Lacuna (gastropod)
Encyclopedia
Lacuna is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of sea snail
Sea snail
Sea snail is a common name for those snails that normally live in saltwater, marine gastropod molluscs....

s, marine
Marine (ocean)
Marine is an umbrella term. As an adjective it is usually applicable to things relating to the sea or ocean, such as marine biology, marine ecology and marine geology...

 gastropod mollusks in the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Littorinidae
Littorinidae
Littorinidae is a taxonomic family of over 200 species of sea snails, marine gastropod molluscs in the clade Littorinimorpha, commonly known as periwinkles and found world-wide.-Names:...

, the winkles or periwinkles.

Description

The spiral animal body is elongated. The mouth has a proboscis
Proboscis
A proboscis is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In simpler terms, a proboscis is the straw-like mouth found in several varieties of species.-Etymology:...

 with two thick lips and a spiral, filiform tongue. There are two slender, contractile tentacles with the eyes on small pedicels near their base. The foot is oval and somewhat broader behind.

The thin shell has an ovate-conical or subglobose shape and is covered with a delicate horny periostracum
Periostracum
The periostracum is a thin organic coating or "skin" which is the outermost layer of the shell of many shelled animals, including mollusks and brachiopods. Among mollusks it is primarily seen in snails and clams, i.e. in bivalves and gastropods, but it is also found in cephalopods such as the...

. It has a short spire with convex whorls
Whorl (mollusc)
A whorl is a single, complete 360° revolution or turn in the spiral growth of a mollusc shell. A spiral configuration of the shell is found in of numerous gastropods, but it is also found in shelled cephalopods including Nautilus, Spirula and the large extinct subclass of cephalopods known as the...

 that are rapidly expanding. The last whorl is ventricose, i.e. inflated in the middle. The apex
Apex (mollusc)
Apex is an anatomical term for the tip of the mollusc shell of a gastropod, scaphopod, or cephalopod mollusc.-Gastropods:The word "apex" is most often used to mean the tip of the spire of the shell of a gastropod...

 is rather obtuse. The oval or roundish aperture
Aperture (mollusc)
The aperture is an opening in certain kinds of mollusc shells: it is the main opening of the shell, where part of the body of the animal emerges for locomotion, feeding, etc....

 is covered with a horny, spirally marked operculum
Operculum (gastropod)
The operculum, meaning little lid, is a corneous or calcareous anatomical structure which exists in many groups of sea snails and freshwater snails, and also in a few groups of land snails...

. The peristome
Peristome
The word peristome is derived from the Greek peri, meaning 'around' or 'about', and stoma, 'mouth'. It is a term used to describe various anatomical features that surround an opening to an organ or structure. The term is used in plants and invertebrate animals, such as in describing the shells of...

 is incomplete behind. The columella is flattened and forms with the peristome an elongated groove continued from the umbilicus. This groove is bounded internally by the decurved margin of the pillar.

Species

Species within the genus Lacuna include:
  • Lacuna abyssicola
    Lacuna abyssicola
    Lacuna abyssicola is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.This name is a nomen nudum...

    Melvill & Standen, 1912 : nomen nudum
  • Lacuna carinifera
    Lacuna carinifera
    Lacuna carinifera is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    A. Adams, 1851
  • Lacuna cleicecabrale Barros, 1994
  • Lacuna crassior
    Lacuna crassior
    Lacuna crassior is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.-Distribution:Lacuna crassior is a circumboreal and subarctic species that has Britain as its southern limit...

    (Montagu, 1803)
  • Lacuna decotata
    Lacuna decotata
    Lacuna decorata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.-References:...

    A. Adams, 1861
  • Lacuna latifasciata
    Lacuna latifasciata
    Lacuna latifasciata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    A. Adams, 1863
  • Lacuna lepidula
    Lacuna lepidula
    Lacuna lepidula is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    A. Adams, 1863
  • Lacuna lukinii
    Lacuna lukinii
    Lacuna lukinii is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Golikov & Gulbin, 1985)
  • Lacuna marmorata
    Lacuna marmorata
    Lacuna marmorata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Dall, 1919
  • Lacuna minor
    Lacuna minor
    Lacuna minor is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Dall, 1919)
  • Lacuna orientalis
    Lacuna orientalis
    Lacuna orientalis is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Golikov & Gulbin, 1985)
  • Lacuna pallidula
    Lacuna pallidula
    Lacuna pallidula, common name the pale lacuna, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (da Costa, 1778)
  • Lacuna parva
    Lacuna parva
    Lacuna parva, common name the tiny lacuna, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (da Costa, 1778)
  • Lacuna pumilio E. A. Smith, 1890
  • Lacuna porrecta
    Lacuna porrecta
    Lacuna porrecta is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Carpenter, 1864
  • Lacuna reflexa
    Lacuna reflexa
    Lacuna reflexa is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Dall, 1884)
  • Lacuna setonaikaiensis
    Lacuna setonaikaiensis
    Lacuna setonaikaiensis is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Habe, 1958)
  • Lacuna smithii
    Lacuna smithii
    Lacuna smithii is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Pilsbry, 1895
  • Lacuna succinea
    Lacuna succinea
    Lacuna succinea is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Berry, 1953
  • Lacuna turneri
    Lacuna turneri
    Lacuna turneri is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Dall, 1886)
  • Lacuna turrita
    Lacuna turrita
    Lacuna turrita is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    A. Adams, 1861
  • Lacuna uchidai
    Lacuna uchidai
    Lacuna uchidai is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Habe, 1953)
  • Lacuna unifasciata
    Lacuna unifasciata
    Lacuna unifasciata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Carpenter, 1856
  • Lacuna variegata
    Lacuna variegata
    Lacuna variegata is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    Carpenter, 1864
  • Lacuna vincta
    Lacuna vincta
    Lacuna vincta, common name the northern lacuna, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles....

    (Montagu, 1803)

Species brought into synonymy :
  • Lacuna cossmanni Locard, 1897 : synonym of Benthobia tryonii
    Benthobia tryonii
    Benthobia tryonii is a species of sea snail, a marine gastropod mollusk in the family Pseudolividae....

    Dall, 1889
  • Lacuna macmurdensis Hedley, 1911 : synonym of Trilirata macmurdensis (Hedley, 1911)
  • Lacuna mediterranea Monterosato, 1869 : synonym of Ersilia mediterranea (Monterosato, 1869)
  • Lacuna neritoidea Gould, 1840 : synonym of Lacuna pallidula (da Costa, 1778)
  • Lacuna notorcadensis Melvill & Standen, 1907 : synonym of Laevilitorina wandelensis
    Laevilitorina wandelensis
    Laevilitorina wandelensis is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles.-References:...

    (Lamy, 1905)
  • Lacuna patula Thorpe, 1844 : synonym of Lacuna pallidula (da Costa, 1778)
  • Lacuna puteolus (Turton, 1819) : synonym of Lacuna parva (da Costa, 1778)
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