Europolemur klatti
Encyclopedia
Europolemur klatti was an medium to large size adapiformes
primate
that lived on the continent
of Europe
from the middle to early Eocene
. The most recent relative to this species,appearing to be so from the Geiseltal localities of the DDR, is a recently discovered primate
, Mahgarita stevensi, whose type specimen is about the size of Lepilemur leucopus. This relationship to M. stevensi suggests that eastern North America
was the possible homeland of lemuriformes with close European phylogenetic ties during the Eocene. Characteristic of most adapines are the reduced or absence of a paraconid and morphology of the paracristid. These and a few other features are synapomorphies that were used to link E. klatti with Leptadapis priscus and Microadapis sciureus, as well as Smilodectes.
and there is evidence that supports that E. klatti may have had nail
s instead of claw
s. This insinuates that stabilizing the tips of the digits and hand must have in some way been an important function for them and their lifestyle in their habitat. Relative to the forearm, the hand of E. klatti was large which may be related to vertical climbing or posture. The shape of the calcaneus resembles that found in Smilodectes
and Notharctus
and E. klatti had an average body mass of 1.7 kilograms.
s while the adults only had three premolars. One of the most distinguishing characteristics of the genus Europolemur is the lack of a metaconule. The dental anatomy of their genus is described in more detail by Franzen as consisting of "upper canines big and pointed; upper molars without postflexus; postprotocrista prominent; no metaconulus; M3 smaller and shorter than M2; P4 much shorter than broad, with a weak parastyle; P4 with a small and unicuspid talonid and a metaconid present to absent; protocristid of M nearly transversely oriented. Protoconid of P3 little higher than that of P4." ^2
Adapiformes
Adapiformes are an extinct group of primitive primates.The adapiformes radiated throughout much of the northern continental mass, reaching as far south as northern Africa and tropical Asia. The adapiformes existed from the Eocene to the Miocene epoch...
primate
Primate
A primate is a mammal of the order Primates , which contains prosimians and simians. Primates arose from ancestors that lived in the trees of tropical forests; many primate characteristics represent adaptations to life in this challenging three-dimensional environment...
that lived on the continent
Continent
A continent is one of several very large landmasses on Earth. They are generally identified by convention rather than any strict criteria, with seven regions commonly regarded as continents—they are : Asia, Africa, North America, South America, Antarctica, Europe, and Australia.Plate tectonics is...
of Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...
from the middle to early Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...
. The most recent relative to this species,appearing to be so from the Geiseltal localities of the DDR, is a recently discovered primate
Primate
A primate is a mammal of the order Primates , which contains prosimians and simians. Primates arose from ancestors that lived in the trees of tropical forests; many primate characteristics represent adaptations to life in this challenging three-dimensional environment...
, Mahgarita stevensi, whose type specimen is about the size of Lepilemur leucopus. This relationship to M. stevensi suggests that eastern North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
was the possible homeland of lemuriformes with close European phylogenetic ties during the Eocene. Characteristic of most adapines are the reduced or absence of a paraconid and morphology of the paracristid. These and a few other features are synapomorphies that were used to link E. klatti with Leptadapis priscus and Microadapis sciureus, as well as Smilodectes.
Morphology
Europolemur klatti is part of a group of long-digited fossils, and most likely approximates early euprimate hand proportions. E. klatti has a grasping halluxHallux
In tetrapods, the hallux is the innermost toe of the foot. Despite its name it may not be the longest toe on the foot of some individuals...
and there is evidence that supports that E. klatti may have had nail
Nail (anatomy)
A nail is a horn-like envelope covering the dorsal aspect of the terminal phalanges of fingers and toes in humans, most non-human primates, and a few other mammals. Nails are similar to claws, which are found on numerous other animals....
s instead of claw
Claw
A claw is a curved, pointed appendage, found at the end of a toe or finger in most mammals, birds, and some reptiles. However, the word "claw" is also often used in reference to an invertebrate. Somewhat similar fine hooked structures are found in arthropods such as beetles and spiders, at the end...
s. This insinuates that stabilizing the tips of the digits and hand must have in some way been an important function for them and their lifestyle in their habitat. Relative to the forearm, the hand of E. klatti was large which may be related to vertical climbing or posture. The shape of the calcaneus resembles that found in Smilodectes
Smilodectes
Smilodectes is an extinct genus of primate that lived in Wyoming. It possesses a post-orbital bar and grasping thumbs and toes. Smilodectes has a small cranium size and the foramen magnum was located at the back of the skull, on the occipatal bone....
and Notharctus
Notharctus
Notharctus was an early primate that inhabited Europe and North America 50 million years ago. Modern lemurs evolved from primates similar to this genus....
and E. klatti had an average body mass of 1.7 kilograms.
Dentition
In 1995, two isolated upper molars belonging to E. klatti (pictured in figure 1 under external link 1) were found in an old lake deposit during excavations done by the "Naturhistorisches Museum Mainz/Landessammlung fur Naturkunde Rheinland-Pfalz". The museum determined that the molars (as well as a mandible with nearly complete dentition belonging to another cercamoiines, Periconodon) were representative of the first primates from the Middle Eocene Eckfeld maar in Southwest Eifel, Germany ^2. E. klatti has a dental formula of 2:1:3:3 (Franzen, 1987; cited in Martin, 1990) and the milk dentition of this species consisted of four premolarPremolar
The premolar teeth or bicuspids are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant, making eight premolars total in the mouth. They have at least two cusps. Premolars can be considered as a 'transitional tooth' during chewing, or...
s while the adults only had three premolars. One of the most distinguishing characteristics of the genus Europolemur is the lack of a metaconule. The dental anatomy of their genus is described in more detail by Franzen as consisting of "upper canines big and pointed; upper molars without postflexus; postprotocrista prominent; no metaconulus; M3 smaller and shorter than M2; P4 much shorter than broad, with a weak parastyle; P4 with a small and unicuspid talonid and a metaconid present to absent; protocristid of M nearly transversely oriented. Protoconid of P3 little higher than that of P4." ^2