Carapace
Encyclopedia
A carapace is a dorsal
Dorsum (biology)
In anatomy, the dorsum is the upper side of animals that typically run, fly, or swim in a horizontal position, and the back side of animals that walk upright. In vertebrates the dorsum contains the backbone. The term dorsal refers to anatomical structures that are either situated toward or grow...

 (upper) section of the exoskeleton
Exoskeleton
An exoskeleton is the external skeleton that supports and protects an animal's body, in contrast to the internal skeleton of, for example, a human. In popular usage, some of the larger kinds of exoskeletons are known as "shells". Examples of exoskeleton animals include insects such as grasshoppers...

 or shell in a number of animal groups, including arthropod
Arthropod
An arthropod is an invertebrate animal having an exoskeleton , a segmented body, and jointed appendages. Arthropods are members of the phylum Arthropoda , and include the insects, arachnids, crustaceans, and others...

s such as crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s and arachnid
Arachnid
Arachnids are a class of joint-legged invertebrate animals in the subphylum Chelicerata. All arachnids have eight legs, although in some species the front pair may convert to a sensory function. The term is derived from the Greek words , meaning "spider".Almost all extant arachnids are terrestrial...

s, as well as vertebrates such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron.

Crustaceans

In crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s, the carapace is a part of the exoskeleton that covers the cephalothorax
Cephalothorax
The cephalothorax is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. The word cephalothorax is derived from the Greek words for head and thorax...

. It is particularly well developed in lobster
Lobster
Clawed lobsters comprise a family of large marine crustaceans. Highly prized as seafood, lobsters are economically important, and are often one of the most profitable commodities in coastal areas they populate.Though several groups of crustaceans are known as lobsters, the clawed lobsters are most...

s and crab
Crab
True crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" , or where the reduced abdomen is entirely hidden under the thorax...

s.

The carapace functions as a protective cover over the cephalothorax. Where it projects forward beyond the eyes, this projection is called a rostrum
Rostrum (anatomy)
The term rostrum is used for a number of unrelated structures in different groups of animals:*In crustaceans, the rostrum is the forward extension of the carapace in front of the eyes....

. The carapace is calcified to varying degrees in different crustaceans.

Zooplankton
Zooplankton
Zooplankton are heterotrophic plankton. Plankton are organisms drifting in oceans, seas, and bodies of fresh water. The word "zooplankton" is derived from the Greek zoon , meaning "animal", and , meaning "wanderer" or "drifter"...

 within the phylum Crustacea also have a carapace. These include Cladocera
Cladocera
Cladocera is an order of small crustaceans commonly called water fleas. Around 620 species have been recognised so far, with many more undescribed. They are ubiquitous in inland aquatic habitats, but rare in the oceans. Most are long, with a down-turned head, and a carapace covering the apparently...

, Ostracods, and Isopods, however Isopods only have a developed "cephalic shield" carapace covering the head.

Arachnida

In arachnid
Arachnid
Arachnids are a class of joint-legged invertebrate animals in the subphylum Chelicerata. All arachnids have eight legs, although in some species the front pair may convert to a sensory function. The term is derived from the Greek words , meaning "spider".Almost all extant arachnids are terrestrial...

s, the carapace is formed by the fusion of prosomal tergites into a single plate which carries the eyes, ocularium, ozopore
Ozopore
An ozopore is a defensive gland present in harvestmen, which are eight-legged arachnids also known as "daddy long-legs". These glands, also called "scent glands", "repugnatorial glands", "odoriferous glands" or "stink glands" by various authors, are located at the anterior sides of the prosoma.The...

s (a pair of openings of the scent gland of Opiliones
Opiliones
Opiliones are an order of arachnids commonly known as harvestmen. , over 6,400 species of harvestmen have been discovered worldwide, although the real number of extant species may exceed 10,000. The order Opiliones can be divided into four suborders: Cyphophthalmi, Eupnoi, Dyspnoi and Laniatores...

) and diverse phaneres.

In a few orders, such as Solifugae
Solifugae
Solifugae are an order of Arachnida, known as camel spiders, wind scorpions and sun spiders or solifuges, comprising more than 1,000 described species in about 153 genera...

 and Schizomida, the carapace may be subdivided. In Opiliones, some authors prefer to use the term carapace interchangeably with the term cephalothorax, which is incorrect usage, because carapace refers only to the dorsal part of the exoskeleton of the cephalothorax.

An alternative term for the carapace of arachnids and their relatives, which avoids confusion with crustaceans, is prosomal dorsal shield.

Turtles and tortoises

The carapace is the dorsal
Dorsum (biology)
In anatomy, the dorsum is the upper side of animals that typically run, fly, or swim in a horizontal position, and the back side of animals that walk upright. In vertebrates the dorsum contains the backbone. The term dorsal refers to anatomical structures that are either situated toward or grow...

 (back) convex part of the shell structure of a turtle
Turtle
Turtles are reptiles of the order Testudines , characterised by a special bony or cartilaginous shell developed from their ribs that acts as a shield...

, consisting primarily of the animal's rib
Rib
In vertebrate anatomy, ribs are the long curved bones which form the rib cage. In most vertebrates, ribs surround the chest, enabling the lungs to expand and thus facilitate breathing by expanding the chest cavity. They serve to protect the lungs, heart, and other internal organs of the thorax...

cage, dermal armor and scutes. See the main article on the turtle shell
Turtle shell
The turtle shell is a highly complicated shield for the ventral and dorsal parts of the turtle, completely enclosing all the vital organs of the turtle and in some cases even the head. It is constructed of modified bony elements such as the ribs, parts of the pelvis and other bones found in most...

for detailed information.
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