Kileskus
Encyclopedia
Kileskus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of tyrannosauroid
Tyrannosauroidea
Tyrannosauroidea is a superfamily of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period...

 dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

 known from partial remains found in Middle Jurassic
Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from 176-161 million years ago. In European lithostratigraphy, rocks of this Middle Jurassic age are called the Dogger....

 (Bathonian
Bathonian
In the geologic timescale the Bathonian is an age or stage of the Middle Jurassic. It lasted from approximately 167.7 Ma to around 164.7 Ma...

 stage) rocks of Krasnoyarsk Krai
Krasnoyarsk Krai
Krasnoyarsk Krai is a federal subject of Russia . It is the second largest federal subject after the Sakha Republic, and Russia's largest krai, occupying an area of , which is 13% of the country's total territory. The administrative center of the krai is the city of Krasnoyarsk...

, Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

. Fossils recovered include the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 maxilla
Maxilla
The maxilla is a fusion of two bones along the palatal fissure that form the upper jaw. This is similar to the mandible , which is also a fusion of two halves at the mental symphysis. Sometimes The maxilla (plural: maxillae) is a fusion of two bones along the palatal fissure that form the upper...

, a premaxilla
Premaxilla
The incisive bone is the portion of the maxilla adjacent to the incisors. It is a pair of small cranial bones at the very tip of the jaws of many animals, usually bearing teeth, but not always. They are connected to the maxilla and the nasals....

, a surangular, and a few bones from the hand and foot. The skull bones are similar to those of Proceratosaurus
Proceratosaurus
Proceratosaurus is a genus of small-sized carnivorous theropod dinosaur from the Middle Jurassic of England. It was originally thought to be an ancestor of Ceratosaurus, due to the similar small crest on its snout...

. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 is K. aristotocus. Kileskus was named in 2010 by Averianov and colleagues. They performed a phylogenetic analysis
Cladistics
Cladistics is a method of classifying species of organisms into groups called clades, which consist of an ancestor organism and all its descendants . For example, birds, dinosaurs, crocodiles, and all descendants of their most recent common ancestor form a clade...

 and found it to be a basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 proceratosaurid
Proceratosauridae
Proceratosauridae is a family or clade of theropod dinosaurs, probably belonging to the tyrannosaur lineage. It was first named in 2010 by Oliver Rauhut and colleagues in their re-evaluation of the type genus, Proceratosaurus...

.
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