Macaronesia
Encyclopedia
Macaronesia is a modern collective name for several groups of island
s in the North Atlantic Ocean near Europe
and North Africa
belonging to three countries: Portugal
, Spain
, and Cape Verde
. The name (which is often misspelled 'Macronesia') comes from the Greek
for "islands of the fortunate
" μακάρων νῆσοι makárōn nêsoi, a term used by Ancient Greek
geographer
s for islands to the west of the Straits of Gibraltar.
Macaronesia consists of five archipelago
s:
The islands of Macaronesia are volcanic
in origin, and are thought to be the product of several geologic hotspot
s.
The climate
of the Macaronesian islands ranges from subtropical
to tropical
. The Portuguese
archipelagos of the Azores
and Madeira
have a generally cooler climate and higher rainfall than the Canaries
and Cape Verde
. The laurisilva
forests of Macaronesia are a type of mountain cloud forest
with relict
s species of a vegetation
type which originally covered much of the Mediterranean Basin
when the climate
of the region was more humid
and evolved to adapt in islanders conditions.
The islands have a unique biogeography
, and are home to several distinct plant and animal communities. None of the Macaronesian islands were part of a continent, so the native plants and animals reached the islands via long-distance dispersal. Laurel-leaved forests
, called laurisilva
, once covered most of the Azores, Madeira, and parts of the Canaries between 400–1200 m altitude (the eastern Canaries and Cape Verde being too dry). These forests resemble the ancient forests that covered the Mediterranean basin and northwestern Africa before cooling and drying of the ice age
s. Trees of the genera Apollonias
(Lauraceae
), Clethra
(Clethra
ceae), Dracaena
(Ruscaceae
), Ocotea
(Lauraceae
), Persea
(Lauraceae
), and Picconia
(Oleaceae
), which are found in the Macaronesian laurel forests, are also known from fossil evidence to have lived around the Mediterranean before the ice ages.
Felling of the forests for timber and firewood, clearing vegetation for grazing and agriculture, and the introduction of exotic plants and animals by humans has displaced much of the native vegetation. The laurisilva has been reduced to small pockets. As a result, many of the endemic biota
of the islands are seriously endangered or extinct.
The jumping spider
genus Macaroeris
is named after Macaronesia.
Island
An island or isle is any piece of sub-continental land that is surrounded by water. Very small islands such as emergent land features on atolls can be called islets, cays or keys. An island in a river or lake may be called an eyot , or holm...
s in the North Atlantic Ocean near Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...
and North Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...
belonging to three countries: Portugal
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
, Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...
, and Cape Verde
Cape Verde
The Republic of Cape Verde is an island country, spanning an archipelago of 10 islands located in the central Atlantic Ocean, 570 kilometres off the coast of Western Africa...
. The name (which is often misspelled 'Macronesia') comes from the Greek
Greek language
Greek is an independent branch of the Indo-European family of languages. Native to the southern Balkans, it has the longest documented history of any Indo-European language, spanning 34 centuries of written records. Its writing system has been the Greek alphabet for the majority of its history;...
for "islands of the fortunate
Fortunate Isles
In the Fortunate Isles, also called the Isles of the Blessed , heroes and other favored mortals in Greek mythology and Celtic mythology were received by the gods into a winterless blissful paradise...
" μακάρων νῆσοι makárōn nêsoi, a term used by Ancient Greek
Ancient Greek
Ancient Greek is the stage of the Greek language in the periods spanning the times c. 9th–6th centuries BC, , c. 5th–4th centuries BC , and the c. 3rd century BC – 6th century AD of ancient Greece and the ancient world; being predated in the 2nd millennium BC by Mycenaean Greek...
geographer
Geographer
A geographer is a scholar whose area of study is geography, the study of Earth's natural environment and human society.Although geographers are historically known as people who make maps, map making is actually the field of study of cartography, a subset of geography...
s for islands to the west of the Straits of Gibraltar.
Macaronesia consists of five archipelago
Archipelago
An archipelago , sometimes called an island group, is a chain or cluster of islands. The word archipelago is derived from the Greek ἄρχι- – arkhi- and πέλαγος – pélagos through the Italian arcipelago...
s:
- AzoresAzoresThe Archipelago of the Azores is composed of nine volcanic islands situated in the middle of the North Atlantic Ocean, and is located about west from Lisbon and about east from the east coast of North America. The islands, and their economic exclusion zone, form the Autonomous Region of the...
(PortugalPortugalPortugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
) - MadeiraMadeiraMadeira is a Portuguese archipelago that lies between and , just under 400 km north of Tenerife, Canary Islands, in the north Atlantic Ocean and an outermost region of the European Union...
(PortugalPortugalPortugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
), including Porto Santo IslandPorto Santo IslandPorto Santo Island is a Portuguese island northeast of Madeira Island in the North Atlantic Ocean; it is the northernmost and easternmost island of the archipelago of Madeira, located in the Atlantic Ocean west of Europe and Africa.- History :...
, the Desertas IslandsDesertas IslandsThe Desertas Islands are a small Portuguese archipelago, located about 25 km to the southeast of Ponta de São Lourenço, the eastern tip of the island of Madeira, the whole chain located roughly between Madeira and the Canary Islands.-Geography:...
and the - Savage IslandsSavage IslandsThe Savage Islands, also referred to as the Salvage Islands or the Selvagens Islands, of Sé. They are designated a Nature Reserve, comprising two areas: one on Selvagem Grande Island and the second on Selvagem Pequena Island.-Geography:...
(PortugalPortugalPortugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
), administratively considered part of the Madeira archipelago - Canary IslandsCanary IslandsThe Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...
(SpainSpainSpain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...
) - Cape VerdeCape VerdeThe Republic of Cape Verde is an island country, spanning an archipelago of 10 islands located in the central Atlantic Ocean, 570 kilometres off the coast of Western Africa...
(Cape VerdeCape VerdeThe Republic of Cape Verde is an island country, spanning an archipelago of 10 islands located in the central Atlantic Ocean, 570 kilometres off the coast of Western Africa...
)
The islands of Macaronesia are volcanic
Volcano
2. Bedrock3. Conduit 4. Base5. Sill6. Dike7. Layers of ash emitted by the volcano8. Flank| 9. Layers of lava emitted by the volcano10. Throat11. Parasitic cone12. Lava flow13. Vent14. Crater15...
in origin, and are thought to be the product of several geologic hotspot
Hotspot (geology)
The places known as hotspots or hot spots in geology are volcanic regions thought to be fed by underlying mantle that is anomalously hot compared with the mantle elsewhere. They may be on, near to, or far from tectonic plate boundaries. There are two hypotheses to explain them...
s.
The climate
Climate
Climate encompasses the statistics of temperature, humidity, atmospheric pressure, wind, rainfall, atmospheric particle count and other meteorological elemental measurements in a given region over long periods...
of the Macaronesian islands ranges from subtropical
Subtropics
The subtropics are the geographical and climatical zone of the Earth immediately north and south of the tropical zone, which is bounded by the Tropic of Cancer and the Tropic of Capricorn, at latitudes 23.5°N and 23.5°S...
to tropical
Tropics
The tropics is a region of the Earth surrounding the Equator. It is limited in latitude by the Tropic of Cancer in the northern hemisphere at approximately N and the Tropic of Capricorn in the southern hemisphere at S; these latitudes correspond to the axial tilt of the Earth...
. The Portuguese
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
archipelagos of the Azores
Azores
The Archipelago of the Azores is composed of nine volcanic islands situated in the middle of the North Atlantic Ocean, and is located about west from Lisbon and about east from the east coast of North America. The islands, and their economic exclusion zone, form the Autonomous Region of the...
and Madeira
Madeira
Madeira is a Portuguese archipelago that lies between and , just under 400 km north of Tenerife, Canary Islands, in the north Atlantic Ocean and an outermost region of the European Union...
have a generally cooler climate and higher rainfall than the Canaries
Canary Islands
The Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...
and Cape Verde
Cape Verde
The Republic of Cape Verde is an island country, spanning an archipelago of 10 islands located in the central Atlantic Ocean, 570 kilometres off the coast of Western Africa...
. The laurisilva
Laurisilva
Laurisilva or laurissilva is a subtropical forest, found in areas with high humidity and relatively stable and mild temperatures. They are characterised by evergreen, glossy-leaved tree species that look alike with leaves of lauroide type...
forests of Macaronesia are a type of mountain cloud forest
Cloud forest
A cloud forest, also called a fog forest, is a generally tropical or subtropical evergreen montane moist forest characterized by a persistent, frequent or seasonal low-level cloud cover, usually at the canopy level. Cloud forests often exhibit an abundance of mosses covering the ground and...
with relict
Relict
A relict is a surviving remnant of a natural phenomenon.* In biology a relict is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas....
s species of a vegetation
Vegetation
Vegetation is a general term for the plant life of a region; it refers to the ground cover provided by plants. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic characteristics. It is broader...
type which originally covered much of the Mediterranean Basin
Mediterranean Basin
In biogeography, the Mediterranean Basin refers to the lands around the Mediterranean Sea that have a Mediterranean climate, with mild, rainy winters and hot, dry summers, which supports characteristic Mediterranean forests, woodlands, and scrub vegetation...
when the climate
Climate
Climate encompasses the statistics of temperature, humidity, atmospheric pressure, wind, rainfall, atmospheric particle count and other meteorological elemental measurements in a given region over long periods...
of the region was more humid
Humidity
Humidity is a term for the amount of water vapor in the air, and can refer to any one of several measurements of humidity. Formally, humid air is not "moist air" but a mixture of water vapor and other constituents of air, and humidity is defined in terms of the water content of this mixture,...
and evolved to adapt in islanders conditions.
The islands have a unique biogeography
Biogeography
Biogeography is the study of the distribution of species , organisms, and ecosystems in space and through geological time. Organisms and biological communities vary in a highly regular fashion along geographic gradients of latitude, elevation, isolation and habitat area...
, and are home to several distinct plant and animal communities. None of the Macaronesian islands were part of a continent, so the native plants and animals reached the islands via long-distance dispersal. Laurel-leaved forests
Laurel forest
Laurel forest is a subtropical or mild temperate forest, found in areas with high humidity and relatively stable and mild temperatures. They are characterized by tree species with evergreen, glossy, enlongated leaves, known as laurophyll or lauroide...
, called laurisilva
Laurisilva
Laurisilva or laurissilva is a subtropical forest, found in areas with high humidity and relatively stable and mild temperatures. They are characterised by evergreen, glossy-leaved tree species that look alike with leaves of lauroide type...
, once covered most of the Azores, Madeira, and parts of the Canaries between 400–1200 m altitude (the eastern Canaries and Cape Verde being too dry). These forests resemble the ancient forests that covered the Mediterranean basin and northwestern Africa before cooling and drying of the ice age
Ice age
An ice age or, more precisely, glacial age, is a generic geological period of long-term reduction in the temperature of the Earth's surface and atmosphere, resulting in the presence or expansion of continental ice sheets, polar ice sheets and alpine glaciers...
s. Trees of the genera Apollonias
Apollonias
Apollonias is a genus of flowering plants belonging to the laurel family, Lauraceae. The genus includes from one to ten species of evergreen trees and shrubs from laurel forest habitat mainly in Macaronesian islands.-Description:...
(Lauraceae
Lauraceae
The Lauraceae or Laurel family comprises a group of flowering plants included in the order Laurales. The family contains about 55 genera and over 3500, perhaps as many as 4000, species world-wide, mostly from warm or tropical regions, especially Southeast Asia and South America...
), Clethra
Clethra
Clethra is a genus of between 30-70 species of flowering shrubs or small trees. It is one of two genera in the family Clethraceae...
(Clethra
Clethra
Clethra is a genus of between 30-70 species of flowering shrubs or small trees. It is one of two genera in the family Clethraceae...
ceae), Dracaena
Dracaena (plant)
Dracaena is a genus of about 40 species of trees and succulent shrubs. In the APG III classification system, it is placed in the family Asparagaceae, subfamily Nolinoideae . It has also formerly been separated into the family Dracaenaceae or placed in the Agavaceae...
(Ruscaceae
Ruscaceae
Nolinoideae is a monocot subfamily of the family Asparagaceae in the APG III system of 2009. It was previously treated as a separate family, Ruscaceae s.l...
), Ocotea
Ocotea
Ocotea is a genus of flowering plants belonging to the family Lauraceae. The genus includes over 200 species of evergreen trees and shrubs, distributed mostly in tropical and subtropical regions of Central and South America, the West Indies, also with a few species in Africa and Madagascar, and...
(Lauraceae
Lauraceae
The Lauraceae or Laurel family comprises a group of flowering plants included in the order Laurales. The family contains about 55 genera and over 3500, perhaps as many as 4000, species world-wide, mostly from warm or tropical regions, especially Southeast Asia and South America...
), Persea
Persea
Persea is a genus of about 150 species of evergreen trees belonging to the laurel family, Lauraceae. The best-known member of the genus is the avocado, P. americana, widely cultivated in subtropical regions for its large, edible fruit.-Overview:...
(Lauraceae
Lauraceae
The Lauraceae or Laurel family comprises a group of flowering plants included in the order Laurales. The family contains about 55 genera and over 3500, perhaps as many as 4000, species world-wide, mostly from warm or tropical regions, especially Southeast Asia and South America...
), and Picconia
Picconia
Picconia is a genus of flowering plants in the family Oleaceae.-Species:*Picconia azorica - endemic to the archipelago of the Azores*Picconia excelsa - tree up to 15m; Madeira, Canaries...
(Oleaceae
Oleaceae
Oleaceae are a family containing 24 extant genera and around 600 species of mesophytic shrubs, trees and occasionally vines. As shrubs, members of this family may be twine climbers, or scramblers.-Leaves:...
), which are found in the Macaronesian laurel forests, are also known from fossil evidence to have lived around the Mediterranean before the ice ages.
Felling of the forests for timber and firewood, clearing vegetation for grazing and agriculture, and the introduction of exotic plants and animals by humans has displaced much of the native vegetation. The laurisilva has been reduced to small pockets. As a result, many of the endemic biota
Biota (ecology)
Biota are the total collection of organisms of a geographic region or a time period, from local geographic scales and instantaneous temporal scales all the way up to whole-planet and whole-timescale spatiotemporal scales. The biota of the Earth lives in the biosphere.-See...
of the islands are seriously endangered or extinct.
The jumping spider
Jumping spider
The jumping spider family contains more than 500 described genera and about 5,000 described species, making it the largest family of spiders with about 13% of all species. Jumping spiders have some of the best vision among invertebrates and use it in courtship, hunting and navigation...
genus Macaroeris
Macaroeris
Macaroeris is a spider genus of the Salticidae family .-Name:The genus name is combined from Macaronesia, where most species of the genus occur, and the salticid spider genus Eris.-Species:...
is named after Macaronesia.