Henonemus
Encyclopedia
Henonemus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of catfish
Catfish
Catfishes are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the heaviest and longest, the Mekong giant catfish from Southeast Asia and the second longest, the wels catfish of Eurasia, to detritivores...

es (order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

 Siluriformes) of the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Trichomycteridae
Trichomycteridae
Trichomycteridae is a family of catfishes commonly known as the pencil or parasitic catfishes. This family includes the infamous candiru fish, feared by some people for its alleged habit of entering into the urethra of humans....

. It includes five species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

, H. intermedius, H. macrops, H. punctatus, H. taxistigma, and H. triacanthopomus.

Though the monophyly
Monophyly
In common cladistic usage, a monophyletic group is a taxon which forms a clade, meaning that it contains all the descendants of the possibly hypothetical closest common ancestor of the members of the group. The term is synonymous with the uncommon term holophyly...

 of Henonemus has previously been in doubt, leading to this genus being placed in synonymy
Synonym (taxonomy)
In scientific nomenclature, a synonym is a scientific name that is or was used for a taxon of organisms that also goes by a different scientific name. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies...

 with Stegophilus
Stegophilus
Stegophilus is a genus of catfishes of the family Trichomycteridae. It includes three species, S. insidiosus, S. panzeri, and S. septentrionalis. S. insidiosus originates from the São Francisco River basin in Brazil, S. panzeri is from the lower Amazon River basin in Brazil, and S...

. However, monophyly has been argued for in 2006 and the genus was also recognized in a 2007 checklist of catfishes.

H. intermedius originates from the Araguaia River
Araguaia River
The Araguaia River or, in Portuguese, Rio Araguaia is one of the major rivers of Brazil, and the principal tributary of the Tocantins. It has a total length of approximately 2,627 km. Araguaia means "river of macaws" in the Tupi language....

 basin in Brazil
Brazil
Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

. H. macrops and H. punctatus both are known from the Amazon River
Amazon River
The Amazon of South America is the second longest river in the world and by far the largest by waterflow with an average discharge greater than the next seven largest rivers combined...

 basin, the former from Brazil and the latter from Brazil, Ecuador
Ecuador
Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

, and Peru
Peru
Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

. H. taxistigma inhabits the Rupununi River basin in Guyana
Guyana
Guyana , officially the Co-operative Republic of Guyana, previously the colony of British Guiana, is a sovereign state on the northern coast of South America that is culturally part of the Anglophone Caribbean. Guyana was a former colony of the Dutch and of the British...

. H. triacanthopomus has been collected in the Apure
Apure River
The Apure River is a river of southwestern Venezuela, formed by the confluence of the Sarare and Uribante near Guasdualito, in Venezuela, at , and flowing across the llanos into the Orinoco...

 and Arauca River
Arauca River
The Arauca River rises in the Andes Mountains of north-central Colombia and ends at the Orinoco in Venezuela. For part of its run it is the boundary between Colombia and Venezuela...

s and in Caño Macareo in the Orinoco
Orinoco
The Orinoco is one of the longest rivers in South America at . Its drainage basin, sometimes called the Orinoquia, covers , with 76.3% of it in Venezuela and the remainder in Colombia...

 Delta, but is probably distributed through the entire middle and lower reaches of the Orinoco River basin.

Fish of the genus Henonemus have an opercle
Ichthyology terms
Ichthyology uses several terms that are unique to the science.Contents: A B C D E F G H I J K L M N O P Q R S T U V W X Y Z- A :* Abdomen: belly* Abdominal: pertaining to the belly* Actinosts: a series of bones at the base of the pectoral rays....

 with two odontode
Odontode
Odontodes, or dermal teeth, are hard structures found on the external surfaces of animals or near internal openings. They comprise a soft pulp surrounded by dentine and covered by a mineralized substance such as enamel, a structure similar to that of teeth. They generally do not have the same...

s (though H. triacanthopomus differs from other Henonemus in that there are three or four odontodes in specimens greater than 80 millimetres or 3.1 in SL
Fish measurement
Fish measurement refers to the measuring of the length of individual fish and of various parts of their anatomy. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.-Overall length:...

) and the teeth of the most posterior row on the premaxilla
Premaxilla
The incisive bone is the portion of the maxilla adjacent to the incisors. It is a pair of small cranial bones at the very tip of the jaws of many animals, usually bearing teeth, but not always. They are connected to the maxilla and the nasals....

 and the dentary proximally
Anatomical terms of location
Standard anatomical terms of location are designations employed in science that deal with the anatomy of animals to avoid ambiguities that might otherwise arise. They are not language-specific, and thus require no translation...

 turned to the midline then abruptly bent laterally in the distal
Anatomical terms of location
Standard anatomical terms of location are designations employed in science that deal with the anatomy of animals to avoid ambiguities that might otherwise arise. They are not language-specific, and thus require no translation...

 half, and arranged in a compact band. They may grow about 4.1–9.4 centimetres (1.6–3.7 in) SL
Fish measurement
Fish measurement refers to the measuring of the length of individual fish and of various parts of their anatomy. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.-Overall length:...

.
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