Cyathea capensis
Encyclopedia
Cyathea capensis, also known as the Forest Tree Fern, is a widespread and highly variable species of tree fern
. It is divided into two subspecies
: C. c. capensis from the Old World
Tropics
and C. c. polypodioides from the New World
Tropics
. It is the only member of the family Cyatheaceae
found in both geographic regions. The erect trunk is up to 4.5 m tall, occasionally even 6 m, and about 15 cm in diameter. Frond
s are pinnate
and 2-3 m in length. The lowest pinnae may be separated from the others along the rachis
and form a clump around the crown, similar to the "wig" of Cyathea baileyana
. The rachis
is smooth to slightly warty and covered in scale
s that range from tan to brown or dark brown. Sori
occur in two rows, one along each side of the pinnule midvein, and are covered by scale-like indusia.
The subspecies C. c. capensis is native to South Africa
(from the Western Cape
province to KwaZulu-Natal
and Mpumalanga
), Swaziland
, Zimbabwe
, Mozambique
, Malawi
and Tanzania
, where it grows in shaded moist forest along rivers and beside waterfalls at an altitude of 360-1820 m. The other subspecies, C. c. polypodioides, grows in southeastern Brazil
in montane forest at 900-2000 m.
The specific epithet
capensis refers to the Cape of Good Hope
in South Africa, where the type
specimen was collected.
C. capensis is relatively easy to grow if provided with a semi-shady, moist and sheltered environment. It can likely tolerate several degrees of frost.
Cyatheales
The order Cyatheales is a taxonomic division of the fern subclass, Cyatheatae, which includes the tree ferns. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicates that the order is monophyletic. Some species in the Cyatheales have tree-like growth...
. It is divided into two subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...
: C. c. capensis from the Old World
Old World
The Old World consists of those parts of the world known to classical antiquity and the European Middle Ages. It is used in the context of, and contrast with, the "New World" ....
Tropics
Tropics
The tropics is a region of the Earth surrounding the Equator. It is limited in latitude by the Tropic of Cancer in the northern hemisphere at approximately N and the Tropic of Capricorn in the southern hemisphere at S; these latitudes correspond to the axial tilt of the Earth...
and C. c. polypodioides from the New World
New World
The New World is one of the names used for the Western Hemisphere, specifically America and sometimes Oceania . The term originated in the late 15th century, when America had been recently discovered by European explorers, expanding the geographical horizon of the people of the European middle...
Tropics
Tropics
The tropics is a region of the Earth surrounding the Equator. It is limited in latitude by the Tropic of Cancer in the northern hemisphere at approximately N and the Tropic of Capricorn in the southern hemisphere at S; these latitudes correspond to the axial tilt of the Earth...
. It is the only member of the family Cyatheaceae
Cyatheaceae
The Cyatheaceae is the scaly tree fern family and includes the world's tallest tree ferns, which reach heights up to 20 m. They are also very ancient plants, appearing in the fossil record in the late Jurassic, though the modern genera likely appeared in the Tertiary. Cyatheaceae is the largest...
found in both geographic regions. The erect trunk is up to 4.5 m tall, occasionally even 6 m, and about 15 cm in diameter. Frond
Frond
The term frond refers to a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group...
s are pinnate
Pinnate
Pinnate is a term used to describe feather-like or multi-divided features arising from both sides of a common axis in plant or animal structures, and comes from the Latin word pinna meaning "feather", "wing", or "fin". A similar term is pectinate, which refers to a comb-like arrangement of parts...
and 2-3 m in length. The lowest pinnae may be separated from the others along the rachis
Rachis
Rachis is a biological term for a main axis or "shaft".-In zoology:In vertebrates a rachis can refer to the series of articulated vertebrae, which encase the spinal cord. In this case the rachis usually form the supporting axis of the body and is then called the spine or vertebral column...
and form a clump around the crown, similar to the "wig" of Cyathea baileyana
Cyathea baileyana
Cyathea baileyana, also known as the wig tree fern, is a species of tree fern native to northeastern Queensland in Australia, where it grows in wet gullies and forest at an altitude of 850-1200 m. It is a rare species that is seldom found in the wild. The erect trunk is 4-5 m tall, approximately 10...
. The rachis
Rachis
Rachis is a biological term for a main axis or "shaft".-In zoology:In vertebrates a rachis can refer to the series of articulated vertebrae, which encase the spinal cord. In this case the rachis usually form the supporting axis of the body and is then called the spine or vertebral column...
is smooth to slightly warty and covered in scale
Scale (zoology)
In most biological nomenclature, a scale is a small rigid plate that grows out of an animal's skin to provide protection. In lepidopteran species, scales are plates on the surface of the insect wing, and provide coloration...
s that range from tan to brown or dark brown. Sori
Sorus
A sorus is a cluster of sporangia .In fungi and lichens, the sorus is surrounded by an external layer. In some red algae it may take the form of a depression into the thallus....
occur in two rows, one along each side of the pinnule midvein, and are covered by scale-like indusia.
The subspecies C. c. capensis is native to South Africa
South Africa
The Republic of South Africa is a country in southern Africa. Located at the southern tip of Africa, it is divided into nine provinces, with of coastline on the Atlantic and Indian oceans...
(from the Western Cape
Western Cape
The Western Cape is a province in the south west of South Africa. The capital is Cape Town. Prior to 1994, the region that now forms the Western Cape was part of the much larger Cape Province...
province to KwaZulu-Natal
KwaZulu-Natal
KwaZulu-Natal is a province of South Africa. Prior to 1994, the territory now known as KwaZulu-Natal was made up of the province of Natal and the homeland of KwaZulu....
and Mpumalanga
Mpumalanga
Mpumalanga , is a province of South Africa. The name means east or literally "the place where the sun rises" in Swazi, Xhosa, Ndebele and Zulu. Mpumalanga lies in eastern South Africa, north of KwaZulu-Natal and bordering Swaziland and Mozambique. It constitutes 6.5% of South Africa's land area...
), Swaziland
Swaziland
Swaziland, officially the Kingdom of Swaziland , and sometimes called Ngwane or Swatini, is a landlocked country in Southern Africa, bordered to the north, south and west by South Africa, and to the east by Mozambique...
, Zimbabwe
Zimbabwe
Zimbabwe is a landlocked country located in the southern part of the African continent, between the Zambezi and Limpopo rivers. It is bordered by South Africa to the south, Botswana to the southwest, Zambia and a tip of Namibia to the northwest and Mozambique to the east. Zimbabwe has three...
, Mozambique
Mozambique
Mozambique, officially the Republic of Mozambique , is a country in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west and Swaziland and South Africa to the southwest...
, Malawi
Malawi
The Republic of Malawi is a landlocked country in southeast Africa that was formerly known as Nyasaland. It is bordered by Zambia to the northwest, Tanzania to the northeast, and Mozambique on the east, south and west. The country is separated from Tanzania and Mozambique by Lake Malawi. Its size...
and Tanzania
Tanzania
The United Republic of Tanzania is a country in East Africa bordered by Kenya and Uganda to the north, Rwanda, Burundi, and the Democratic Republic of the Congo to the west, and Zambia, Malawi, and Mozambique to the south. The country's eastern borders lie on the Indian Ocean.Tanzania is a state...
, where it grows in shaded moist forest along rivers and beside waterfalls at an altitude of 360-1820 m. The other subspecies, C. c. polypodioides, grows in southeastern Brazil
Brazil
Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...
in montane forest at 900-2000 m.
The specific epithet
Epithet
An epithet or byname is a descriptive term accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, divinities, objects, and binomial nomenclature. It is also a descriptive title...
capensis refers to the Cape of Good Hope
Cape of Good Hope
The Cape of Good Hope is a rocky headland on the Atlantic coast of the Cape Peninsula, South Africa.There is a misconception that the Cape of Good Hope is the southern tip of Africa, because it was once believed to be the dividing point between the Atlantic and Indian Oceans. In fact, the...
in South Africa, where the type
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
specimen was collected.
C. capensis is relatively easy to grow if provided with a semi-shady, moist and sheltered environment. It can likely tolerate several degrees of frost.