Glomalin
Encyclopedia
Glomalin is a glycoprotein
produced abundantly on hyphae and spore
s of arbuscular
mycorrhizal fungi in soil
and in root
s.
As a glycoprotein, glomalin stores carbon
in both its protein
and its carbohydrate
(glucose
or sugar
) subunits. It permeates organic matter
, binding it to silt
, sand
, and clay
particles. Not only does glomalin contain 30 to 40 percent carbon but it also forms clumps of soil granules called aggregate
s. These add structure to soil
, and keep other stored soil carbon from escaping.
Glomalin was discovered in 1996 by Sara F. Wright, which she named after the Glomales order of fungi. Glomalin is causing a complete reexamination of what makes up soil organic matter. It is increasingly being included in studies of carbon storage and soil quality.
Glycoprotein
Glycoproteins are proteins that contain oligosaccharide chains covalently attached to polypeptide side-chains. The carbohydrate is attached to the protein in a cotranslational or posttranslational modification. This process is known as glycosylation. In proteins that have segments extending...
produced abundantly on hyphae and spore
Spore
In biology, a spore is a reproductive structure that is adapted for dispersal and surviving for extended periods of time in unfavorable conditions. Spores form part of the life cycles of many bacteria, plants, algae, fungi and some protozoa. According to scientist Dr...
s of arbuscular
Arbuscular mycorrhiza
An arbuscular mycorrhiza is a type of mycorrhiza in which the fungus penetrates the cortical cells of the roots of a vascular plant....
mycorrhizal fungi in soil
Soil
Soil is a natural body consisting of layers of mineral constituents of variable thicknesses, which differ from the parent materials in their morphological, physical, chemical, and mineralogical characteristics...
and in root
Root
In vascular plants, the root is the organ of a plant that typically lies below the surface of the soil. This is not always the case, however, since a root can also be aerial or aerating . Furthermore, a stem normally occurring below ground is not exceptional either...
s.
As a glycoprotein, glomalin stores carbon
Carbon
Carbon is the chemical element with symbol C and atomic number 6. As a member of group 14 on the periodic table, it is nonmetallic and tetravalent—making four electrons available to form covalent chemical bonds...
in both its protein
Protein
Proteins are biochemical compounds consisting of one or more polypeptides typically folded into a globular or fibrous form, facilitating a biological function. A polypeptide is a single linear polymer chain of amino acids bonded together by peptide bonds between the carboxyl and amino groups of...
and its carbohydrate
Carbohydrate
A carbohydrate is an organic compound with the empirical formula ; that is, consists only of carbon, hydrogen, and oxygen, with a hydrogen:oxygen atom ratio of 2:1 . However, there are exceptions to this. One common example would be deoxyribose, a component of DNA, which has the empirical...
(glucose
Glucose
Glucose is a simple sugar and an important carbohydrate in biology. Cells use it as the primary source of energy and a metabolic intermediate...
or sugar
Sugar
Sugar is a class of edible crystalline carbohydrates, mainly sucrose, lactose, and fructose, characterized by a sweet flavor.Sucrose in its refined form primarily comes from sugar cane and sugar beet...
) subunits. It permeates organic matter
Organic matter
Organic matter is matter that has come from a once-living organism; is capable of decay, or the product of decay; or is composed of organic compounds...
, binding it to silt
Silt
Silt is granular material of a size somewhere between sand and clay whose mineral origin is quartz and feldspar. Silt may occur as a soil or as suspended sediment in a surface water body...
, sand
Sand
Sand is a naturally occurring granular material composed of finely divided rock and mineral particles.The composition of sand is highly variable, depending on the local rock sources and conditions, but the most common constituent of sand in inland continental settings and non-tropical coastal...
, and clay
Clay
Clay is a general term including many combinations of one or more clay minerals with traces of metal oxides and organic matter. Geologic clay deposits are mostly composed of phyllosilicate minerals containing variable amounts of water trapped in the mineral structure.- Formation :Clay minerals...
particles. Not only does glomalin contain 30 to 40 percent carbon but it also forms clumps of soil granules called aggregate
Aggregate (composite)
Aggregate is the component of a composite material that resists compressive stress and provides bulk to the composite material. For efficient filling, aggregate should be much smaller than the finished item, but have a wide variety of sizes...
s. These add structure to soil
Soil structure
Soil structure is determined by how individual soil granules clump or bind together and aggregate, and therefore, the arrangement of soil pores between them...
, and keep other stored soil carbon from escaping.
Glomalin was discovered in 1996 by Sara F. Wright, which she named after the Glomales order of fungi. Glomalin is causing a complete reexamination of what makes up soil organic matter. It is increasingly being included in studies of carbon storage and soil quality.