taxonomy of the Gastropoda as it was revised by Philippe Bouchet and Jean-Pierre Rocroi is currently the most up-to-date overall system for classifying gastropod mollusks...

.

Distribution

Distribution of Bulimulinae includes Ecuador (9 genera) and other South American countries.

Anatomy

Members of this subfamily have a haploid chromosome
Chromosome
A chromosome is an organized structure of DNA and protein found in cells. It is a single piece of coiled DNA containing many genes, regulatory elements and other nucleotide sequences. Chromosomes also contain DNA-bound proteins, which serve to package the DNA and control its functions.Chromosomes...

 number between 26 to 30 (according to the values in this table).

Taxonomy

Previously, the members of the Orthalicidae
Orthalicidae
Orthalicidae are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the informal group Sigmurethra of the clade Stylommatophora...

 were also included in this subfamily, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.

2005 taxonomy

The subfamily Bulimulinae includes 3 tribes:
  • Tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
  • Tribe Odontostomini Pilsbry
    Henry Augustus Pilsbry
    Henry Augustus Pilsbry was an American biologist, malacologist and carcinologist, among other areas of study. He was a dominant presence in many fields of invertebrate taxonomy for the better part of a century...

     & Vanatta, 1898 - synonym: Tomogeridae Jousseaume, 1877
  • Tribe Simpulopsini Schileyko, 1999

Genera

Genera in the subfamily Bulimulinae include:
  • tribe Bulimulini
    • Bulimulus
      Bulimulus
      Bulimulus is a genus of small to medium-sized tropical or sub-tropical, air-breathing land snails, pulmonate gastropod mollusks in the subfamily Bulimulinae.-Shell description:...

      Leach, 1814 - type genus
      Type genus
      In biological classification, a type genus is a representative genus, as with regard to a biological family. The term and concept is used much more often and much more formally in zoology than it is in botany, and the definition is dependent on the nomenclatural Code that applies:* In zoological...

       of the subfamily Bulimulinae

  • tribe Odontostomini
    • Odontostomus
      Odontostomus
      Odontostomus is a genus of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the tribe Odontostomini, in the subfamily Bulimulinae, within the family Orthalicidae.Odontostomus is the type genus for the tribe Odontostomini....

      Beck, 1837 - type genus of the tribe Odontostomini
    • Tomigerus
      Tomigerus
      Tomigerus is a genus of air-breathing tropical land snails, terrestrial pulmonate gastropod mollusks in the tribe Odontostomini within the subfamily Bulimulinae, in the family Orthalicidae.- Species :This genus contains the following species:...

      Spix, 1827
    • Anostoma
      Anostoma
      Anostoma, common name the up-mouth snails, is a genus of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Bulimulinae, in the family Orthalicidae...

      Fischer von Waldheim, 1807
    • Ringicella
      Ringicella
      Ringicella is a genus of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Bulimulinae, in the family Orthalicidae....

      Gray, 1847

  • tribe Simpulopsini
    • Simpulopsis Beck, 1837 - type genus of the tribe Simpulopsini

  • unsorted:
    • Auris Spix, 1827
    • Bostryx
      Bostryx
      Bostryx is a genus of air-breathing land snails, a terrestrial pulmonate gastropod mollusks in the family Orthalicidae.- Distribution :There are 29 species from the genus Bostryx in Chile and numerous species in other countries.- Species :...

      Troschel, 1847
    • Cochlorina Jan, 1830
    • Diplomorpha Ancey, 1884
    • Drymaeus
      Drymaeus
      Drymaeus is a genus of medium-sized air-breathing, tropical land snails, terrestrial pulmonate gastropod molluscs in the subfamily Bulimulinae and the family Orthalicidae, according to the taxonomy of the Gastropoda ....

      Albers, 1850
    • Dryptus Martens, 1860
    • Eudolichotis
      Eudolichotis
      Eudolichotis is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orthalicidae.- Species :Species in the genus Eudolichotis include:...

      Pilsbry, 1896
    • Naesiotus
      Naesiotus
      Naesiotus is a genus of tropical air-breathing land snails, a pulmonate gastropod mollusks in the family Orthalicidae.- Species :Species within the genus Naesiotus include:* Naesiotus quitensis -...

      Albers, 1850
    • Neopetraeus Martens, 1885
    • Newboldius Pilsbry, 1932
    • Otostomus Beck, 1837
    • Oxychona Moerch, 1852
    • Plectostylus Beck, 1837
    • Plekocheilus
      Plekocheilus
      Plekocheilus is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Orthalicidae.- Species :Species within the genus Plekocheilus include:subgenus Plekocheilus Guilding, 1828...

      Guilding, 1828
    • Rabdotus Albers, 1850
    • Scutalus
      Scutalus
      Scutalus is a genus of air-breathing land snails, a terrestrial pulmonate gastropod mollusks in the family Orthalicidae.- Species :Species in the genus Scutalus include:subgenus Scutalus Albers, 1850...

      Albers, 1850
    • Stenostylus Pilsbry, 1898
    • Tasmanembryon Iredale, 1933
    • Thaumastus
      Thaumastus (gastropod)
      Thaumastus is a genus of tropical air-breathing land snails, pulmonate gastropod mollusks in the family Orthalicidae.- Distribution :The distribution of the genus Thaumastus includes Colombia, Peru, Ecuador, ...- Species :...

      Albers, 1860 - synonym: Tholus Strebel, 1909
    • Xenothauma Fulton 1896

Odontostomini

Species in tribe Odontostomini have the aperture obstructed by internal lamellae, folds or teeth (rarely absent by degeneration); the base is perforate
or has an umbilical suture; and the genitalia are extremely lengthened. Jaw either plaited or solid.

Odontostomini is clearly a natural group of genera, confined to South America east of the Andes, and with the exception of some species, south of the Amazon. That the whole series had its inception in a form in which the characteristic apertural teeth had already been developed, is demonstrated by
the fact that these lamella and folds are clearly homologous throughout the species of the several genera. It follows from this that the toothless forms, such as Moricandia
Moricandia
Moricandia is a genus of plants belonging to the family Brassicaceae. Moricandia ramburii commonly known as violet cabbage, a native of the Mediterranean, is cultivated as a garden flower.-Species:* Moricandia arvensis DC....

, are secondarily so, by degeneration of the teeth of their ancestors. Many species show the various stages of tooth
degeneration.

Further reading

Neubert E. & Janssen R. (2004). "Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae". Archiv für Molluskenkunde 133: 193-297. abstract.
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