Engrailed
Encyclopedia
The Engrailed and Small Engrailed (Ectropis crepuscularia) are moth
Moth
A moth is an insect closely related to the butterfly, both being of the order Lepidoptera. Moths form the majority of this order; there are thought to be 150,000 to 250,000 different species of moth , with thousands of species yet to be described...

s of the family Geometridae. They are distributed across most of Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

. There is an on-going debate as to whether they make up one species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

, or whether E. crepuscularia actually refers only to the Small Engrailed, with the Engrailed proper being separable as E. bistortata.

The ground colour of the wings is buff or grey variably marked with darker fascia
Fascia
A fascia is a layer of fibrous tissue that permeates the human body. A fascia is a connective tissue that surrounds muscles, groups of muscles, blood vessels, and nerves, binding those structures together in much the same manner as plastic wrap can be used to hold the contents of sandwiches...

. The darker markings are not usually as strong as in the rather similar Willow Beauty
Willow Beauty
| image = Peribatodes.rhomboidaria.7197.jpg| image_width = 240px| image_caption = Adult male from Dresden | regnum = Animalia| phylum = Arthropoda| classis = Insecta| ordo = Lepidoptera| zoodivisio = Ditrysia| familia = Geometridae...

. Melanic
Melanism
Melanism is an undue development of dark-colored pigment in the skin or its appendages, and the opposite of albinism. It is also the medical term for black jaundice.The word is deduced from the , meaning black pigment....

 forms occur fairly frequently. The wingspan
Wingspan
The wingspan of an airplane or a bird, is the distance from one wingtip to the other wingtip. For example, the Boeing 777 has a wingspan of about ; and a Wandering Albatross caught in 1965 had a wingspan of , the official record for a living bird.The term wingspan, more technically extent, is...

 is 38–45 mm. One or two broods are produced each year. In the British Isles the adults can be seen at any time between March and August; this time range may vary in other parts of this moth's range. The species flies at night and is attracted to light.

The greyish caterpillar
Caterpillar
Caterpillars are the larval form of members of the order Lepidoptera . They are mostly herbivorous in food habit, although some species are insectivorous. Caterpillars are voracious feeders and many of them are considered to be pests in agriculture...

 is truly polyphagous, feeding on a huge range of plant
Plant
Plants are living organisms belonging to the kingdom Plantae. Precise definitions of the kingdom vary, but as the term is used here, plants include familiar organisms such as trees, flowers, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The group is also called green plants or...

s (see list below). The species overwinters as a pupa
Pupa
A pupa is the life stage of some insects undergoing transformation. The pupal stage is found only in holometabolous insects, those that undergo a complete metamorphosis, going through four life stages; embryo, larva, pupa and imago...

.

Engrailed/Small Engrailed

There is lots of debate as to whether Small Engrailed (Ectropis crepuscularia) and The Engrailed (Ectropis bistortata) are the same species, despite DNA profiling confirming they are conspecific.

Recorded food plants

  • Acer, Maple
    Maple
    Acer is a genus of trees or shrubs commonly known as maple.Maples are variously classified in a family of their own, the Aceraceae, or together with the Hippocastanaceae included in the family Sapindaceae. Modern classifications, including the Angiosperm Phylogeny Group system, favour inclusion in...

  • Aconitum
    Aconitum
    Aconitum , known as aconite, monkshood, wolfsbane, leopard's bane, women's bane, Devil's helmet or blue rocket, is a genus of over 250 species of flowering plants belonging to the buttercup family .-Overview:These herbaceous perennial plants are chiefly natives of the mountainous parts of the...

    , Monkshood
  • Alnus, Alder
    Alder
    Alder is the common name of a genus of flowering plants belonging to the birch family . The genus comprises about 30 species of monoecious trees and shrubs, few reaching large size, distributed throughout the North Temperate Zone and in the Americas along the Andes southwards to...

  • Aquilegia
    Aquilegia
    Aquilegia is a genus of about 60-70 species of perennial plants that are found in meadows, woodlands, and at higher altitudes throughout the Northern Hemisphere, known for the spurred petals of their flowers.-Etymology:The genus name Aquilegia is derived from the Latin word for eagle , because...

    , Columbine
  • Betula, Birch
    Birch
    Birch is a tree or shrub of the genus Betula , in the family Betulaceae, closely related to the beech/oak family, Fagaceae. The Betula genus contains 30–60 known taxa...

  • Calluna vulgaris, Heather
  • Camellia japonica
    Camellia japonica
    The Japanese Camellia is one of the best known species of camellia. Sometimes called the rose of winter, it is a member of the Theaceae family or tea family. It is a flowering shrub or a small tree native to Japan, Korea and China. It is the official state flower of Alabama.-Description:In the...

    , Japanese Camellia
  • Castanea, Chestnut
    Chestnut
    Chestnut , some species called chinkapin or chinquapin, is a genus of eight or nine species of deciduous trees and shrubs in the beech family Fagaceae, native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce.-Species:The chestnut belongs to the...

  • Centaurea
  • Cirsium arvense
    Cirsium arvense
    Cirsium arvense is a species of Cirsium, native throughout Europe and northern Asia, and widely introduced elsewhere. The standard English name in its native area is Creeping Thistle.-Alternate names:...

    , Creeping Thistle
  • Cornus, Dogwood
    Dogwood
    The genus Cornus is a group of about 30-60 species of woody plants in the family Cornaceae, commonly known as dogwoods. Most dogwoods are deciduous trees or shrubs, but a few species are nearly herbaceous perennial subshrubs, and a few of the woody species are evergreen...

  • Daphniphyllum
    Daphniphyllum
    Daphniphyllum is a genus of flowering plants in the family Daphniphyllaceae, including about 25 species, all evergreen shrubs and trees native to east and southeast Asia...

  • Diervilla, Bush Honeysuckle
  • Frangula, Alder Buckthorn
    Buckthorn
    The Buckthorns are a genus of about 100 species of shrubs or small trees from 1-10 m tall , in the buckthorn family Rhamnaceae...

  • Fraxinus, Ash
    Ash tree
    Fraxinus is a genus flowering plants in the olive and lilac family, Oleaceae. It contains 45-65 species of usually medium to large trees, mostly deciduous though a few subtropical species are evergreen. The tree's common English name, ash, goes back to the Old English æsc, while the generic name...

  • Genista, Broom
    Broom (shrub)
    Brooms are a group of evergreen, semi-evergreen, and deciduous shrubs in the subfamily Faboideae of the legume family Fabaceae, mainly in the three genera Chamaecytisus, Cytisus and Genista, but also in many other small genera . All genera in this group are from the tribe Genisteae...

  • Glycine
    Glycine (plant)
    Glycine is a genus in the bean family Fabaceae. The best known species is the soybean . While the majority of the species are found only in Australia, the soybean's native range is in East Asia. A few species extend from Australia to East Asia Glycine is a genus in the bean family Fabaceae. The...

    , Soybean
  • Hypericum maculatum
    Hypericum maculatum
    Hypericum maculatum Crantz is a species of perennial herbaceous plants in the genus Hypericum, native to Europe and Western Asia and occurring in meadows....

    , Imperforate St. John's Wort
  • Ilex, Holly
    Holly
    Ilex) is a genus of 400 to 600 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. The species are evergreen and deciduous trees, shrubs, and climbers from tropics to temperate zones world wide....

  • Juglans, Walnut
    Walnut
    Juglans is a plant genus of the family Juglandaceae, the seeds of which are known as walnuts. They are deciduous trees, 10–40 meters tall , with pinnate leaves 200–900 millimetres long , with 5–25 leaflets; the shoots have chambered pith, a character shared with the wingnuts , but not the hickories...

  • Larix, Larch
    Larch
    Larches are conifers in the genus Larix, in the family Pinaceae. Growing from 15 to 50m tall, they are native to much of the cooler temperate northern hemisphere, on lowlands in the north and high on mountains further south...

  • Lindera
    Lindera
    Lindera is a genus of about 80-100 species of flowering plants in the family Lauraceae, mostly native to eastern Asia but with three species in eastern North America. The species are shrubs and small trees; common names include Spicewood, Spicebush, and Benjamin Bush.-Overview:They are dioecious,...

    , Spice Bush
  • Lonicera, Honeysuckle
    Honeysuckle
    Honeysuckles are arching shrubs or twining vines in the family Caprifoliaceae, native to the Northern Hemisphere. There are about 180 species of honeysuckle, 100 of which occur in China; Europe, India and North America have only about 20 native species each...

  • Lythrum salicaria, Purple loosestrife
    Purple loosestrife
    Lythrum salicaria is a flowering plant belonging to the family Lythraceae, native to Europe, Asia, northwest Africa, and southeastern Australia. It should not be confused with other plants sharing the name loosestrife that are members of the family Primulaceae...

  • Malus
    Malus
    Malus , the apples, are a genus of about 30–35 species of small deciduous trees or shrubs in the family Rosaceae. Other studies go as far as 55 species including the domesticated Orchard Apple, or Table apple as it was formerly called...

    , Apple
  • Philadelphus, Mock-orange
    Mock-orange
    Mock-orange is a genus of about 60 species of shrubs from 1 to 6 m tall, native to North America, Central America, Asia and in southeast Europe....

  • Picea, Spruce
    Spruce
    A spruce is a tree of the genus Picea , a genus of about 35 species of coniferous evergreen trees in the Family Pinaceae, found in the northern temperate and boreal regions of the earth. Spruces are large trees, from tall when mature, and can be distinguished by their whorled branches and conical...

  • Pieris
    Pieris (plant)
    Pieris is a genus of seven species of shrubs in the family Ericaceae, native to mountain regions of eastern and southern Asia, eastern North America and Cuba. Known commonly as andromedas or fetterbushes, they are broad-leaved evergreen shrubs growing to 1–6 m tall and wide...

  • Pinus, Pine
    Pine
    Pines are trees in the genus Pinus ,in the family Pinaceae. They make up the monotypic subfamily Pinoideae. There are about 115 species of pine, although different authorities accept between 105 and 125 species.-Etymology:...

  • Plectranthus
    Plectranthus
    Plectranthus, with some 350 species, is a genus of warm-climate plants occurring largely in the southern hemisphere, in sub-Saharan Africa, Madagascar, India and the Indonesian archipelago down to Australia and some Pacific islands. It is closely related to Solenostemon, sometimes known as the...

  • Populus, Poplar
    Poplar
    Populus is a genus of 25–35 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar , aspen, and cottonwood....

  • Pseudotsuga, Douglas-fir
    Douglas-fir
    Douglas-fir is one of the English common names for evergreen coniferous trees of the genus Pseudotsuga in the family Pinaceae. Other common names include Douglas tree, and Oregon pine. There are five species, two in western North America, one in Mexico, and two in eastern Asia...

  • Quercus, Oak
    Oak
    An oak is a tree or shrub in the genus Quercus , of which about 600 species exist. "Oak" may also appear in the names of species in related genera, notably Lithocarpus...

  • Ribes rubrum, Redcurrant
    Redcurrant
    The redcurrant , Ribes rubrum, is a member of the genus Ribes in the gooseberry family Grossulariaceae, native to parts of western Europe...

  • Rosa, Rose
    Rose
    A rose is a woody perennial of the genus Rosa, within the family Rosaceae. There are over 100 species. They form a group of erect shrubs, and climbing or trailing plants, with stems that are often armed with sharp prickles. Flowers are large and showy, in colours ranging from white through yellows...

  • Rubus idaeus, Raspberry
    Raspberry
    The raspberry or hindberry is the edible fruit of a multitude of plant species in the genus Rubus, most of which are in the subgenus Idaeobatus; the name also applies to these plants themselves...

  • Rumex
    Rumex
    The docks and sorrels, genus Rumex L., are a genus of about 200 species of annual, biennial and perennial herbs in the buckwheat family Polygonaceae....

    , Dock
  • Salix, Willow
    Willow
    Willows, sallows, and osiers form the genus Salix, around 400 species of deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions of the Northern Hemisphere...

  • Sambucus, Elder
    Elderberry
    Sambucus is a genus of between 5 and 30 species of shrubs or small trees in the moschatel family, Adoxaceae. It was formerly placed in the honeysuckle family, Caprifoliaceae, but was reclassified due to genetic evidence...

  • Shepherdia canadensis, Canada Buffaloberry
    Canada Buffaloberry
    The Canada Buffaloberry , also known as Russet Buffaloberry, Soopolallie, Soapberry, or Foamberry, is one of a small number of shrubs of the genus Shepherdia bearing edible red berries. One recognized form however bears yellow fruits...

  • Sorbus spp., rowan
    Rowan
    The rowans or mountain-ashes are shrubs or small trees in genus Sorbus of family Rosaceae. They are native throughout the cool temperate regions of the Northern Hemisphere, with the highest species diversity in the mountains of western China and the Himalaya, where numerous apomictic microspecies...

    s
  • Spartina, Cordgrass
    Cordgrass
    Spartina, commonly known as cordgrass or cord-grass, is a genus of 14 species of grasses in the family Poaceae. They are native to the coasts of the Atlantic Ocean in western and southern Europe, northwest and southern Africa, the Americas and the southern Atlantic Ocean islands; one or two species...

  • Thuja
    Thuja
    Thuja is a genus of coniferous trees in the Cupressaceae . There are five species in the genus, two native to North America and three native to eastern Asia...

  • Trifolium, Clover
    Clover
    Clover , or trefoil, is a genus of about 300 species of plants in the leguminous pea family Fabaceae. The genus has a cosmopolitan distribution; the highest diversity is found in the temperate Northern Hemisphere, but many species also occur in South America and Africa, including at high altitudes...

  • Tsuga
    Tsuga
    Tsuga is a genus of conifers in the family Pinaceae. The common name hemlock is derived from a perceived similarity in the smell of its crushed foliage to that of the unrelated plant poison hemlock....

    , Hemlock
  • Ulmus, Elm
    Elm
    Elms are deciduous and semi-deciduous trees comprising the genus Ulmus in the plant family Ulmaceae. The dozens of species are found in temperate and tropical-montane regions of North America and Eurasia, ranging southward into Indonesia. Elms are components of many kinds of natural forests...

  • Vaccinium
    Vaccinium
    Vaccinium is a genus of shrubs or dwarf shrubs in the plant Family Ericaceae. The fruit of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry or whortleberry, lingonberry or cowberry, and huckleberry...

  • Zanthoxylum
    Zanthoxylum
    Zanthoxylum is a genus of about 250 species of deciduous and evergreen trees and shrubs in the citrus or rue family, Rutaceae, native to warm temperate and subtropical areas worldwide. Common names include Prickly-ash and Hercules' Club.The fruit of several species are used to make the spice...


External links

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