Eocarcinus praecursor is a Jurassic

Jurassic

The Jurassic is a geologic period and system that extends from about Mya to  Mya, that is, from the end of the Triassic to the beginning of the Cretaceous. The Jurassic constitutes the middle period of the Mesozoic era, also known as the age of reptiles. The start of the period is marked by...

 species of decapod

Decapoda

The decapods or Decapoda are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crayfish, crabs, lobsters, prawns and shrimp. Most decapods are scavengers. It is estimated that the order contains nearly 15,000 species in around 2,700 genera, with...

 crustacean

Crustacean

Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

, sufficiently distinct from its relatives to be placed in its own family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 (Eocarcinidae). Once considered the oldest true crab

Crab

True crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" , or where the reduced abdomen is entirely hidden under the thorax...

, it is now considered to be an early member of the Anomura

Anomura

Anomura is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans includes the word crab, all true crabs are in the sister group to the Anomura, the Brachyura .-Description:The name Anomala reflects the unusual variety of forms in this group;...

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Distribution

It lived during the Pliensbachian

Pliensbachian

The Pliensbachian is an age of the geologic timescale or stage in the stratigraphic column. It is part of the Early or Lower Jurassic epoch or series and spans the time between 189.6 ± 1.5 Ma and 183 ± 1.5 Ma . The Pliensbachian is preceded by the Sinemurian and followed by the Toarcian.The...

 age (Lower Jurassic), and has been found in rocks at two sites in the United Kingdom

United Kingdom

The United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...

 – Mickelton Tunnel (near Aston Magna

Aston Magna

Aston Magna is a hamlet located off Fosse Way in north Gloucestershire between Moreton-in-Marsh and Shipston-on-Stour.- See also :*Corpse roads...

), Gloucestershire

Gloucestershire

Gloucestershire is a county in South West England. The county comprises part of the Cotswold Hills, part of the flat fertile valley of the River Severn, and the entire Forest of Dean....

 and Runswick Bay, Yorkshire

Yorkshire

Yorkshire is a historic county of northern England and the largest in the United Kingdom. Because of its great size in comparison to other English counties, functions have been increasingly undertaken over time by its subdivisions, which have also been subject to periodic reform...

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Description

In many of its characters, it represents a transitional stage

Transitional fossil

A transitional fossil is any fossilized remains of a lifeform that exhibits characteristics of two distinct taxonomic groups. A transitional fossil is the fossil of an organism near the branching point where major individual lineages diverge...

 between the Glypheoidea

Glypheoidea

The Glypheoidea , is a group of lobster-like decapod crustaceans which forms an important part of fossil faunas, such as the Solnhofen limestone. These fossils included taxa such as Glyphea , and Mecochirus, mostly with elongated chelipeds...

 and the Middle Jurassic

Middle Jurassic

The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from 176-161 million years ago. In European lithostratigraphy, rocks of this Middle Jurassic age are called the Dogger....

 crabs in the Prosopidae. Since its ancestors were long-tailed decapods

Decapoda

The decapods or Decapoda are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crayfish, crabs, lobsters, prawns and shrimp. Most decapods are scavengers. It is estimated that the order contains nearly 15,000 species in around 2,700 genera, with...

, and its successors were short-tailed crabs, Eocarcinus has been described as "the lobster who decided to become a crab". Previously considered to be the oldest known true crab

Crab

True crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" , or where the reduced abdomen is entirely hidden under the thorax...

, a 2010 revision concluded that Eocarcinus could not be accommodated among the Brachyura, and was instead transferred to the Anomura

Anomura

Anomura is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans includes the word crab, all true crabs are in the sister group to the Anomura, the Brachyura .-Description:The name Anomala reflects the unusual variety of forms in this group;...

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