Gamebird hybrids
Encyclopedia
Gamebird hybrids are the result of crossing species of game birds, including ducks, with each other and with domestic poultry. These hybrid species may occur both naturally or though the intervention of man.

Charles Darwin
Charles Darwin
Charles Robert Darwin FRS was an English naturalist. He established that all species of life have descended over time from common ancestry, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection.He published his theory...

 described hybrids of game birds and domestic fowl in The Variation of Animals And Plants Under Domestication
Domestication
Domestication or taming is the process whereby a population of animals or plants, through a process of selection, becomes accustomed to human provision and control. In the Convention on Biological Diversity a domesticated species is defined as a 'species in which the evolutionary process has been...

:

Mr. Hewitt, who has had great experience in crossing tame cock-pheasants with fowls belonging to five breeds, gives as the character of all 'extraordinary wildness' (13/42. 'The Poultry Book' by Tegetmeier 1866 pages 165, 167.); but I have myself seen one exception to this rule. Mr. S. J. Salter (13/43. 'Natural History Review' 1863 April page 277.) who raised a large number of hybrids from a bantam-hen by Gallus sonneratii, states that 'all were exceedingly wild.' [...] utterly sterile
Infertility
Infertility primarily refers to the biological inability of a person to contribute to conception. Infertility may also refer to the state of a woman who is unable to carry a pregnancy to full term...

 male hybrids from the pheasant and the fowl act in the same manner, "their delight being to watch when the hens leave their nests, and to take on themselves the office of a sitter." (13/57. 'Cottage Gardener' 1860 page 379.) [...] Mr. Hewitt gives it as a general rule with fowls, that crossing the breed increases their size. He makes this remark after stating that hybrids from the pheasant and fowl are considerably larger than either progenitor: so again, hybrids from the male golden pheasant and female common pheasant "are of far larger size than either parent-bird.' (17/39. Ibid 1866 page 167; and 'Poultry Chronicle' volume 3 1855 page 15.)"

Pheasant and Grouse hybrids

Hybrids have been obtained between the "ornamental" species of pheasants e.g. Lady Amherst's, Silver and Reeves Pheasants.

Natural pheasant
Pheasant
Pheasants refer to some members of the Phasianinae subfamily of Phasianidae in the order Galliformes.Pheasants are characterised by strong sexual dimorphism, males being highly ornate with bright colours and adornments such as wattles and long tails. Males are usually larger than females and have...

 and grouse
Grouse
Grouse are a group of birds from the order Galliformes. They are sometimes considered a family Tetraonidae, though the American Ornithologists' Union and many others include grouse as a subfamily Tetraoninae in the family Phasianidae...

 hybrids have been reported:
  • Capercaillie or Wood Grouse (Tetrao urogallus) and Black Grouse
    Black Grouse
    The Black Grouse or Blackgame is a large bird in the grouse family. It is a sedentary species, breeding across northern Eurasia in moorland and bog areas near to woodland, mostly boreal...

     (Lyrorus tetrix) in the UK
  • Dusky or Blue Grouse (Dendragapus obscurus) and Common Pheasant
    Common Pheasant
    The Common Pheasant , is a bird in the pheasant family . It is native to Georgia and has been widely introduced elsewhere as a game bird. In parts of its range, namely in places where none of its relatives occur such as in Europe , it is simply known as the "pheasant"...

     (Phasianus colchicus) near Portland, Oregon, USA
  • Sharp-tailed Grouse (Pedioecetes phasianellus) and Prairie Chicken (Tympanuchus cupido)
  • Willow Ptarmigan (Lagopus lagopus) and Spruce Grouse (Canachites canadensis)
  • Dusky or Blue Grouse (Dendragapus obscurus) and Sharp-tailed grouse (Pedioecetes phasianellus) at Osoyoos, British Columbia, 1906 (Dr. John W. Aldrich)
  • Common Pheasant
    Common Pheasant
    The Common Pheasant , is a bird in the pheasant family . It is native to Georgia and has been widely introduced elsewhere as a game bird. In parts of its range, namely in places where none of its relatives occur such as in Europe , it is simply known as the "pheasant"...

     (Phasianus colchicus) and Prairie Chicken (Tympanuchus cupido) at Ellendale, North Dakota, 1933

Chicken Hybrids

Charles Darwin mentioned crosses between domestic fowl and pheasants in Origin of Species 
[...] from observations communicated to me by Mr. Hewitt, who has had great experience in hybridising pheasants and fowls
and later in The Variation of Animals and Plants under Domestication
The Variation of Animals and Plants under Domestication
The Variation of Animals and Plants under Domestication is a book written by Charles Darwin that was first published in January 1868.A large proportion of the book contains detailed information on the domestication of animals and plants but it also contains in Chapter XXVII a description of...

 (top of this page), where he mentioned effeminate behaviour in the male hybrids.

In her book Bird Hybrids, A. P. Gray lists numerous crosses between chicken
Chicken
The chicken is a domesticated fowl, a subspecies of the Red Junglefowl. As one of the most common and widespread domestic animals, and with a population of more than 24 billion in 2003, there are more chickens in the world than any other species of bird...

s (Gallus gallus) and other types of fowl. Domestic fowl can be crossed, and produce fertile offspring, with Silver Pheasant
Silver Pheasant
The Silver Pheasant is a species of pheasant found in forests, mainly in mountains, of mainland Southeast Asia, and eastern and southern China, with introduced populations in Hawaii and various locations in the US mainland. The male is black and white, while the female is mainly brown...

s, Red Junglefowl
Red Junglefowl
The Red Junglefowl is a tropical member of the Pheasant family. They are thought to be ancestors of the domestic chicken with some hybridisation with the Grey Junglefowl...

 and Green Junglefowl
Green Junglefowl
The Green Junglefowl, Gallus varius also known as Javan Junglefowl, Forktail or Green Javanese Junglefowl is a medium-sized, up to 75 cm long, bird in the pheasant family Phasianidae...

. They have also produced hybrids with peafowl
Peafowl
Peafowl are two Asiatic species of flying birds in the genus Pavo of the pheasant family, Phasianidae, best known for the male's extravagant eye-spotted tail, which it displays as part of courtship. The male is called a peacock, the female a peahen, and the offspring peachicks. The adult female...

, chachalaca
Chachalaca
Chachalacas are mainly brown birds from the genus Ortalis. These cracids are found in wooded habitats in far southern United States , Mexico, and Central and South America. They are social, can be very noisy and often remain fairly common even near humans, as their relatively small size makes them...

s, Capercaillie
Capercaillie
The Western Capercaillie , also known as the Wood Grouse, Heather Cock or Capercaillie , is the largest member of the grouse family, reaching over 100 cm in length and 6.7 kg in weight. The largest one ever recorded in captivity had a weight of 7.2 kg....

, Grouse
Grouse
Grouse are a group of birds from the order Galliformes. They are sometimes considered a family Tetraonidae, though the American Ornithologists' Union and many others include grouse as a subfamily Tetraoninae in the family Phasianidae...

, Quail
Quail
Quail is a collective name for several genera of mid-sized birds generally considered in the order Galliformes. Old World quail are found in the family Phasianidae, while New World quail are found in the family Odontophoridae...

, curassows
Cracidae
The chachalacas, guans and curassows are birds in the family Cracidae.These are species of tropical and subtropical Central and South America. One species, the Plain Chachalaca, just reaches southernmost Texas in the USA...

, pheasant
Pheasant
Pheasants refer to some members of the Phasianinae subfamily of Phasianidae in the order Galliformes.Pheasants are characterised by strong sexual dimorphism, males being highly ornate with bright colours and adornments such as wattles and long tails. Males are usually larger than females and have...

s and guans
Guan (bird)
The guans are a number of bird genera which make up the largest group in the family Cracidae. They are found mainly in northern South America, southern Central America, and a few adjacent Caribbean islands...

.

Domestic fowl have been crossed with guineafowl
Guineafowl
The guineafowl are a family of birds in the Galliformes order, although some authorities include the guineafowl as a subfamily, Numidinae, of the family Phasianidae...

 and also with Common Pheasant
Common Pheasant
The Common Pheasant , is a bird in the pheasant family . It is native to Georgia and has been widely introduced elsewhere as a game bird. In parts of its range, namely in places where none of its relatives occur such as in Europe , it is simply known as the "pheasant"...

 (Phasianus colchicus). Domestic Fowl/Pheasant hybrids have also occurred naturally. Domestic chickens and quail (Coturnix coturnix subsp. japonica) have been hybridised using artificial insemination
Artificial insemination
Artificial insemination, or AI, is the process by which sperm is placed into the reproductive tract of a female for the purpose of impregnating the female by using means other than sexual intercourse or natural insemination...

.

The Peafowl
Peafowl
Peafowl are two Asiatic species of flying birds in the genus Pavo of the pheasant family, Phasianidae, best known for the male's extravagant eye-spotted tail, which it displays as part of courtship. The male is called a peacock, the female a peahen, and the offspring peachicks. The adult female...

 (Pavo cristatus) from Asia and the Common Guinea fowl (Numida meleagris) from Africa have been crossed.

Chicken and Turkey hybrids

There have been attempted crosses between domestic turkeys (Meleagris gallapavo) and chickens. According to Gray in 12 studies no hybrids hatched. Other reports found only a few fertile eggs were produced and very few resulted in advance embryos. According to Olsen, 23 hybrids were obtained from 302 embryos which resulted from 2132 eggs. Dark Cornish cockerels and Rhode Island Red cockerels successfully fertilised turkey eggs. Harada & Buss reported hybridisation experiments between Beltsville Small White Turkeys and two strains of chickens. When male chickens inseminated
Insemination
Insemination is the deliberate introduction of sperm into the uterus of a mammal or the oviduct of an oviparous animal for the objective of impregnating a female for reproduction...

 female turkeys, both male and female embryos form, but the males are much less viable and usually die in the early stages of development. When male turkeys inseminated female chickens, no hybrids resulted although the unfertilised chicken eggs began to divide. According to Olson turkey-chicken crosses produced all males.

A supposed Turkey x Pheasant hybrid was reported by Edwards in 1761.

Duck hybrids

Charles Darwin
Charles Darwin
Charles Robert Darwin FRS was an English naturalist. He established that all species of life have descended over time from common ancestry, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection.He published his theory...

 also described duck hybrids in The Variation of Animals And Plants Under Domestication:
Hybrids are often raised between the common and musk duck, and I have been assured by three persons, who have kept these crossed birds, that they were not wild; but Mr. Garnett (13/45. As stated by Mr. Orton in his 'Physiology of Breeding' page 12.) observed that his hybrids were wild, and exhibited 'migratory propensities' of which there is not a vestige in the common or musk duck.


Hybrids between mallard
Mallard
The Mallard , or Wild Duck , is a dabbling duck which breeds throughout the temperate and subtropical Americas, Europe, Asia, and North Africa, and has been introduced to New Zealand and Australia....

 ducks and Aylesbury ducks (a white domestic breed derived from the mallard) are frequently seen in British parks where the two types are present. The hybrids often resemble a dark coloured mallard with a white breast. Mallard ducks also hybridise with the Muscovy duck producing pied offspring.

Hybrids between the Ruddy Duck
Ruddy Duck
The Ruddy Duck is a small stiff-tailed duck.Their breeding habitat is marshy lakes and ponds throughout much of North America, and in South America in the Andes. They nest in dense marsh vegetation near water. The female builds her nest out of grass, locating it in tall vegetation to hide it from...

 and White-headed Duck
White-headed Duck
The White-headed Duck is a small stiff-tailed duck.Adult males have a grey and reddish body, a blue bill and a largely white head with a black cap and neck. Adult females have a grey-brown body with a white face and a darker bill, cap and a cheek stripe.This duck breeds in Spain and North Africa,...

 are undesirable in parts of Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 http://www.wwt.org.uk/threatsp/hybrid/spain.htm http://www.rspb.org.uk/policy/species/ruddyduck/problem.asp where the introduced Ruddy Duck has bred with native White-headed ducks. The increasing number of Ruddy ducks and hybrids threatens the existence of the White-headed ducks, resulting in shooting campaigns to remove the introduced species. This is controversial as some believe that nature should be allowed to take its course, even though this favours the more successful introduced species
Introduced species
An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...

.

Hybrid duck
Duck
Duck is the common name for a large number of species in the Anatidae family of birds, which also includes swans and geese. The ducks are divided among several subfamilies in the Anatidae family; they do not represent a monophyletic group but a form taxon, since swans and geese are not considered...

s of the genus Aythya
Aythya
Aythya is a genus of diving ducks. It has twelve described species.Aythya shihuibas was described from the Late Miocene of China. An undescribed prehistoric species is known only from Early Pleistocene fossil remains found at Dursunlu, Turkey; it might however be referrable to a paleosubspecies of...

, include birds which are a mixture of Tufted Duck
Tufted Duck
The Tufted Duck, Aythya fuligula, is a medium-sized diving duck with a population of close to one million birds.- Description :The adult male is all black except for white flanks and a blue-grey bill. It has an obvious head tuft that gives the species its name.The adult female is brown with paler...

, Greater Scaup
Greater Scaup
The Greater Scaup , just Scaup in Europe, or colloquially known as "Bluebill", for its bright blue bill, is small compared to other diving ducks, however it is larger than the closely related Lesser Scaup...

, Pochard
Pochard
The Common Pochard, Aythya ferina, is a medium-sized diving duck.The adult male has a long dark bill with a grey band, a red head and neck, a black breast, red eyes and a grey back. The adult female has a brown head and body and a narrower grey bill band. The triangular head shape is distinctive...

, Ferruginous Duck
Ferruginous Duck
The Ferruginous Duck is a medium-sized diving duck from Eurasia. The species is known colloquially by birders as "Fudge Duck"....

 and Ring-necked Duck
Ring-necked Duck
The Ring-necked Duck is a smaller diving duck from North America.The adult male is similar in color pattern to the Eurasian Tufted Duck, its relative. It has a grey bill with a white band, a shiny purple head, a white breast, yellow eyes and a dark grey back...



List of duck hybrids:
  • Northern pintail
    Northern Pintail
    The Pintail or Northern Pintail is a widely occurring duck which breeds in the northern areas of Europe, Asia and North America. It is strongly migratory and winters south of its breeding range to the equator...

     and Mallard
    Mallard
    The Mallard , or Wild Duck , is a dabbling duck which breeds throughout the temperate and subtropical Americas, Europe, Asia, and North Africa, and has been introduced to New Zealand and Australia....

  • Ruddy duck
    Ruddy Duck
    The Ruddy Duck is a small stiff-tailed duck.Their breeding habitat is marshy lakes and ponds throughout much of North America, and in South America in the Andes. They nest in dense marsh vegetation near water. The female builds her nest out of grass, locating it in tall vegetation to hide it from...

     and White-headed duck
    White-headed Duck
    The White-headed Duck is a small stiff-tailed duck.Adult males have a grey and reddish body, a blue bill and a largely white head with a black cap and neck. Adult females have a grey-brown body with a white face and a darker bill, cap and a cheek stripe.This duck breeds in Spain and North Africa,...

  • Ruddy shelduck
    Ruddy Shelduck
    The Ruddy Shelduck, Tadorna ferruginea, is a member of the duck, goose and swan family Anatidae. It is in the shelduck subfamily Tadorninae...

     and Shelduck
    Shelduck
    The shelducks, genus Tadorna, are a group of large birds in the Tadorninae subfamily of the Anatidae, the biological family that includes the ducks and most duck-like waterfowl such as the geese and swans....

  • White-faced whistling duck
    White-faced Whistling Duck
    The White-faced Whistling Duck, Dendrocygna viduata, is a whistling duck which breeds in sub-Saharan Africa and much of South America.This species is gregarious, and at favoured sites, the flocks of a thousand or more birds arriving at dawn are an impressive sight...

     and Plumed whistling duck
    Plumed Whistling Duck
    The Plumed Whistling Duck , also called the Grass Whistle Duck , is a whistling duck which breeds in New Guinea and Australia. It is a predominantly brown-coloured duck with a long neck and characteristic plumes arising from its flanks...

  • Baikal teal
    Baikal Teal
    The Baikal Teal , also called the Bimaculate Duck or Squawk Duck, is a dabbling duck that breeds in eastern Russia and winters in East Asia.- Description :...

     and Northern pintail
    Northern Pintail
    The Pintail or Northern Pintail is a widely occurring duck which breeds in the northern areas of Europe, Asia and North America. It is strongly migratory and winters south of its breeding range to the equator...

  • Hooded merganser
    Hooded Merganser
    The Hooded Merganser is a small duck and is the only member of the genus Lophodytes.Hooded Mergansers have a crest at the back of the head which can be expanded or contracted. In adult males, this crest has a large white patch, the head is black and the sides of the duck are reddish-brown...

     and Smew
    Smew
    The Smew is a small duck, which is somewhat intermediate between the typical mergansers and the goldeneyes . It is the only member of the genus Mergellus; sometimes included in Mergus, this genus is distinct and might actually be a bit closer to the goldeneyes...

  • Eurasian wigeon and American wigeon
    American Wigeon
    The American Wigeon, also American Widgeon or Baldpate, is a species of wigeon in the dabbling duck genus Anas. If this is split up, all wigeons will go into their old genus Mareca again...

  • Mallard
    Mallard
    The Mallard , or Wild Duck , is a dabbling duck which breeds throughout the temperate and subtropical Americas, Europe, Asia, and North Africa, and has been introduced to New Zealand and Australia....

     and Grey Duck, a subspecies of the Pacific Black Duck
    Pacific Black Duck
    The Pacific Black Duck is a dabbling duck found in much of Indonesia, New Guinea, Australia, New Zealand, and many islands in the southwestern Pacific, reaching to the Caroline Islands in the north and French Polynesia in the east. It is usually called the Grey Duck in New Zealand...

    .

See also Mariana Mallard
Mariana Mallard
The Mariana Mallard or Oustalet's Duck is an extinct type of duck of the genus Anas that was endemic to the Mariana Islands...

.

Goose hybrids

Charles Darwin described hybrid geese in Origin of Species:
The hybrids from the common and Chinese geese (A. cygnoides), species which are so different that they are generally ranked in distinct genera, have often bred in this country with either pure parent, and in one single instance they have bred inter se.


Goose
Goose
The word goose is the English name for a group of waterfowl, belonging to the family Anatidae. This family also includes swans, most of which are larger than true geese, and ducks, which are smaller....

 hybrids include Canada Goose
Canada Goose
The Canada Goose is a wild goose belonging to the genus Branta, which is native to arctic and temperate regions of North America, having a black head and neck, white patches on the face, and a brownish-gray body....

 x Greylag Goose
Greylag Goose
The Greylag Goose , Anser anser, is a bird with a wide range in the Old World. It is the type species of the genus Anser....

, Canada Goose x Domesticated Geese, Emperor goose
Emperor Goose
The Emperor Goose is a species of goose. It breeds around the Bering Sea, mostly in Alaska, USA, but also in Kamchatka, Russia...

 x Canada goose, Red-breasted goose
Red-breasted Goose
The Red-breasted Goose is an endangered species of goose in the genus Branta. It is sometimes separated in Rufibrenta but appears close enough to the Brent Goose to make this unnecessary, despite its distinct appearance.- Description :All the species of the Branta genus are distinguished by their...

 x Canada goose, Canada goose x White-fronted goose
White-fronted Goose
The Greater White-fronted Goose is a species of goose. The Greater White-fronted Goose is more closely related to the smaller Lesser White-fronted Goose...

 and Barnacle goose
Barnacle Goose
The Barnacle Goose belongs to the genus Branta of black geese, which contains species with largely black plumage, distinguishing them from the grey Anser species...

x Canada goose.

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