Eonatator
Encyclopedia
Eonatator is a genus
of halisaurine mosasaur
from the Upper Cretaceous
of North America
and Europe
. Originally, this taxon
was included within Halisaurus, but was placed in its own genus . Eonatator is presently known from the Smoky Hill Chalk
Member of the Niobrara Chalk Formation (Upper Coniacian
- Lower Campanian
) of Kansas
, from the Eutaw Formation
(Santonian
) and Mooreville Chalk Formation
(Selma Group
; Santonian-Lower Campanian) of Alabama
, and from the Kristianstad Basin of southern Sweden
(late early Campanian). The name Eonatator means "dawn swimmer" (Greek
eos = dawn + Latin
natator = swimmer). At this time it contains only a single species
, E. sternbergii. The species is named in honour of Charles H. Sternberg and his son, Levi, who discovered the type specimen in the Niobrara Chalk during the summer of 1918.
Like many mosasaurs, this genus has a complicated taxonomic history. The type specimen (UPI R 163, Uppsala University Palaeontological Institute, Uppsala
, Sweden), a nearly complete skeleton, was originally referred to the genus Clidastes by Wiman and then to Halisaurus by Russell . Hence, Clidastes sternbergii became Halisaurus sternbergii. However, by the late 1980s, some paleontologists began to suggest that H. sternbergii belonged in its own genus and that Halisaurus was polyphyletic .
Bardet et al. (2005, p. 465 ) diagnose Eonatator sternbergii as follows: "Ambiguous characters: premaxilla
-maxilla
lateral suture ending posterior to 9th maxillary teeth; tail about 40% of the head and trunk length (convergent in mosasaurines); caudal vertebra length greater than width; fewer than four pygal vertebrae; femur
length about twice distal width (convergent in Clidastes). Autapomorphies: parietal
with smooth triangular table extending very far posteriorly, bearing medium-sized circular foramen, located at distance twice its diameter from the frontal-parietal suture, and surrounded anteriorly and posteriorly by two parallel ridges; rounded quadrate with regularly convex tympanic ala; vertebral formula: seven cervicals, 24 dorsals, four pygals, 28 median caudals and at least 41 terminal caudals; humerus
length approximately 2.5x distal width." A cladistic analysis of the Halisaurinae
has indicated that Eonatator is the sister group to the clade
Halisaurus and that it represents one of the most basal of known mosasaur taxa.
Eonatator was among the smaller mosasaurs. The length of the type skeleton, which represents an adult, is only 2.65 meters in length. Like many mosasaurs, it likely fed primarily on fish
and smaller marine reptile
s.
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of halisaurine mosasaur
Mosasaur
Mosasaurs are large extinct marine lizards. The first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764...
from the Upper Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...
of North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
and Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...
. Originally, this taxon
Taxon
|thumb|270px|[[African elephants]] form a widely-accepted taxon, the [[genus]] LoxodontaA taxon is a group of organisms, which a taxonomist adjudges to be a unit. Usually a taxon is given a name and a rank, although neither is a requirement...
was included within Halisaurus, but was placed in its own genus . Eonatator is presently known from the Smoky Hill Chalk
Smoky Hill Chalk
The Smoky Hill Chalk Member of the Niobrara Chalk formation is a Cretaceous conservation Lagerstätte, or fossil rich geological formation, known primarily for its exceptionally well-preserved marine reptiles. The Smoky Hill Chalk Member is the uppermost of the two structural units of the Niobrara...
Member of the Niobrara Chalk Formation (Upper Coniacian
Coniacian
The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous epoch or Upper Cretaceous series and spans the time between 89.3 ± 1 Ma and 85.8 ± 0.7 Ma...
- Lower Campanian
Campanian
The Campanian is, in the ICS' geologic timescale, the fifth of six ages of the Late Cretaceous epoch . The Campanian spans the time from 83.5 ± 0.7 Ma to 70.6 ± 0.6 Ma ...
) of Kansas
Kansas
Kansas is a US state located in the Midwestern United States. It is named after the Kansas River which flows through it, which in turn was named after the Kansa Native American tribe, which inhabited the area. The tribe's name is often said to mean "people of the wind" or "people of the south...
, from the Eutaw Formation
Eutaw Formation
The Eutaw Formation is a geological formation in North America, within the U.S. states of Alabama, Georgia, and Mississippi. The strata date from the late Coniacian to the early Santonian stage of the Late Cretaceous. It consists of the upper Tombigbee Sand Member and an unnamed lower member...
(Santonian
Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous epoch or Upper Cretaceous series. It spans the time between 85.8 ± 0.7 mya and 83.5 ± 0.7 mya...
) and Mooreville Chalk Formation
Mooreville Chalk Formation
The Mooreville Chalk Formation is a geological formation in North America, within the U.S. states of Alabama and Mississippi. The strata date back to the early Santonian to the early Campanian stage of the Late Cretaceous. The chalk was formed by pelagic sediments deposited along the eastern edge...
(Selma Group
Selma Group
The Selma Group is a geological formation in North America, within the U.S. states of Alabama, Mississippi, and Tennessee. The strata date from the Santonian to the Maastrichtian stages of the Late Cretaceous. The group is composed of, in ascending order, the Mooreville Chalk Formation, Demopolis...
; Santonian-Lower Campanian) of Alabama
Alabama
Alabama is a state located in the southeastern region of the United States. It is bordered by Tennessee to the north, Georgia to the east, Florida and the Gulf of Mexico to the south, and Mississippi to the west. Alabama ranks 30th in total land area and ranks second in the size of its inland...
, and from the Kristianstad Basin of southern Sweden
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....
(late early Campanian). The name Eonatator means "dawn swimmer" (Greek
Greek language
Greek is an independent branch of the Indo-European family of languages. Native to the southern Balkans, it has the longest documented history of any Indo-European language, spanning 34 centuries of written records. Its writing system has been the Greek alphabet for the majority of its history;...
eos = dawn + Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...
natator = swimmer). At this time it contains only a single species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...
, E. sternbergii. The species is named in honour of Charles H. Sternberg and his son, Levi, who discovered the type specimen in the Niobrara Chalk during the summer of 1918.
Like many mosasaurs, this genus has a complicated taxonomic history. The type specimen (UPI R 163, Uppsala University Palaeontological Institute, Uppsala
Uppsala
- Economy :Today Uppsala is well established in medical research and recognized for its leading position in biotechnology.*Abbott Medical Optics *GE Healthcare*Pfizer *Phadia, an offshoot of Pharmacia*Fresenius*Q-Med...
, Sweden), a nearly complete skeleton, was originally referred to the genus Clidastes by Wiman and then to Halisaurus by Russell . Hence, Clidastes sternbergii became Halisaurus sternbergii. However, by the late 1980s, some paleontologists began to suggest that H. sternbergii belonged in its own genus and that Halisaurus was polyphyletic .
Bardet et al. (2005, p. 465 ) diagnose Eonatator sternbergii as follows: "Ambiguous characters: premaxilla
Premaxilla
The incisive bone is the portion of the maxilla adjacent to the incisors. It is a pair of small cranial bones at the very tip of the jaws of many animals, usually bearing teeth, but not always. They are connected to the maxilla and the nasals....
-maxilla
Maxilla
The maxilla is a fusion of two bones along the palatal fissure that form the upper jaw. This is similar to the mandible , which is also a fusion of two halves at the mental symphysis. Sometimes The maxilla (plural: maxillae) is a fusion of two bones along the palatal fissure that form the upper...
lateral suture ending posterior to 9th maxillary teeth; tail about 40% of the head and trunk length (convergent in mosasaurines); caudal vertebra length greater than width; fewer than four pygal vertebrae; femur
Femur
The femur , or thigh bone, is the most proximal bone of the leg in tetrapod vertebrates capable of walking or jumping, such as most land mammals, birds, many reptiles such as lizards, and amphibians such as frogs. In vertebrates with four legs such as dogs and horses, the femur is found only in...
length about twice distal width (convergent in Clidastes). Autapomorphies: parietal
Parietal
Parietal may refer to:*Parietal placentation*Parietal lobe of the brain*Parietal bone of the skull*Parietal scales of a snake lie in the general region of the parietal bone*Parietal cell in the stomach*Parietal pleura...
with smooth triangular table extending very far posteriorly, bearing medium-sized circular foramen, located at distance twice its diameter from the frontal-parietal suture, and surrounded anteriorly and posteriorly by two parallel ridges; rounded quadrate with regularly convex tympanic ala; vertebral formula: seven cervicals, 24 dorsals, four pygals, 28 median caudals and at least 41 terminal caudals; humerus
Humerus
The humerus is a long bone in the arm or forelimb that runs from the shoulder to the elbow....
length approximately 2.5x distal width." A cladistic analysis of the Halisaurinae
Halisaurinae
Halisaurinae is a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates.Bardet et al. diagnosed the Halsaurinae as all mosasaurs more closely related to Halisaurus than Moanasaurus...
has indicated that Eonatator is the sister group to the clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...
Halisaurus and that it represents one of the most basal of known mosasaur taxa.
Eonatator was among the smaller mosasaurs. The length of the type skeleton, which represents an adult, is only 2.65 meters in length. Like many mosasaurs, it likely fed primarily on fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...
and smaller marine reptile
Reptile
Reptiles are members of a class of air-breathing, ectothermic vertebrates which are characterized by laying shelled eggs , and having skin covered in scales and/or scutes. They are tetrapods, either having four limbs or being descended from four-limbed ancestors...
s.
Further reading
- Lindgren J, Siverson M. 2005. Halisaurus sternbergii, a small mosasaur with an intercontinental distribution. Journal of Paleontology 79 (4): 763–773.