Cetacea
Encyclopedia
The order
Cetacea (icon) includes the marine mammals commonly known as whale
s, dolphin
s, and porpoise
s. Cetus is Latin
and is used in biological names to mean "whale"; its original meaning, "large sea animal", was more general. It comes from Ancient Greek
(), meaning "whale" or "any huge fish
or sea monster
". In Greek mythology
the monster Perseus
defeated was called Ceto
, which is depicted by the constellation
of Cetus
. Cetology
is the branch of marine science associated with the study of cetaceans.
Cetaceans are the mammal
s best adapted to aquatic life
. Their body is fusiform (spindle-shaped). The forelimbs are modified into flippers. The tiny hindlimbs are vestigial; they do not attach to the backbone and are hidden within the body. The tail has horizontal flukes. Cetaceans are nearly hairless, and are insulated from the cooler water they inhabit by a thick layer of blubber
. Some species are noted for their high intelligence
.
and inhale a fresh supply of oxygen
. During diving, a muscular action closes the blowholes
(nostril
s), which remain closed until the cetacean next breaks the surface; when it surfaces, the muscles open the blowholes and warm air is exhaled.
Cetaceans' blowholes have evolved
to a position at the top of the head, simplifying breathing in sometimes rough seas. When the stale air, warmed from the lungs, is exhaled, it condenses as it meets colder external air. As with a terrestrial mammal breathing out on a cold day, a small cloud of 'steam' appears. This is called the 'blow' or 'spout' and varies by species in terms of shape, angle and height. Species can be identified at a distance using this characteristic.
Cetaceans can remain under water for much longer periods than most other mammals, (approximately 7–30 minutes, varying by species) due to large physiological differences. Two studied advantages of cetacean physiology let this order (and other marine mammals) forage underwater for extended periods without breathing:
As with the eyes, cetacean ears
are also small. Life in the sea accounts for the cetacean's loss of its external ears, whose function is to collect and focus airborne sound waves. However, water conducts sound better than air, so the external ear is unneeded: it is a tiny hole in the skin, just behind the eye. The highly developed inner ear can detect sounds from dozens of miles away and discern from which direction the sound comes.
Odontoceti are generally capable of echolocation
. From this, Odontoceti can discern the size, shape, surface characteristics, distance and movement of an object. With this ability cetaceans can search for, chase and catch fast-swimming prey in total darkness. Echolocation is so advanced in most Odontoceti that they can distinguish between prey and non-prey (such as humans or boats); captive Odontoceti can be trained to distinguish between, for example, balls of different sizes or shapes. Mysticeti have exceptionally thin, wide basilar membranes in their cochleae without stiffening agents - making their ears adapted for processing low to infrasonic frequencies.
Cetaceans also use sound to communicate
, whether it be groans, moans, whistles, clicks or the complex 'singing' of the humpback whale
.
Besides hearing and vision, at least one species, the Tucuxi
or Guiana Dolphin, is able to use electroreception to sense prey.
, squid
or other marine life. They do not chew but swallow prey whole. When they catch large prey, such as when the orca
(Orcinus orca) catches a seal
, they bite off and swallow one chunk at a time.
Mysticeti instead have baleen
plates made of keratin
(the same substance as human fingernails) which hang from the upper jaw. These plates filter small animals (such as krill
and fish) from the seawater. Cetaceans included in this group include the Blue, Humpback, Bowhead and Minke whales.
Not all Mysticeti feed on plankton
: the larger species eat small shoaling fish, such as herring
and sardine
, called micronecton. The gray whale
(Eschrichtius robustus), is a benthic
feeder, primarily eating sea floor crustacean
s.
Mammalia. Their closest living relatives are the even-toed ungulate
s, such as the hippopotamus
and deer
.
Mammalian characteristics include warm-blooded
ness, breathing air through their lungs, and suckling
their young, and growing hair
, although very little of it.
Another way of distinguishing a cetacean from a fish is the shape of the tail. Fish tails are vertical and move from side to side when the fish swims. Cetacea tails—called a fluke—are horizontal and move up and down, because cetacea spines bend in the same manner as a human
spine.
. The order contains two suborders, Mysticeti (baleen whales) and Odontoceti (toothed whales, which includes dolphins and porpoises). The species range in size from Commerson's dolphin
, smaller than a human, to the Blue Whale
, the largest animal ever known to have lived.
s, dolphin
s and porpoise
s) are marine mammal
descendants of the artiodactyl family Raoellidae, a group of land mammals characterized by an even-toed ungulate skull, slim limbs, and an ear with significant similarities to that of early whales. The terrestrial origins of cetaceans are indicated by their need to breathe air from the surface or the bones of their fins, which resemble the limbs of land mammals, including the vestigial remains of hind legs inherited from the four-legged ancestors.
The question of how land animals evolved into ocean-going leviathan
s was a mystery until discoveries starting in the late 1970s in Pakistan
revealed several stages in the transition of cetaceans from land to sea:
This series of images does not capture the true phylogenetic evolution
of a particular species, but they are illustrative representation of the evolution of cetaceans from terrestrial four-legged mammals, from their probable ancestor, through different stages of adaptation to aquatic life to modern cetaceans type; aerodynamic body shape, fully developed caudal fin and vestigial hind legs. The separation of cetaceans in suborder baleen whales and suborder toothed whales, occurred during the Oligocene
(Janjucetus and Squalodon represent the early forms of their suborders).
The classification here closely follows Dale W. Rice, Marine Mammals of the World: Systematics and Distribution (1998), which has become the standard taxonomy reference in the field. There is very close agreement between this classification and that of Mammal Species of the World: 3rd Edition (Wilson and Reeder eds., 2005). Any differences are noted using the abbreviations "Rice" and "MSW3" respectively. Further differences due to recent discoveries are also noted.
Discussion of synonyms and subspecies are relegated to the relevant genus and species articles.
†Recently extinct
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...
Cetacea (icon) includes the marine mammals commonly known as whale
Whale
Whale is the common name for various marine mammals of the order Cetacea. The term whale sometimes refers to all cetaceans, but more often it excludes dolphins and porpoises, which belong to suborder Odontoceti . This suborder also includes the sperm whale, killer whale, pilot whale, and beluga...
s, dolphin
Dolphin
Dolphins are marine mammals that are closely related to whales and porpoises. There are almost forty species of dolphin in 17 genera. They vary in size from and , up to and . They are found worldwide, mostly in the shallower seas of the continental shelves, and are carnivores, mostly eating...
s, and porpoise
Porpoise
Porpoises are small cetaceans of the family Phocoenidae; they are related to whales and dolphins. They are distinct from dolphins, although the word "porpoise" has been used to refer to any small dolphin, especially by sailors and fishermen...
s. Cetus is Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...
and is used in biological names to mean "whale"; its original meaning, "large sea animal", was more general. It comes from Ancient Greek
Ancient Greek
Ancient Greek is the stage of the Greek language in the periods spanning the times c. 9th–6th centuries BC, , c. 5th–4th centuries BC , and the c. 3rd century BC – 6th century AD of ancient Greece and the ancient world; being predated in the 2nd millennium BC by Mycenaean Greek...
(), meaning "whale" or "any huge fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...
or sea monster
Sea monster
Sea monsters are sea-dwelling mythical or legendary creatures, often believed to be of immense size.Marine monsters can take many forms, including sea dragons, sea serpents, or multi-armed beasts. They can be slimy or scaly and are often pictured threatening ships or spouting jets of water...
". In Greek mythology
Greek mythology
Greek mythology is the body of myths and legends belonging to the ancient Greeks, concerning their gods and heroes, the nature of the world, and the origins and significance of their own cult and ritual practices. They were a part of religion in ancient Greece...
the monster Perseus
Perseus
Perseus ,Perseos and Perseas are not used in English. the legendary founder of Mycenae and of the Perseid dynasty of Danaans there, was the first of the mythic heroes of Greek mythology whose exploits in defeating various archaic monsters provided the founding myths of the Twelve Olympians...
defeated was called Ceto
Ceto
In ancient Greek, the word ketos - Latinized as cetus - denotes a large fish, a whale, a shark, or a sea monster. The sea monsters slain by Perseus and Heracles were each referred to as a cetus by ancient sources. The term cetacean originates from cetus. In Greek art, cetea were depicted as...
, which is depicted by the constellation
Constellation
In modern astronomy, a constellation is an internationally defined area of the celestial sphere. These areas are grouped around asterisms, patterns formed by prominent stars within apparent proximity to one another on Earth's night sky....
of Cetus
Cetus
Cetus is a constellation. Its name refers to Cetus, a sea monster in Greek mythology, although it is often called 'the whale' today. Cetus is located in the region of the sky that contains other water-related constellations such as Aquarius, Pisces, and Eridanus.-Ecliptic:Although Cetus is not...
. Cetology
Cetology
Cetology is the branch of marine mammal science that studies the approximately eighty species of whales, dolphins, and porpoise in the scientific order Cetacea...
is the branch of marine science associated with the study of cetaceans.
Cetaceans are the mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...
s best adapted to aquatic life
Aquatic adaptation
Several animal groups have undergone aquatic adaptation, going from being purely terrestrial animals to living at least part of the time in water. The adaptations in early speciation tend to develop as the animal ventures into water in order to find available food. As successive generations spend...
. Their body is fusiform (spindle-shaped). The forelimbs are modified into flippers. The tiny hindlimbs are vestigial; they do not attach to the backbone and are hidden within the body. The tail has horizontal flukes. Cetaceans are nearly hairless, and are insulated from the cooler water they inhabit by a thick layer of blubber
Blubber
Blubber is a thick layer of vascularized adipose tissue found under the skin of all cetaceans, pinnipeds and sirenians.-Description:Lipid-rich, collagen fiber–laced blubber comprises the hypodermis and covers the whole body, except for parts of the appendages, strongly attached to the musculature...
. Some species are noted for their high intelligence
Cetacean intelligence
Cetacean intelligence denotes the cognitive capabilities of the Cetacea order of mammals, which includes whales, porpoises, and dolphins.-Brain size:...
.
Respiration
Cetaceans breathe air. They surface periodically to exhale carbon dioxideCarbon dioxide
Carbon dioxide is a naturally occurring chemical compound composed of two oxygen atoms covalently bonded to a single carbon atom...
and inhale a fresh supply of oxygen
Oxygen
Oxygen is the element with atomic number 8 and represented by the symbol O. Its name derives from the Greek roots ὀξύς and -γενής , because at the time of naming, it was mistakenly thought that all acids required oxygen in their composition...
. During diving, a muscular action closes the blowholes
Blowhole (biology)
In biology, a blowhole is the hole at the top of a cetacean's head through which the animal breathes air. It is homologous with the nostril of other mammals. As whales reach the water surface to breathe, they will forcefully expel air through the blowhole. Not only is air expelled, but mucus and...
(nostril
Nostril
A nostril is one of the two channels of the nose, from the point where they bifurcate to the external opening. In birds and mammals, they contain branched bones or cartilages called turbinates, whose function is to warm air on inhalation and remove moisture on exhalation...
s), which remain closed until the cetacean next breaks the surface; when it surfaces, the muscles open the blowholes and warm air is exhaled.
Cetaceans' blowholes have evolved
Evolution of cetaceans
The cetaceans are marine mammal descendants of land mammals. Their terrestrial origins are indicated by:* Their need to breathe air from the surface;* The bones of their fins, which resemble the limbs of land mammals...
to a position at the top of the head, simplifying breathing in sometimes rough seas. When the stale air, warmed from the lungs, is exhaled, it condenses as it meets colder external air. As with a terrestrial mammal breathing out on a cold day, a small cloud of 'steam' appears. This is called the 'blow' or 'spout' and varies by species in terms of shape, angle and height. Species can be identified at a distance using this characteristic.
Cetaceans can remain under water for much longer periods than most other mammals, (approximately 7–30 minutes, varying by species) due to large physiological differences. Two studied advantages of cetacean physiology let this order (and other marine mammals) forage underwater for extended periods without breathing:
- Mammalian myoglobinMyoglobinMyoglobin is an iron- and oxygen-binding protein found in the muscle tissue of vertebrates in general and in almost all mammals. It is related to hemoglobin, which is the iron- and oxygen-binding protein in blood, specifically in the red blood cells. The only time myoglobin is found in the...
concentrations in skeletalSkeletonThe skeleton is the body part that forms the supporting structure of an organism. There are two different skeletal types: the exoskeleton, which is the stable outer shell of an organism, and the endoskeleton, which forms the support structure inside the body.In a figurative sense, skeleton can...
muscleMuscleMuscle is a contractile tissue of animals and is derived from the mesodermal layer of embryonic germ cells. Muscle cells contain contractile filaments that move past each other and change the size of the cell. They are classified as skeletal, cardiac, or smooth muscles. Their function is to...
have much variation. New ZealandNew ZealandNew Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...
white rabbits have 0.08 gram (0.00282191696840896 oz) +/- 0.6 gram (0.0211643772630672 oz) myoglobin in 100 grams (3.5 oz) of wet muscle, whereas a northern bottlenose whale has 6.34 gram (0.22363691974641 oz). Myoglobin, by nature, has a higher oxygen affinity than hemoglobinHemoglobinHemoglobin is the iron-containing oxygen-transport metalloprotein in the red blood cells of all vertebrates, with the exception of the fish family Channichthyidae, as well as the tissues of some invertebrates...
. The higher the myoglobin concentration in skeletal muscle, the longer the animal can stay underwater.
- Increased body size also increases maximum dive duration. Greater body size implies increased muscle mass and increased oxygen stores. Cetaceans also obey Kleiber's lawKleiber's lawKleiber's law, named after Max Kleiber's biological work in the early 1930s, is the observation that, for the vast majority of animals, an animal's metabolic rate scales to the ¾ power of the animal's mass. Symbolically: if q0 is the animal's metabolic rate, and M the animal's mass, then Kleiber's...
, which states that mass and metabolic rate are inversely related. I.e., larger animals consume less oxygen than smaller animals per unit mass.
Vision, hearing and echolocation
Cetacean eyes are set on the side rather than the front of the head. This means that only cetaceans with pointed 'beaks' (such as dolphins) have good binocular vision forward and downward. Tear glands secrete greasy tears, which protect the eyes from the salt in the water. The lens is almost spherical, which is most efficient at focusing the minimal light that reaches deep water. Cetaceans make up for their generally poor vision (with the exception of the dolphin) with excellent hearing.As with the eyes, cetacean ears
EARS
EARS may refer to:* Electoral software* Electronic Arts, Redwood Shores campus.* Emirates Amateur Radio Society...
are also small. Life in the sea accounts for the cetacean's loss of its external ears, whose function is to collect and focus airborne sound waves. However, water conducts sound better than air, so the external ear is unneeded: it is a tiny hole in the skin, just behind the eye. The highly developed inner ear can detect sounds from dozens of miles away and discern from which direction the sound comes.
Odontoceti are generally capable of echolocation
Animal echolocation
Echolocation, also called biosonar, is the biological sonar used by several kinds of animals.Echolocating animals emit calls out to the environment and listen to the echoes of those calls that return from various objects near them. They use these echoes to locate and identify the objects...
. From this, Odontoceti can discern the size, shape, surface characteristics, distance and movement of an object. With this ability cetaceans can search for, chase and catch fast-swimming prey in total darkness. Echolocation is so advanced in most Odontoceti that they can distinguish between prey and non-prey (such as humans or boats); captive Odontoceti can be trained to distinguish between, for example, balls of different sizes or shapes. Mysticeti have exceptionally thin, wide basilar membranes in their cochleae without stiffening agents - making their ears adapted for processing low to infrasonic frequencies.
Cetaceans also use sound to communicate
Communication
Communication is the activity of conveying meaningful information. Communication requires a sender, a message, and an intended recipient, although the receiver need not be present or aware of the sender's intent to communicate at the time of communication; thus communication can occur across vast...
, whether it be groans, moans, whistles, clicks or the complex 'singing' of the humpback whale
Humpback Whale
The humpback whale is a species of baleen whale. One of the larger rorqual species, adults range in length from and weigh approximately . The humpback has a distinctive body shape, with unusually long pectoral fins and a knobbly head. It is an acrobatic animal, often breaching and slapping the...
.
Besides hearing and vision, at least one species, the Tucuxi
Tucuxi
The Tucuxi , alternately bufeo gris or bufeo negro is a dolphin found in the rivers of the Amazon Basin. The word "tucuxi" is derived from the Tupi language word tuchuchi-ana and has now been adopted as the species' common name...
or Guiana Dolphin, is able to use electroreception to sense prey.
Feeding
The toothed whales such as the sperm whale, beluga, dolphins and porpoises, have teeth that they use for catching fishFish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...
, squid
Squid
Squid are cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, bilateral symmetry, a mantle, and arms. Squid, like cuttlefish, have eight arms arranged in pairs and two, usually longer, tentacles...
or other marine life. They do not chew but swallow prey whole. When they catch large prey, such as when the orca
Orca
The killer whale , commonly referred to as the orca, and less commonly as the blackfish, is a toothed whale belonging to the oceanic dolphin family. Killer whales are found in all oceans, from the frigid Arctic and Antarctic regions to tropical seas...
(Orcinus orca) catches a seal
Pinniped
Pinnipeds or fin-footed mammals are a widely distributed and diverse group of semiaquatic marine mammals comprising the families Odobenidae , Otariidae , and Phocidae .-Overview: Pinnipeds are typically sleek-bodied and barrel-shaped...
, they bite off and swallow one chunk at a time.
Mysticeti instead have baleen
Baleen
Baleen or whalebone is a filter-feeder system inside the mouths of baleen whales. The baleen system works when a whale opens its mouth underwater and then water pours into the whale's mouth. The whale then pushes the water out, and animals such as krill are filtered by the baleen and remain as food...
plates made of keratin
Keratin
Keratin refers to a family of fibrous structural proteins. Keratin is the key of structural material making up the outer layer of human skin. It is also the key structural component of hair and nails...
(the same substance as human fingernails) which hang from the upper jaw. These plates filter small animals (such as krill
Krill
Krill is the common name given to the order Euphausiacea of shrimp-like marine crustaceans. Also known as euphausiids, these small invertebrates are found in all oceans of the world...
and fish) from the seawater. Cetaceans included in this group include the Blue, Humpback, Bowhead and Minke whales.
Not all Mysticeti feed on plankton
Plankton
Plankton are any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. That is, plankton are defined by their ecological niche rather than phylogenetic or taxonomic classification...
: the larger species eat small shoaling fish, such as herring
Herring
Herring is an oily fish of the genus Clupea, found in the shallow, temperate waters of the North Pacific and the North Atlantic oceans, including the Baltic Sea. Three species of Clupea are recognized. The main taxa, the Atlantic herring and the Pacific herring may each be divided into subspecies...
and sardine
Sardine
Sardines, or pilchards, are several types of small, oily fish related to herrings, family Clupeidae. Sardines are named after the Mediterranean island of Sardinia, around which they were once abundant....
, called micronecton. The gray whale
Gray Whale
The gray whale, Eschrichtius robustus, is a baleen whale that migrates between feeding and breeding grounds yearly. It reaches a length of about , a weight of , and lives 50–70 years. The common name of the whale comes from the gray patches and white mottling on its dark skin. Gray whales were...
(Eschrichtius robustus), is a benthic
Benthos
Benthos is the community of organisms which live on, in, or near the seabed, also known as the benthic zone. This community lives in or near marine sedimentary environments, from tidal pools along the foreshore, out to the continental shelf, and then down to the abyssal depths.Many organisms...
feeder, primarily eating sea floor crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...
s.
Mammalian nature
Cetaceans are mammals, that is, members of the classClass (biology)
In biological classification, class is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, order, family, genus, and species, with class fitting between phylum and order...
Mammalia. Their closest living relatives are the even-toed ungulate
Even-toed ungulate
The even-toed ungulates are ungulates whose weight is borne about equally by the third and fourth toes, rather than mostly or entirely by the third as in odd-toed ungulates such as horses....
s, such as the hippopotamus
Hippopotamus
The hippopotamus , or hippo, from the ancient Greek for "river horse" , is a large, mostly herbivorous mammal in sub-Saharan Africa, and one of only two extant species in the family Hippopotamidae After the elephant and rhinoceros, the hippopotamus is the third largest land mammal and the heaviest...
and deer
Deer
Deer are the ruminant mammals forming the family Cervidae. Species in the Cervidae family include white-tailed deer, elk, moose, red deer, reindeer, fallow deer, roe deer and chital. Male deer of all species and female reindeer grow and shed new antlers each year...
.
Mammalian characteristics include warm-blooded
Warm-blooded
The term warm-blooded is a colloquial term to describe animal species which have a relatively higher blood temperature, and maintain thermal homeostasis primarily through internal metabolic processes...
ness, breathing air through their lungs, and suckling
Breastfeeding
Breastfeeding is the feeding of an infant or young child with breast milk directly from female human breasts rather than from a baby bottle or other container. Babies have a sucking reflex that enables them to suck and swallow milk. It is recommended that mothers breastfeed for six months or...
their young, and growing hair
Hair
Hair is a filamentous biomaterial, that grows from follicles found in the dermis. Found exclusively in mammals, hair is one of the defining characteristics of the mammalian class....
, although very little of it.
Another way of distinguishing a cetacean from a fish is the shape of the tail. Fish tails are vertical and move from side to side when the fish swims. Cetacea tails—called a fluke—are horizontal and move up and down, because cetacea spines bend in the same manner as a human
Human
Humans are the only living species in the Homo genus...
spine.
Taxonomy
The order Cetacea contains about 90 species, all marine except for 4 species of freshwater dolphinsRiver dolphin
River dolphins are the four living species of dolphin that reside in freshwater rivers and estuaries. River dolphins inhabit areas of Asia and South America. They are classed in the Platanistoidea superfamily of cetaceans. Three species live in fresh water rivers. The fourth species, the La Plata...
. The order contains two suborders, Mysticeti (baleen whales) and Odontoceti (toothed whales, which includes dolphins and porpoises). The species range in size from Commerson's dolphin
Commerson's Dolphin
Commerson's Dolphin is one of four dolphins in the Cephalorhynchus genus. The species has also the common names Skunk Dolphin, Piebald Dolphin and Panda Dolphin...
, smaller than a human, to the Blue Whale
Blue Whale
The blue whale is a marine mammal belonging to the suborder of baleen whales . At in length and or more in weight, it is the largest known animal to have ever existed....
, the largest animal ever known to have lived.
Evolution
The cetaceans (whaleWhale
Whale is the common name for various marine mammals of the order Cetacea. The term whale sometimes refers to all cetaceans, but more often it excludes dolphins and porpoises, which belong to suborder Odontoceti . This suborder also includes the sperm whale, killer whale, pilot whale, and beluga...
s, dolphin
Dolphin
Dolphins are marine mammals that are closely related to whales and porpoises. There are almost forty species of dolphin in 17 genera. They vary in size from and , up to and . They are found worldwide, mostly in the shallower seas of the continental shelves, and are carnivores, mostly eating...
s and porpoise
Porpoise
Porpoises are small cetaceans of the family Phocoenidae; they are related to whales and dolphins. They are distinct from dolphins, although the word "porpoise" has been used to refer to any small dolphin, especially by sailors and fishermen...
s) are marine mammal
Marine mammal
Marine mammals, which include seals, whales, dolphins, and walruses, form a diverse group of 128 species that rely on the ocean for their existence. They do not represent a distinct biological grouping, but rather are unified by their reliance on the marine environment for feeding. The level of...
descendants of the artiodactyl family Raoellidae, a group of land mammals characterized by an even-toed ungulate skull, slim limbs, and an ear with significant similarities to that of early whales. The terrestrial origins of cetaceans are indicated by their need to breathe air from the surface or the bones of their fins, which resemble the limbs of land mammals, including the vestigial remains of hind legs inherited from the four-legged ancestors.
The question of how land animals evolved into ocean-going leviathan
Leviathan
Leviathan , is a sea monster referred to in the Bible. In Demonology, Leviathan is one of the seven princes of Hell and its gatekeeper . The word has become synonymous with any large sea monster or creature...
s was a mystery until discoveries starting in the late 1970s in Pakistan
Pakistan
Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...
revealed several stages in the transition of cetaceans from land to sea:
Pakicetus Pakicetus Pakicetus is a genus of extinct terrestrial carnivorous mammal of the family Pakicetidae which was endemic to Pakistan from the Eocene .Pakicetus existed for approximately... |
Ambulocetus Ambulocetus Ambulocetus was an early cetacean that could walk as well as swim. It lived during early Eocene some 50-49 million years ago. It is a transitional fossil that shows how whales evolved from land-living mammals. The Ambulocetus fossils were found in Pakistan by anthropologist Johannes Thewissen... |
Kutchicetus Kutchicetus Kutchicetus is an extinct genus of early carnivorous freshwater whales of the family Remingtonocetidae endemic to the coastline of the ancient Tethys Ocean during the Eocene living from 55.8—40.4 mya, existing for approximately .-Taxonomy:... |
Protocetus Protocetus Protocetus atavus is an extinct species of primitive cetacean from Egypt. It lived during the middle Eocene period 45 million years ago.... |
Janjucetus Janjucetus Janjucetus is an extinct genus of whale, and a basal form of the Mysticeti, a clade which includes the extant baleen whales. The only known species, Janjucetus hunderi, lived during the late Oligocene, about 25 million years ago in coastal seas off southeast Australia. Unlike modern mysticetes, it... |
Squalodon Squalodon Squalodon is an extinct genus of whales, belonging to the family Squalodontidae. Named by Grateloup in 1840, it was originally believed to be an iguanodontid dinosaur but has since been reclassified. The name Squalodon comes from Squalus, a genus of shark... |
This series of images does not capture the true phylogenetic evolution
Evolution
Evolution is any change across successive generations in the heritable characteristics of biological populations. Evolutionary processes give rise to diversity at every level of biological organisation, including species, individual organisms and molecules such as DNA and proteins.Life on Earth...
of a particular species, but they are illustrative representation of the evolution of cetaceans from terrestrial four-legged mammals, from their probable ancestor, through different stages of adaptation to aquatic life to modern cetaceans type; aerodynamic body shape, fully developed caudal fin and vestigial hind legs. The separation of cetaceans in suborder baleen whales and suborder toothed whales, occurred during the Oligocene
Oligocene
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 34 million to 23 million years before the present . As with other older geologic periods, the rock beds that define the period are well identified but the exact dates of the start and end of the period are slightly...
(Janjucetus and Squalodon represent the early forms of their suborders).
Mysticeti vs Odontoceti
Fossils indicate that before evolving baleen, Mysticeti also had teeth, so defining the Odontoceti via teeth alone is problematic. Instead, paleontologists have identified other features uniting fossil and modern odontocetes that are not shared by Mysticetes. It was also assumed that toothed whales evolved their assymetrical skulls as an adaptation to their echolocation, but newer discoveries indicate that the common ancestor of the present whales actually had a contorted skull as well. Cranial asymmetry is now known to have evolved in ancient whales as part of a set of traits linked to directional hearing, including pan-bone thinning of the lower jaws, the development of mandibular fat pads, and the isolation of the ear region . This likely means that while the asymmetry in the Odontoceti skull has increased over time, the Mysticeti skull has evolved from asymmetrical to a symmetrical.Characteristic | Odontoceti | Mysticeti |
Feeding | Echolocation Animal echolocation Echolocation, also called biosonar, is the biological sonar used by several kinds of animals.Echolocating animals emit calls out to the environment and listen to the echoes of those calls that return from various objects near them. They use these echoes to locate and identify the objects... , fast |
Filter feeder, not fast |
Size | |Smaller (except Sperm whale Sperm Whale The sperm whale, Physeter macrocephalus, is a marine mammal species, order Cetacea, a toothed whale having the largest brain of any animal. The name comes from the milky-white waxy substance, spermaceti, found in the animal's head. The sperm whale is the only living member of genus Physeter... and Beaked Whale Beaked whale Beaked whales are 21 species of toothed whales, members of the family Ziphiidae, are notable for their elongated beaks. Beaked whales are one of the world's most extreme divers. They can dive for long periods—20 to 30 minutes is common, and 85 minute dives have been recorded—and to... ) |
Larger (except pygmy right whale Pygmy Right Whale The pygmy right whale is a baleen whale, the sole member of the family Neobalaenidae. First described by John Edward Gray in 1846, it is the smallest of the baleen whales, ranging between and in length and 3,000 and 3,500 kg in mass... ) |
Blowhole | One | Two |
Dentition Dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, the characteristic arrangement, kind, and number of teeth in a given species at a given age... |
|Teeth | Baleen Baleen Baleen or whalebone is a filter-feeder system inside the mouths of baleen whales. The baleen system works when a whale opens its mouth underwater and then water pours into the whale's mouth. The whale then pushes the water out, and animals such as krill are filtered by the baleen and remain as food... plates |
Melon Melon (whale) The melon is an ovoid-shaped, fatty organ found in the forehead of all toothed whales , including dolphins and porpoises and believed to be used in echolocation.... |
Ovoid, in anterior facial region | Vestigial or none |
Skull Skull The skull is a bony structure in the head of many animals that supports the structures of the face and forms a cavity for the brain.The skull is composed of two parts: the cranium and the mandible. A skull without a mandible is only a cranium. Animals that have skulls are called craniates... and facial tissue |
Dorsally asymmetric | |Symmetric |
Sexual dimorphism Sexual dimorphism Sexual dimorphism is a phenotypic difference between males and females of the same species. Examples of such differences include differences in morphology, ornamentation, and behavior.-Examples:-Ornamentation / coloration:... |
Some species have larger males | Females always larger |
Mandible Mandible The mandible pronunciation or inferior maxillary bone forms the lower jaw and holds the lower teeth in place... |
Symphyseal (fused anteriorly) | Nonsymphyseal |
Pan bone of lower jaw | Yes | No |
Maxillae projection | Outward over expanded supraorbital processes | Under eye orbit, with bony protuberance anterior to eye orbit |
Olfactory nerve Olfactory nerve The olfactory nerve, or cranial nerve I, is the first of twelve cranial nerves. It is instrumental in the sense of smell. Derived from the embryonic nasal placode, the olfactory nerve is capable of regeneration.-Anatomy:... and bulb Olfactory bulb The olfactory bulb is a structure of the vertebrate forebrain involved in olfaction, the perception of odors.-Anatomy:In most vertebrates, the olfactory bulb is the most rostral part of the brain. In humans, however, the olfactory bulb is on the inferior side of the brain... |
Absent | Vestigal |
Periotic bone Periotic bone Periotic bone is the single bone that surrounds the ear of mammals. It is formed from the fusion of the pro-otic, epiotic, and opisthotic bones.- External links :* http://www.encyclopedia.com/doc/1O8-perioticbone.html... |
External to skull Skull The skull is a bony structure in the head of many animals that supports the structures of the face and forms a cavity for the brain.The skull is composed of two parts: the cranium and the mandible. A skull without a mandible is only a cranium. Animals that have skulls are called craniates... , fused with tympanic bulla |
Fused with skull Skull The skull is a bony structure in the head of many animals that supports the structures of the face and forms a cavity for the brain.The skull is composed of two parts: the cranium and the mandible. A skull without a mandible is only a cranium. Animals that have skulls are called craniates... |
Classification
The classification here closely follows Dale W. Rice, Marine Mammals of the World: Systematics and Distribution (1998), which has become the standard taxonomy reference in the field. There is very close agreement between this classification and that of Mammal Species of the World: 3rd Edition (Wilson and Reeder eds., 2005). Any differences are noted using the abbreviations "Rice" and "MSW3" respectively. Further differences due to recent discoveries are also noted.
Discussion of synonyms and subspecies are relegated to the relevant genus and species articles.
- ORDER CETACEA
- Suborder Mysticeti: Baleen whales
- Family BalaenidaeBalaenidaeBalaenidae is a family of mysticete whales that contains two living genera. Commonly called the right whales as it contains mainly right whale species...
: Right whaleRight whaleRight whales are three species of large baleen whales consisting of two genera in the family Balaenidae of order Cetacea. Their bodies are very dark gray or black and rotund....
s and bowhead whaleBowhead WhaleThe bowhead whale is a baleen whale of the right whale family Balaenidae in suborder Mysticeti. A stocky dark-colored whale without a dorsal fin, it can grow to in length. This thick-bodied species can weigh to , second only to the blue whale, although the bowhead's maximum length is less than...
- Genus BalaenaBalaenaBalaena is a genus of cetacean in the Balaenidae family. This family is also known as the Right Whales, which can cause confusion as there is also a species of whale called the Right Whale. The only extant species is the Bowhead whale. The fossil record of Balaena, dating to the late Miocene,...
- Bowhead whaleBowhead WhaleThe bowhead whale is a baleen whale of the right whale family Balaenidae in suborder Mysticeti. A stocky dark-colored whale without a dorsal fin, it can grow to in length. This thick-bodied species can weigh to , second only to the blue whale, although the bowhead's maximum length is less than...
, Balaena mysticetus
- Bowhead whale
- Genus Eubalaena
- North Atlantic right whaleNorth Atlantic Right WhaleThe North Atlantic right whale which means "good, or true, whale of the ice") is a baleen whale, one of three right whale species belonging to the genus Eubalaena, formerly classified as a single species. With only 400 in existence, North Atlantic right whales are among the most endangered whales...
, Eubalaena glacialis - North Pacific right whaleNorth Pacific Right WhaleThe North Pacific right whale is a very large, robust baleen whale species that is now extremely rare and endangered. The Northeast Pacific subpopulation, that summers in the southeastern Bering Sea and Gulf of Alaska, may have no more than 50 animals...
, Eubalaena japonica - Southern right whaleSouthern Right WhaleThe southern right whale is a baleen whale, one of three species classified as right whales belonging to the genus Eubalaena. Like other right whales, the southern right whale is readily distinguished from others by the callosities on its head, a broad back without a dorsal fin, and a long arching...
, Eubalaena australis
- North Atlantic right whale
- Genus Balaena
- Family Balaenopteridae: Rorquals
- Subfamily Balaenopterinae
- Genus BalaenopteraBalaenopteraBalaenoptera is a genus of Balaenopteridae, the Rorqual whales, and contains eight species. The species Balaenoptera omurai was published in 2003...
- Common minke whaleCommon Minke WhaleThe common minke whale or northern minke whale, , is a species of minke whale within the suborder of baleen whales.-Taxonomy:...
, Balaenoptera acutorostrata - Antarctic minke whaleAntarctic Minke WhaleThe Antarctic minke whale or southern minke whale , is a species of minke whale within the suborder of baleen whales.-Taxonomy:...
, Balaenoptera bonaerensis - Sei whaleSei WhaleThe sei whale , Balaenoptera borealis, is a baleen whale, the third-largest rorqual after the blue whale and the fin whale. It inhabits most oceans and adjoining seas, and prefers deep offshore waters. It avoids polar and tropical waters and semi-enclosed bodies of water...
, Balaenoptera borealis - Bryde's whaleBryde's WhaleBryde's whales are baleen whales, one of the "great whales" or rorquals. They prefer tropical and temperate waters over the polar seas that other whales in their family frequent. They are largely coastal rather than pelagic. Bryde's whales are very similar in appearance to sei whales and almost as...
, Balaenoptera brydei - Eden's whale Balaenoptera edeni – Rice lists this as a separate species, MSW3 does not
- Omura's whale – MSW3 lists this is a synonym of Bryde's whale but suggests this may be temporary.
- Blue whaleBlue WhaleThe blue whale is a marine mammal belonging to the suborder of baleen whales . At in length and or more in weight, it is the largest known animal to have ever existed....
, Balaenoptera musculus - Fin whaleFin WhaleThe fin whale , also called the finback whale, razorback, or common rorqual, is a marine mammal belonging to the suborder of baleen whales. It is the second longest whale and the sixth largest living animal after the blue whale, bowhead whale, and right whales, growing to nearly 27 metres long...
, Balaenoptera physalus
- Common minke whale
- Genus Balaenoptera
- Subfamily Megapterinae
- Genus Megaptera
- Humpback whaleHumpback WhaleThe humpback whale is a species of baleen whale. One of the larger rorqual species, adults range in length from and weigh approximately . The humpback has a distinctive body shape, with unusually long pectoral fins and a knobbly head. It is an acrobatic animal, often breaching and slapping the...
, Megaptera novaeangliae
- Humpback whale
- Genus Megaptera
- Subfamily Balaenopterinae
- Family EschrichtiidaeEschrichtiidaeEschrichtiidae is a family of baleen whales in the suborder Mysticeti.At least five genera are recognised, but only a single species from one genus is still alive, the gray whale, Eschrichtius robustus....
- Genus Eschrichtius
- Gray whaleGray WhaleThe gray whale, Eschrichtius robustus, is a baleen whale that migrates between feeding and breeding grounds yearly. It reaches a length of about , a weight of , and lives 50–70 years. The common name of the whale comes from the gray patches and white mottling on its dark skin. Gray whales were...
, Eschrichtius robustus
- Gray whale
- Genus Eschrichtius
- Family Neobalaenidae: Pygmy Right Whale
- Genus Caperea
- Pygmy right whalePygmy Right WhaleThe pygmy right whale is a baleen whale, the sole member of the family Neobalaenidae. First described by John Edward Gray in 1846, it is the smallest of the baleen whales, ranging between and in length and 3,000 and 3,500 kg in mass...
, Caperea marginata
- Pygmy right whale
- Genus Caperea
- Family Balaenidae
- Suborder Odontoceti: toothed whales
- Family Delphinidae: DolphinDolphinDolphins are marine mammals that are closely related to whales and porpoises. There are almost forty species of dolphin in 17 genera. They vary in size from and , up to and . They are found worldwide, mostly in the shallower seas of the continental shelves, and are carnivores, mostly eating...
- Genus CephalorhynchusCephalorhynchusCephalorhynchus is a genus in the dolphin family Delphinidae. It consists of four species:*Commerson's Dolphin, Cephalorhyncus commersonii*Chilean Dolphin, Cephalorhyncus eutropia*Heaviside's Dolphin, Cephalorhyncus heavisidii...
- Commerson's dolphinCommerson's DolphinCommerson's Dolphin is one of four dolphins in the Cephalorhynchus genus. The species has also the common names Skunk Dolphin, Piebald Dolphin and Panda Dolphin...
, Cephalorhyncus commersonii - Chilean dolphinChilean DolphinThe Chilean Dolphin , also known as the Black Dolphin, is one of four dolphins in the Cephalorhynchus genus. The dolphin is only found off the coast of Chile, it is commonly referred to in the country as Tunina....
, Cephalorhyncus eutropia - Heaviside's dolphinHeaviside's DolphinThe Haviside's Dolphin , or mistakenly the Heaviside's Dolphin, is a small dolphin that is found off the coast of Namibia and the west coast of South Africa...
, Cephalorhyncus heavisidii - Hector's dolphinHector's DolphinHector's dolphin is the best-known of the four dolphins in the genus Cephalorhynchus and is found only in New Zealand. At about 1.4 m in length, it is one of the smallest cetaceans....
, Cephalorhyncus hectori
- Commerson's dolphin
- Genus Delphinus
- Long-beaked common dolphinLong-beaked Common DolphinThe Long-beaked Common Dolphin is a species of common dolphin. It has a more restricted range than the Short-beaked Common Dolphin . It has a disjointed range in coastal areas in tropical and warmer temperate oceans...
, Delphinus capensis - Short-beaked common dolphinShort-beaked Common DolphinThe short-beaked common dolphin is a species of common dolphin. It has a larger range than the long-beaked common dolphin , occurring throughout warm-temperate and tropical oceans, with the possible exception of the Indian Ocean...
, Delphinus delphis - Arabian common dolphin, Delphinus tropicalis. Rice recognises this as a separate species. MSW3 does not.
- Long-beaked common dolphin
- Genus Feresa
- Pygmy killer whalePygmy Killer WhaleThe pygmy killer whale is a small, rarely seen cetacean of the oceanic dolphin family . It derives its common name from sharing some physical characteristics with the orca It is the smallest species that has "whale" in its common name. In fact, "killer" may be more apt in the case of the pygmy...
, Feresa attenuata
- Pygmy killer whale
- Genus Globicephala
- Short-finned pilot whaleShort-finned Pilot WhaleThe Short-finned Pilot Whale is one of the two species of cetacean in the genus Globicephala. It is part of the oceanic dolphin family , though its behaviour is closer to that of the larger whales....
, Globicephala macrorhyncus - Long-finned pilot whaleLong-finned Pilot WhaleThe long-finned pilot whale is one of the two species of cetacean in the genus Globicephala. It belongs to the oceanic dolphin family , though its behavior is closer to that of the larger whales.-Description:...
, Globicephala melas
- Short-finned pilot whale
- Genus Grampus
- Risso's dolphinRisso's DolphinRisso's dolphin is the only species of dolphin in the genus Grampus.-Taxonomy:Risso's dolphin is named after Antoine Risso, whose description formed the basis of the first public description of the animal, by Georges Cuvier, in 1812...
, Grampus griseus
- Risso's dolphin
- Genus Lagenodelphis
- Fraser's dolphinFraser's DolphinFraser's Dolphin or Sarawak Dolphin is a cetacean in the family Delphinidae found in deep waters in the Pacific Ocean and to a lesser extent in the Indian and Atlantic Oceans.-Taxonomy:...
, Lagenodelphis hosei
- Fraser's dolphin
- Genus LagenorhynchusLagenorhynchusLagenorhynchus is a genus in the order Cetacea, traditionally containing six species:* white-beaked dolphin, Lagenorhynchus albirostris* Atlantic white-sided dolphin, Lagenorhynchus acutus...
- Atlantic white-sided dolphinAtlantic White-sided DolphinThe Atlantic White-sided Dolphin is a distinctively coloured dolphin found in the cool to temperate waters of the North Atlantic Ocean.-Taxonomy:...
, Lagenorhynchus acutus - White-beaked dolphinWhite-beaked DolphinThe White-beaked dolphin is a marine mammal belonging to the family Delphinidae in the suborder Odontoceti .-Taxonomy:...
, Lagenorhynchus albirostris - Peale's dolphinPeale's DolphinPeale's Dolphin is a small dolphin found in the waters around Tierra del Fuego at the foot of South America. It is also commonly known as the Black-chinned Dolphin or even Peale's Black-chinned Dolphin...
, Lagenorhynchus australis - Hourglass dolphinHourglass DolphinThe hourglass dolphin is a small dolphin in the family Delphinidae that inhabits Antarctic and sub-Antarctic waters....
, Lagenorhynchus cruciger - Pacific white-sided dolphinPacific White-sided DolphinThe Pacific White-sided Dolphin is a very active dolphin found in the cool to temperate waters of the North Pacific Ocean.-Taxonomy:...
, Lagenorhynchus obliquidens - Dusky dolphinDusky DolphinThe dusky dolphin is a dolphin found in coastal waters in the Southern Hemisphere. Its specific epithet is Latin for "dark" or "dim". It is very closely genetically related to the Pacific white-sided dolphin, but current scientific consensus is that they are distinct species...
, Lagenorhynchus obscurus
- Atlantic white-sided dolphin
- Genus Lissodelphis
- Northern right whale dolphinNorthern Right Whale DolphinThe Northern right whale dolphin is a small and slender species of marine mammal found in the North Pacific Ocean. The Northern right whale dolphin travels in groups of up to 2000, often with other cetaceans, in deep waters of the North Pacific...
, Lissodelphis borealis - Southern right whale dolphinSouthern Right Whale DolphinThe southern right whale dolphin, Lissodelphis peronii, is a small and slender species of mammal found in cool waters of the southern hemisphere...
, Lissodelphis peronii
- Northern right whale dolphin
- Genus OrcaellaOrcaellaThe snubfin dolphins are a genus of dolphins containing two members: the Irrawaddy Dolphin and Australian Snubfin Dolphin. The genus was long believed to be monotypic with the only species being the Irrawaddy Dolphin; however, in 2005, genetic analysis showed that the Australian Snubfin Dolphin is...
- Irrawaddy DolphinIrrawaddy dolphinThe Irrawaddy dolphin is a euryhaline species of oceanic dolphin found in discontinuous subpopulations near sea coasts and in estuaries and rivers in parts of the Bay of Bengal and Southeast Asia.-Etymology and taxonomic history:...
, Orcaella brevirostris- Australian snubfin dolphinAustralian Snubfin DolphinThe Australian snubfin dolphin is a dolphin found off the northern coasts of Australia. It closely resembles the Irrawaddy dolphin and was not described as a separate species until 2005. The Australian snubfin is tri-coloured, while the Irrawaddy dolphin only has two colours on its skin...
, Orcaella heinsohni. 2005 discovery, thus not recognized by Rice or MSW3 and subject to revision.
- Australian snubfin dolphin
- Irrawaddy Dolphin
- Genus Orcinus
- Killer Whale, Orcinus orca
- Genus Peponocephala
- Melon-headed whaleMelon-headed WhaleThe melon-headed whale is a cetacean of the oceanic dolphin family . It is closely related to the pygmy killer whale and pilot whale, and collectively these dolphin species are known by the common name blackfish. It is also related to the false killer whale...
, Peponocephala electra
- Melon-headed whale
- Genus Pseudorca
- False killer whaleFalse Killer WhaleThe False Killer Whale is a cetacean, and the third largest member of the oceanic dolphin family . It lives in temperate and tropical waters throughout the world. As its name implies, the False Killer Whale shares characteristics, such as appearance, with the more widely known Orca...
, Pseudorca crassidens
- False killer whale
- Genus Sotalia
- TucuxiTucuxiThe Tucuxi , alternately bufeo gris or bufeo negro is a dolphin found in the rivers of the Amazon Basin. The word "tucuxi" is derived from the Tupi language word tuchuchi-ana and has now been adopted as the species' common name...
, Sotalia fluviatilis, see the species article for a discussion - CosteroCosteroThe Costero is found in the coastal waters to the north and east of South America. The common name "costero" has been suggested by Caballero and colleagues due to the species' affinity for coastal habitats. The Costero is a member of the oceanic dolphin family . Physically it resembles the...
, Sotalia guianensis, see the species article for a discussion
- Tucuxi
- Genus Sousa
- Pacific humpback dolphin, Sousa chinensis
- Indian humpback dolphin, Sousa plumbea
- Atlantic humpback dolphin, Sousa teuszii
- Genus StenellaStenellaStenella is a genus of aquatic mammals in Delphinidae, the family informally known as the oceanic dolphins.Currently five species are recognised in this genus:*Pantropical Spotted Dolphin, S. attenuata*Atlantic Spotted Dolphin, S. frontalis...
- Pantropical spotted dolphinPantropical Spotted DolphinThe Pantropical Spotted Dolphin is a species of dolphin found in all the world's temperate and tropical oceans. The species was beginning to come under threat due to the killing of millions of individuals in tuna purse seines...
, Stenella attenuata - Clymene dolphinClymene DolphinThe Clymene dolphin , in older texts known as the Short-snouted spinner dolphin, is a dolphin endemic to the Atlantic Ocean.- Taxonomy :...
, Stenella clymene - Striped dolphinStriped DolphinThe Striped Dolphin is an extensively studied dolphin that is found in temperate and tropical waters of all the world's oceans.-Taxonomy:...
, Stenella coeruleoalba - Atlantic spotted dolphinAtlantic Spotted DolphinThe Atlantic Spotted Dolphin is a dolphin found in the Gulf Stream of the North Atlantic Ocean. Older members of the species have a very distinctive spotted coloration all over their body.-Taxonomy:...
, Stenella frontalis - Spinner dolphinSpinner DolphinThe Spinner Dolphin is a small dolphin found in off-shore tropical waters around the world. It is famous for its acrobatic displays in which they spin longitudinally along their axis as they leap through the air.-Taxonomy:...
, Stenella longirostris
- Pantropical spotted dolphin
- Genus Steno
- Rough-toothed dolphinRough-toothed DolphinThe Rough-toothed dolphin is species of dolphin that can be found in deep warm and tropical waters around the world.The species was first described by Georges Cuvier in 1823...
, Steno bredanensis
- Rough-toothed dolphin
- Genus Tursiops – Rice and MSW3 tentatively agree on this classification but see species article for more detail.
- Indian Ocean bottlenose dolphin, Tursiops aduncus
- Burrunan dolphinBurrunan dolphinThe Burrunan dolphin is a recently-described species of bottlenose dolphin found in parts of Victoria, Australia. By size, the Burrunan dolphin is between the other two species of bottlenose dolphin and only around 150 individuals have been found in two locations.-Taxonomy:The species was formally...
, Tursiops australis - Common bottlenose dolphinCommon Bottlenose DolphinTursiops truncatus, commonly known as the Common Bottlenose Dolphin, is the most well-known species from the family Delphinidae.Common bottlenose dolphins are the most familiar dolphins due to the wide exposure they receive in captivity in marine parks, dolphinarias, in movies, and television...
, Tursiops truncatus
- Genus Cephalorhynchus
- Family MonodontidaeMonodontidaeThe cetacean family Monodontidae comprises two unusual whale species, the narwhal, in which the male has a long tusk, and the white beluga whale...
- Genus Delphinapterus
- Beluga, Delphinapterus leucas
- Genus Monodon
- NarwhalNarwhalThe narwhal, Monodon monoceros, is a medium-sized toothed whale that lives year-round in the Arctic. One of two living species of whale in the Monodontidae family, along with the beluga whale, the narwhal males are distinguished by a characteristic long, straight, helical tusk extending from their...
, Monodon monoceros
- Narwhal
- Genus Delphinapterus
- Family Phocoenidae: PorpoisePorpoisePorpoises are small cetaceans of the family Phocoenidae; they are related to whales and dolphins. They are distinct from dolphins, although the word "porpoise" has been used to refer to any small dolphin, especially by sailors and fishermen...
s- Genus Neophocaena
- Finless PorpoiseFinless PorpoiseThe finless porpoise is one of six porpoise species. In the waters around Japan, at the northern end of its range, it is known as the sunameri . A freshwater population found in the Yangtze River in China is known locally as the jiangzhu or "river pig". There is a degree of taxonomic uncertainty...
, Neophocaena phocaenoides
- Finless Porpoise
- Genus Phocoena
- Spectacled porpoiseSpectacled PorpoiseThe Spectacled Porpoise is a rarely seen member of the porpoise family. The species is readily distinguished from other porpoises by a characteristic dark ring around the eyes, which gives the animals their name. This ring is commonly surrounded by a farther lighter ring...
, Phocoena dioptrica - Harbour porpoiseHarbour PorpoiseThe harbour porpoise is one of six species of porpoise. It is one of the smallest marine mammals. As its name implies, it stays close to coastal areas or river estuaries, and as such, is the most familiar porpoise to whale watchers. This porpoise often ventures up rivers, and has been seen...
, Phocoena phocaena - VaquitaVaquitaThe vaquita is a rare species of porpoise. It is endemic to the northern part of the Gulf of California. Estimates of the number of individuals alive range from 100 to 300. The word "vaquita" is Spanish for little cow...
, Phocoena sinus - Burmeister's porpoiseBurmeister's PorpoiseBurmeister's Porpoise is a species of porpoise endemic to the coast of South America. It was first described by Hermann Burmeister, for whom the species is named, in 1865...
, Phocoena spinipinnis
- Spectacled porpoise
- Genus Phocoenoides
- Dall's porpoiseDall's PorpoiseDall's porpoise is a species of porpoise found on the North Pacific. It came to worldwide attention in the 1970s when it was disclosed for the first time to the public that salmon fishing trawls were killing a lot, thousands of Dall's porpoises and other cetaceans each year by accidentally...
, Phocoenoides dalli
- Dall's porpoise
- Genus Neophocaena
- Family Physeteridae: Sperm whale familySperm whale familyPhyseteroidea is a superfamily including just three living species of whale; the Sperm Whale, in the genus Physeter, and the Pygmy Sperm Whale and Dwarf Sperm Whale, in the genus Kogia...
- Genus Physeter
- Sperm whaleSperm WhaleThe sperm whale, Physeter macrocephalus, is a marine mammal species, order Cetacea, a toothed whale having the largest brain of any animal. The name comes from the milky-white waxy substance, spermaceti, found in the animal's head. The sperm whale is the only living member of genus Physeter...
, Physeter catodon (syn. P. macrocephalus)
- Sperm whale
- Genus Physeter
- Family Kogiidae – MSW3 treats Kogia as a member of Physeteridae
- Genus Kogia
- Pygmy sperm whalePygmy Sperm WhaleThe Pygmy Sperm Whale is one of three species of toothed whale in the sperm whale family. They are not often sighted at sea, and most of what is known about them comes from the examination of stranded specimens.-Taxonomy:...
, Kogia breviceps - Dwarf sperm whaleDwarf Sperm WhaleThe Dwarf Sperm Whale is one of three species in the sperm whale family. They are not often sighted at sea. As such, most information is a result of the study of stranded carcasses.-Taxonomy:...
, Kogia sima
- Pygmy sperm whale
- Genus Kogia
- Superfamily Platanistoidea: River dolphins
- Family IniidaeIniidaeIniidae is a family of river dolphins containing one living and three extinct genera.-Taxonomy:The family was described by John Edward Gray in 1846.Current classifications include a single living genera, Inia, with one species and three subspecies...
- Genus IniaIniaInia is a genus of river dolphin containing one or possibly two species.-Taxonomy:The genus was described by Alcide d'Orbigny in 1834 when Delphinus geoffrensis, described by Henri Marie Ducrotay de Blainville in 1817, was recognized to be a unique taxon...
- Amazon river dolphin, Inia geoffrensis
- Bolivian river dolphinBolivian River DolphinThe Bolivian river dolphin is a subspecies of the Amazon river dolphin. Although older publications as well as some recent publications consider the boliviensis population as distinct species from Inia geoffrensis, the great majority of the scientific community including the IUCN consider...
, Inia boliviensis
- Bolivian river dolphin
- Amazon river dolphin, Inia geoffrensis
- Genus Inia
- † Family Lipotidae – MSW3 treats Lipotes as a member of Iniidae
- † Genus Lipotes
- † Baiji, Lipotes vexillifer
- † Genus Lipotes
- Family Pontoporiidae – MSW3 treats Pontoporia as a member of Iniidae
- Genus PontoporiaLa Plata DolphinThe La Plata Dolphin or Franciscana is found in coastal Atlantic waters of southeastern South America. Taxonomically it is a member of the river dolphin group and the only one that actually lives in the ocean and saltwater estuaries, rather than inhabiting exclusively freshwater...
- La Plata dolphinLa Plata DolphinThe La Plata Dolphin or Franciscana is found in coastal Atlantic waters of southeastern South America. Taxonomically it is a member of the river dolphin group and the only one that actually lives in the ocean and saltwater estuaries, rather than inhabiting exclusively freshwater...
, Pontoporia blainvillei
- La Plata dolphin
- Genus Pontoporia
- Family Platanistidae
- Genus Platanista
- Ganges and Indus River dolphinGanges and Indus River DolphinThe South Asian River Dolphin is a freshwater or river dolphin found in India, Nepal and Pakistan which is split into two sub-species, the Ganges River Dolphin and Indus River Dolphin...
, Platanista gangetica. MSW3 treats Platanista minor as a separate species, with common names Ganges River Dolphin and Indus River DolphinIndus River dolphinThe Indus River dolphin is a sub-species of freshwater or river dolphin found in the Indus River of Pakistan...
, respectively.
- Ganges and Indus River dolphin
- Genus Platanista
- Family Iniidae
- Family Delphinidae: Dolphin
- Superfamily Ziphioidea: Beaked whales
- Family Ziphidae,
- Genus Berardius
- Arnoux's beaked whale, Berardius arnuxii
- Baird's beaked whale (North Pacific Bottlenose Whale), Berardius bairdii
- Subfamily Hyperoodontidae
- Genus Hyperoodon
- Northern bottlenose whale, Hyperoodon ampullatus
- Southern bottlenose whale, Hyperoodon planifrons
- Genus Indopacetus
- Indo-Pacific beaked whale (Longman's Beaked Whale), Indopacetus pacificus
- Genus Mesoplodon, Mesoplodont WhaleMesoplodont whaleMesoplodont whales are fourteen species of whale in the genus Mesoplodon, making it the largest genus in the cetacean order. Two species were described as recently as 1991 and 2002 , and marine biologists predict the discovery of more species in the future. They are the most poorly known group of...
- Sowerby's beaked whaleSowerby's Beaked WhaleSowerby's beaked whale , also known as the North Atlantic/North Sea beaked whale, was the first beaked whale to be described. James Sowerby, an English naturalist and artist, first described the species in 1804 from a skull obtained from a male that had stranded in the Moray Firth, Scotland, in 1800...
, Mesoplodon bidens - Andrews' beaked whaleAndrews' Beaked WhaleAndrews' beaked whale , sometimes known as the deep-crest beaked whale or splay-toothed whale, is one of the most poorly known members of a poorly known genus...
, Mesoplodon bowdoini - Hubbs' beaked whaleHubbs' Beaked WhaleHubbs' beaked whale was initially thought to be an Andrews' beaked whale when discovered by ichthyologist Carl Hubbs; however, it was named in his honor when it was discovered to be a new species. This species has the typical dentition found in the genus, but its main outstanding features are a...
, Mesoplodon carlhubbsi - Blainville's beaked whaleBlainville's Beaked WhaleBlainville's beaked whale , or the dense-beaked whale, is the widest ranging mesoplodont whale and perhaps the most documented. Henri de Blainville first described the species in 1817 from a small piece of jaw—the heaviest bone he had ever come across—which resulted in the name densirostris...
, Mesoplodon densirostris - Gervais' beaked whaleGervais' Beaked WhaleGervais' beaked whale , sometimes known as the Antillian beaked whale, Gulf Stream beaked whale, or European beaked whale is the most frequently stranding type of mesoplodont whale off the coast of North America...
, Mesoplodon europaeus - Ginkgo-toothed beaked whaleGinkgo-toothed Beaked WhaleThe ginkgo-toothed beaked whale is a poorly known species of whale even for a beaked whale, and was named for the unusual shape of its dual teeth...
, Mesoplodon ginkgodens - Gray's beaked whaleGray's Beaked WhaleGray's beaked whale , sometimes known as Haast's beaked whale, the Scamperdown whale, or the southern beaked whale, is one of the better-known members of the genus Mesoplodon. The scientific name refers to John Edward Gray, a zoologist at the British Museum. This species is fairly gregarious and...
, Mesoplodon grayi - Hector's beaked whaleHector's Beaked WhaleHector's beaked whale , is a small mesoplodont living in the Southern Hemisphere. This whale is named after Sir James Hector, a founder of the colonial museum in Wellington, New Zealand...
, Mesoplodon hectori - Strap-toothed whale, Mesoplodon layardii
- True's beaked whaleTrue's Beaked WhaleThe True's Beaked Whale is a medium sized whale in the Mesoplodont genus. The common name is in reference to Frederick W. True, a curator at the United States National Museum...
, Mesoplodon mirus - Perrin's beaked whalePerrin's Beaked WhalePerrin's beaked whale is the newest species of beaked whale to be described. The first two specimens were found in May 1975 stranded on the California coast, with two more specimens being found in 1978 and 1979, and the last in September 1997...
, Mesoplodon perrini. This species was recognised in 2002 and as such is listed by MSW3 but not Rice. - Pygmy beaked whalePygmy Beaked WhaleThe pygmy beaked whale , also known as the bandolero beaked whale, Peruvian beaked whale and lesser beaked whale, is the smallest of the mesoplodonts and one of the newest discoveries. There were at least two dozen sightings of an unknown beaked whale named Mesoplodon sp...
, Mesoplodon peruvianus - Stejneger's beaked whaleStejneger's Beaked WhaleStejneger's beaked whale , sometimes known as the Bering Sea beaked whale or the Saber-toothed whale, is a poorly-known member of the genus Mesoplodon inhabiting the northern North Pacific Ocean. Leonhard Hess Stejneger initially described the species in 1885 from a skull, and nothing more of the...
, Mesoplodon stejnegeri - Spade-toothed whale, Mesoplodon traversii
- Sowerby's beaked whale
- Genus Hyperoodon
- Genus Tasmacetus
- Shepherd's beaked whaleShepherd's Beaked WhaleShepherd's beaked whale , also commonly called Tasman's beaked whale or simply the Tasman whale, is a cetacean of the family Ziphidae. The whale has been little studied. Only four confirmed at sea sightings have been made and 42 strandings recorded . It was first known to science in 1937, being...
, Tasmacetus shepherdi
- Shepherd's beaked whale
- Genus Ziphius
- Cuvier's beaked whaleCuvier's Beaked WhaleCuvier's beaked whale is the most widely distributed of all the beaked whales. It is the only member of the genus Ziphius. Another common name for the species is goose-beaked whale because its head is said to be shaped like the beak of a goose. Georges Cuvier first described it in 1823 from part...
, Ziphius cavirostris
- Cuvier's beaked whale
- Genus Berardius
- Family Ziphidae,
- Suborder Mysticeti: Baleen whales
†Recently extinct
External links
Including a page on taxonomy- EIA Cetacean campaign: Reports and latest info.
- EIA in USA: reports etc.