Chinkapin oak
Encyclopedia
Quercus muehlenbergii, the chinkapin oak (or chinquapin oak), is an oak in the white oak group (Quercus sect. Quercus). The scientific name is often incorrectly spelled Q. muhlenbergii, and the species was often called Quercus acuminata in older literature. Quercus muehlenbergii is native to eastern and central North America
, ranging from Vermont
west to Wisconsin
and south to South Carolina
, western Florida
, New Mexico
, and northeastern Mexico
from Coahuila
south to Hidalgo.
The tree's scientific name honors Gotthilf Heinrich Ernst Muhlenberg
(1753–1815), a Lutheran pastor and amateur botanist in Pennsylvania
. In publishing the name Quercus mühlenbergii, German
-American
botanist George Engelmann
mistakenly used an umlaut
in spelling Muhlenberg's name, even though Pennsylvania-born Muhlenberg himself did not use an umlaut in his name. Under the modern rules of botanical nomenclature, umlauts are transliterated, with "ü" becoming "ue", hence Engelmann's Quercus mühlenbergii is now presented as Quercus muehlenbergii. In lack of evidence that Engelmann's use of the umlaut was an unintended error, and hence correctable, the muehlenbergii spelling is considered correct, although the more appropriate orthographic variant
Quercus muhlenbergii is often seen.
The low-growing, cloning
Q. prinoides (dwarf chinkapin oak)
is similar to Q. muehlenbergii and has been confused with it in the past, but is now generally accepted as a distinct species. If the two are considered to be conspecific, the earlier-published name Quercus prinoides has priority over Q. muehlenbergii, and the larger chinkapin oak can then be classified as Quercus prinoides var. acuminata, with the dwarf chinkapin oak being Quercus prinoides var. prinoides. Q. prinoides was named and described by the German botanist Karl (Carl) Ludwig Willdenow
in 1801, in a German journal article by Muhlenberg.
Chinkapin oak is monoecious in flowering habit; flowers emerge in April to late May or early June. The staminate flowers are borne in catkins that develop from the leaf axils of the previous year, and the pistillate flowers develop from the axils of the current year's leaves. The fruit, an acorn or nut, is borne singly or in pairs, matures in 1 year, and ripens in September or October. About half of the acorn is enclosed in a thin cup and is chestnut brown to nearly black.
Chinkapin oak is closely related to the smaller but generally similar dwarf chinkapin oak (Quercus prinoides)
. Chinkapin oak is usually a tree, but occasionally shrubby, while dwarf chinkapin oak is a low-growing, clone-forming
shrub. The two species generally occur in different habitats: chinquapin oak is typically found on calcareous soils
and rocky slopes, while dwarf chinkapin oak is usually found on acidic substrates
, primarily sand or sandy soils, and also dry shales.
Chinkapin oak is also sometimes confused with the related chestnut oak (Quercus montana), which it closely resembles. However, unlike the pointed teeth on the leaves of the chinkapin oak, chestnut oak leaves generally have rounded teeth. The two species have contrasting kinds of bark: Chinkapin oak has a gray, flaky bark very similar to that of white oak (Q. alba) but with a more yellow-brown cast to it (hence the occasional name yellow oak for this species), while chestnut oak has dark, solid, deeply ridged bark. The chinkapin oak also has smaller acorns than the chestnut oak or another similar species, the swamp chestnut oak (Q. michauxii), which have some of the largest acorns of any oaks.
It grows in association with white oak (Quercus alba), black oak (Q. velutina), northern red oak (Q. rubra), scarlet oak (Q. coccinea), sugar maple (Acer saccharum), red maple (A. rubrum), hickories (Carya spp.), black cherry (Prunus serotina), cucumbertree (Magnolia acuminata), white ash (Fraxinus americana), American basswood (Tilia americana), black walnut (Juglans nigra), butternut (J. cinerea), and yellow-poplar (Liriodendron tulipifera). American beech (Fagus grandifolia), shortleaf pine (Pinus echinata), pitch pine (P. rigida), Virginia pine (P. uirginiana), Ozark chinkapin (Castanea ozarkensis), eastern redcedar (Juniperus virginiana), bluejack oak (Quercus incana), southern red oak (Q. falcata), blackgum (Nyssa sylvatica), and winged elm (Ulmus alata) also grow in association with chinkapin oak. In the Missouri Ozarks a redcedar-chinkapin oak association has been described.
The most common small tree and shrub species found in association with chinkapin oak include flowering dogwood (Cornus ), sassafras (Sassafras albidum), sourwood (Oxydendron arboreum), eastern hophornbeam (Ostrya virginiana), Vaccinium spp., Viburnum spp., hawthorns (Crataegus spp.), and sumacs (Rhus spp.). The most common woody vines are wild grape (Vitis spp.) and greenbrier (Smilax spp.).
, and the resulting decay can cause serious losses.
Oak wilt (Ceratocystis fagacearum), a vascular disease, attacks Chinkapin oak and usually kills the tree within two to four years. Other diseases that attack Chinkapin oak include the cankers Strumella coryneoidea and Nectria galligena, shoestring root rot (Armillarea mellea), anthracnose (Gnomonia veneta), and leaf blister (Taphrina spp.).
The most serious defoliating insect
s that attack Chinkapin oak are the gypsy moth (Lymantria dispar)
, the orangestriped oakworm (Anisota senatoria), and the variable oakleaf caterpillar (Heterocampa manteo). Insects that bore into the bole and seriously degrade the products cut from infested trees include the carpenterworm (Prionoyxstus robiniae), little carpenterworm (P. macmurtrei), white oak borer (Goes tigrinus), Columbian timber beetle (Corthylus columbianus), oak timberworm (Arrhenodes minutus), and twolined chestnut borer (Agrilus bilineatus). The acorn weevils (Curculio spp.), larvae of moths (Valentinia glandulella and Melissopus latiferreanus), and gall forming cynipids (Callirhytis spp.) attack and destroy the acorns.
The chinkapin oak is especially known for its sweet and palatable acorns. Indeed, the nuts contained inside of the thin shell are among the sweetest of any oak, with an excellent taste even when eaten raw, providing an excellent source of food for both wildlife and people. The acorns are eaten by squirrels, mice, voles, chipmunks, deer, turkey, and other birds.
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
, ranging from Vermont
Vermont
Vermont is a state in the New England region of the northeastern United States of America. The state ranks 43rd in land area, , and 45th in total area. Its population according to the 2010 census, 630,337, is the second smallest in the country, larger only than Wyoming. It is the only New England...
west to Wisconsin
Wisconsin
Wisconsin is a U.S. state located in the north-central United States and is part of the Midwest. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michigan to the northeast, and Lake Superior to the north. Wisconsin's capital is...
and south to South Carolina
South Carolina
South Carolina is a state in the Deep South of the United States that borders Georgia to the south, North Carolina to the north, and the Atlantic Ocean to the east. Originally part of the Province of Carolina, the Province of South Carolina was one of the 13 colonies that declared independence...
, western Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...
, New Mexico
New Mexico
New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...
, and northeastern Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...
from Coahuila
Coahuila
Coahuila, formally Coahuila de Zaragoza , officially Estado Libre y Soberano de Coahuila de Zaragoza is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico...
south to Hidalgo.
Classification and nomenclature
Since its recognition as a different species from the similar-appearing chestnut oak (Quercus prinus), Q. muehlenbergii has generally been regarded as a distinct species; no subspecies or varieties are currently recognized within it, although a few infraspecific variants had been accepted in the past.The tree's scientific name honors Gotthilf Heinrich Ernst Muhlenberg
Gotthilf Heinrich Ernst Muhlenberg
Gotthilf Heinrich Ernst Muhlenberg was an American clergyman and botanist.-Biography:The son of Heinrich Melchior Muhlenberg, he was born in Trappe, Pennsylvania. He was educated at in Halle starting in 1763 and in 1769 at the University of Halle. He returned to Pennsylvania in September 1770...
(1753–1815), a Lutheran pastor and amateur botanist in Pennsylvania
Pennsylvania
The Commonwealth of Pennsylvania is a U.S. state that is located in the Northeastern and Mid-Atlantic regions of the United States. The state borders Delaware and Maryland to the south, West Virginia to the southwest, Ohio to the west, New York and Ontario, Canada, to the north, and New Jersey to...
. In publishing the name Quercus mühlenbergii, German
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...
-American
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...
botanist George Engelmann
George Engelmann
George Engelmann, also known as Georg Engelmann, was a German-American botanist. He was instrumental in describing the flora of the west of North America, then very poorly-known; he was particularly active in the Rocky Mountains and northern Mexico.-Origins:George Engelmann was born in Frankfurt...
mistakenly used an umlaut
Umlaut
Umlaut may refer to:*Diaeresis , a pair of dots above a vowel, used in German to mark umlaut**Metal umlaut, the same diacritic in names of heavy metal or hard rock bands for effect...
in spelling Muhlenberg's name, even though Pennsylvania-born Muhlenberg himself did not use an umlaut in his name. Under the modern rules of botanical nomenclature, umlauts are transliterated, with "ü" becoming "ue", hence Engelmann's Quercus mühlenbergii is now presented as Quercus muehlenbergii. In lack of evidence that Engelmann's use of the umlaut was an unintended error, and hence correctable, the muehlenbergii spelling is considered correct, although the more appropriate orthographic variant
Orthography
The orthography of a language specifies a standardized way of using a specific writing system to write the language. Where more than one writing system is used for a language, for example Kurdish, Uyghur, Serbian or Inuktitut, there can be more than one orthography...
Quercus muhlenbergii is often seen.
The low-growing, cloning
Cloning
Cloning in biology is the process of producing similar populations of genetically identical individuals that occurs in nature when organisms such as bacteria, insects or plants reproduce asexually. Cloning in biotechnology refers to processes used to create copies of DNA fragments , cells , or...
Q. prinoides (dwarf chinkapin oak)
Quercus prinoides
Quercus prinoides, commonly known as Dwarf Chinkapin Oak, Dwarf Chinquapin Oak, Dwarf Chestnut Oak or Scrub Chestnut Oak, is a shrubby, clone-forming oak native to eastern North America, ranging from Massachusetts to the Carolinian forest zone of southern Ontario to Minnesota, south to North...
is similar to Q. muehlenbergii and has been confused with it in the past, but is now generally accepted as a distinct species. If the two are considered to be conspecific, the earlier-published name Quercus prinoides has priority over Q. muehlenbergii, and the larger chinkapin oak can then be classified as Quercus prinoides var. acuminata, with the dwarf chinkapin oak being Quercus prinoides var. prinoides. Q. prinoides was named and described by the German botanist Karl (Carl) Ludwig Willdenow
Carl Ludwig Willdenow
Carl Ludwig Willdenow was a German botanist, pharmacist, and plant taxonomist. He is considered one of the founders of phytogeography, the study of the geographic distribution of plants...
in 1801, in a German journal article by Muhlenberg.
Description
Key characteristics of Quercus muehlenbergii (chinkapin oak):- The leaf base is typically more rounded.
- The veins and sinuses are regular.
- Acorns on short stalks and turn chestnut brown in the fall.
- The leaves have sharp teeth but no bristles, as a member of the white oak subgenus of Quercus.
Chinkapin oak is monoecious in flowering habit; flowers emerge in April to late May or early June. The staminate flowers are borne in catkins that develop from the leaf axils of the previous year, and the pistillate flowers develop from the axils of the current year's leaves. The fruit, an acorn or nut, is borne singly or in pairs, matures in 1 year, and ripens in September or October. About half of the acorn is enclosed in a thin cup and is chestnut brown to nearly black.
Chinkapin oak is closely related to the smaller but generally similar dwarf chinkapin oak (Quercus prinoides)
Quercus prinoides
Quercus prinoides, commonly known as Dwarf Chinkapin Oak, Dwarf Chinquapin Oak, Dwarf Chestnut Oak or Scrub Chestnut Oak, is a shrubby, clone-forming oak native to eastern North America, ranging from Massachusetts to the Carolinian forest zone of southern Ontario to Minnesota, south to North...
. Chinkapin oak is usually a tree, but occasionally shrubby, while dwarf chinkapin oak is a low-growing, clone-forming
Cloning
Cloning in biology is the process of producing similar populations of genetically identical individuals that occurs in nature when organisms such as bacteria, insects or plants reproduce asexually. Cloning in biotechnology refers to processes used to create copies of DNA fragments , cells , or...
shrub. The two species generally occur in different habitats: chinquapin oak is typically found on calcareous soils
Calcareous
Calcareous is an adjective meaning mostly or partly composed of calcium carbonate, in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines.-In zoology:...
and rocky slopes, while dwarf chinkapin oak is usually found on acidic substrates
Soil pH
The soil pH is a measure of the acidity or basicity in soils. pH is defined as the negative logarithm of the activity of hydrogen ions in solution. It ranges from 0 to 14, with 7 being neutral. A pH below 7 is acidic and above 7 is basic. Soil pH is considered a master variable in soils as it...
, primarily sand or sandy soils, and also dry shales.
Chinkapin oak is also sometimes confused with the related chestnut oak (Quercus montana), which it closely resembles. However, unlike the pointed teeth on the leaves of the chinkapin oak, chestnut oak leaves generally have rounded teeth. The two species have contrasting kinds of bark: Chinkapin oak has a gray, flaky bark very similar to that of white oak (Q. alba) but with a more yellow-brown cast to it (hence the occasional name yellow oak for this species), while chestnut oak has dark, solid, deeply ridged bark. The chinkapin oak also has smaller acorns than the chestnut oak or another similar species, the swamp chestnut oak (Q. michauxii), which have some of the largest acorns of any oaks.
Soil and topography
Chinkapin oak is generally found on well-drained upland soils derived from limestone or where limestone outcrops occur. Occasionally it is found on well-drained limestone soils along streams. The Chinkapin oak is generally found on soils that are weakly acid (pH about 6.5) to alkaline (above pH 7.0). It grows on both northerly and southerly aspects but is more common on the warmer southerly aspects. It is absent or rare at high elevations in the Appalachians.Associated cover
It is rarely a predominant tree, but it grows in association with many other species. It is a component of the forest cover type White Oak-Black Oak-Northern Red Oak (Society of American Foresters Type 52) and the Post Oak-Blackjack Oak (Type 40) (2).It grows in association with white oak (Quercus alba), black oak (Q. velutina), northern red oak (Q. rubra), scarlet oak (Q. coccinea), sugar maple (Acer saccharum), red maple (A. rubrum), hickories (Carya spp.), black cherry (Prunus serotina), cucumbertree (Magnolia acuminata), white ash (Fraxinus americana), American basswood (Tilia americana), black walnut (Juglans nigra), butternut (J. cinerea), and yellow-poplar (Liriodendron tulipifera). American beech (Fagus grandifolia), shortleaf pine (Pinus echinata), pitch pine (P. rigida), Virginia pine (P. uirginiana), Ozark chinkapin (Castanea ozarkensis), eastern redcedar (Juniperus virginiana), bluejack oak (Quercus incana), southern red oak (Q. falcata), blackgum (Nyssa sylvatica), and winged elm (Ulmus alata) also grow in association with chinkapin oak. In the Missouri Ozarks a redcedar-chinkapin oak association has been described.
The most common small tree and shrub species found in association with chinkapin oak include flowering dogwood (Cornus ), sassafras (Sassafras albidum), sourwood (Oxydendron arboreum), eastern hophornbeam (Ostrya virginiana), Vaccinium spp., Viburnum spp., hawthorns (Crataegus spp.), and sumacs (Rhus spp.). The most common woody vines are wild grape (Vitis spp.) and greenbrier (Smilax spp.).
Reaction to competition
Chinkapin oak is classed as intolerant of shade. It withstands moderate shading when young but becomes more intolerant of shade with age. It is regarded as a climax species on dry, drought prone soils, especially those of limestone origin. On more moist sites it is subclimax to climax. It is often found as a component of the climax vegetation in stands on mesic sites with limestone soils. However, many oak-hickory stands on moist sites that contain chinkapin oak are succeeded by a climax forest including beech, maple, and ash.Diseases and pests
Severe wildfire kills chinkapin oak saplings and small pole-size trees, but these often resprout. However, fire scars serve as entry points for decay-causing fungiFungus
A fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from plants, animals, and bacteria...
, and the resulting decay can cause serious losses.
Oak wilt (Ceratocystis fagacearum), a vascular disease, attacks Chinkapin oak and usually kills the tree within two to four years. Other diseases that attack Chinkapin oak include the cankers Strumella coryneoidea and Nectria galligena, shoestring root rot (Armillarea mellea), anthracnose (Gnomonia veneta), and leaf blister (Taphrina spp.).
The most serious defoliating insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...
s that attack Chinkapin oak are the gypsy moth (Lymantria dispar)
Gypsy moth
The gypsy moth, Lymantria dispar, is a moth in the family Lymantriidae of Eurasian origin. Originally ranging from Europe to Asia, it was introduced to North America in the late 1860s and has been expanding its range ever since...
, the orangestriped oakworm (Anisota senatoria), and the variable oakleaf caterpillar (Heterocampa manteo). Insects that bore into the bole and seriously degrade the products cut from infested trees include the carpenterworm (Prionoyxstus robiniae), little carpenterworm (P. macmurtrei), white oak borer (Goes tigrinus), Columbian timber beetle (Corthylus columbianus), oak timberworm (Arrhenodes minutus), and twolined chestnut borer (Agrilus bilineatus). The acorn weevils (Curculio spp.), larvae of moths (Valentinia glandulella and Melissopus latiferreanus), and gall forming cynipids (Callirhytis spp.) attack and destroy the acorns.
Uses
Like that of other white oak species, the wood of the chinkapin oak (Quercus muehlenbergii) is a durable hardwood prized for many types of construction.The chinkapin oak is especially known for its sweet and palatable acorns. Indeed, the nuts contained inside of the thin shell are among the sweetest of any oak, with an excellent taste even when eaten raw, providing an excellent source of food for both wildlife and people. The acorns are eaten by squirrels, mice, voles, chipmunks, deer, turkey, and other birds.
Other references
- Flora of North America: Quercus muhlenbergii RangeMap:
- Quercus muehlenbergii images at bioimages.vanderbilt.edu
- http://www.na.fs.fed.us/spfo/pubs/silvics_manual/volume_2/quercus/muehlenbergii.htm