Peloridiidae is a family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of true bugs

Hemiptera

Hemiptera is an order of insects most often known as the true bugs , comprising around 50,000–80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, and others...

, comprising 17 genera and 36 species. They are small, ranging in length from 2 to 4 mm, rarely seen, peculiarly lumpy, flattened bugs found in Patagonia

Patagonia

Patagonia is a region located in Argentina and Chile, integrating the southernmost section of the Andes mountains to the southwest towards the Pacific ocean and from the east of the cordillera to the valleys it follows south through Colorado River towards Carmen de Patagones in the Atlantic Ocean...

 (Argentina

Argentina

Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

 and Chile

Chile

Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

), New Zealand

New Zealand

New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

, eastern Australia

Australia

Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

, Lord Howe Island

Lord Howe Island

Lord Howe Island is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, directly east of mainland Port Macquarie, and about from Norfolk Island. The island is about 11 km long and between 2.8 km and 0.6 km wide with an area of...

 and New Caledonia

New Caledonia

New Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...

. All the Peloridiidae species are flightless, except one. Their present distribution suggests that they have existed before the breakup of Gondwana, and their relation to Heteroptera

Heteroptera

Heteroptera is a group of about 40,000 species of insects in the Hemiptera. Sometimes called "true bugs", that name more commonly refers to Hemiptera as a whole, and "typical bugs" might be used as a more unequivocal alternative since among the Hemiptera the heteropterans are most consistently and...

 (true bugs) dates back to at least the upper Permian

Permian

The PermianThe term "Permian" was introduced into geology in 1841 by Sir Sir R. I. Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil; Murchison asserted in 1841 that he named his "Permian...

, more than 230 Myr ago. Peloridiids are found amongst moss

Moss

Mosses are small, soft plants that are typically 1–10 cm tall, though some species are much larger. They commonly grow close together in clumps or mats in damp or shady locations. They do not have flowers or seeds, and their simple leaves cover the thin wiry stems...

es and liverwort

Marchantiophyta

The Marchantiophyta are a division of bryophyte plants commonly referred to as hepatics or liverworts. Like other bryophytes, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information....

s, commonly in association with southern beech

Nothofagus

Nothofagus, also known as the southern beeches, is a genus of 35 species of trees and shrubs native to the temperate oceanic to tropical Southern Hemisphere in southern South America and Australasia...

 forests. They have become known as moss bugs for their habit of feeding on mosses.

Peloridiidae is the only family in the suborder Coleorrhyncha, although there are alternative classifications in which the group is treated as an infraorder within the suborder Prosorrhyncha

Prosorrhyncha

The name Prosorrhyncha is a name for a suborder of Hemiptera, comprising a grouping of the traditional taxon "Heteroptera" plus its sister taxon, the family Peloridiidae...

, in which case it is called Peloridiomorpha (see the Heteroptera

Heteroptera

Heteroptera is a group of about 40,000 species of insects in the Hemiptera. Sometimes called "true bugs", that name more commonly refers to Hemiptera as a whole, and "typical bugs" might be used as a more unequivocal alternative since among the Hemiptera the heteropterans are most consistently and...

 and Prosorrhyncha

Prosorrhyncha

The name Prosorrhyncha is a name for a suborder of Hemiptera, comprising a grouping of the traditional taxon "Heteroptera" plus its sister taxon, the family Peloridiidae...

pages for a discussion).

Images

• Peloridium hammoniorum http://homepage.ruhr-uni-bochum.de/Andre.Mursch/14.htm