Common names: blunt-nosed viper, Lebetine viper, Levant viper, more.

Macrovipera lebetina is a venomous

Venomous snake

"Poisonous snake" redirects here. For true poisonous snakes, see Rhabdophis.Venomous snakes are snakes which have venom glands and specialized teeth for the injection of venom...

 viper

Viperinae

The Viperinae, or viperines, are a subfamily of venomous vipers found in Europe, Asia and Africa. They are distinguished by their lack of the heat-sensing pit organs that characterize their sister group, the Crotalinae. Currently, 12 genera and 66 species are recognized...

 species

Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 found in North Africa

North Africa

North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

, much of the Middle East

Middle East

The Middle East is a region that encompasses Western Asia and Northern Africa. It is often used as a synonym for Near East, in opposition to Far East...

, and as far east as Kashmir

Kashmir

Kashmir is the northwestern region of the Indian subcontinent. Until the mid-19th century, the term Kashmir geographically denoted only the valley between the Great Himalayas and the Pir Panjal mountain range...

. Five subspecies are currently recognized, including the nominate race described here.

Description

This is a large snake, with females reaching 150 cm (59.1 in) in length and males a little less. Sizes vary between different populations, with M. l. lebetina being somewhat smaller.

It will be found under short trees with thick shade close to the roots if the outside temperatures reaches 45 Celsius plus.

The head is broad, triangular and distinct from the neck. The snout is rounded and blunt when viewed from above, which is why it is also called the blunt-nosed viper. The nasal and nasorostral scales are almost completely fused into a single plate, although some variation occurs.

The dorsal scales

Dorsal scales

In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales.When counting dorsal scales, numbers are often given for three points along the body, for example 19:21:17...

 are strongly keeled, except for those bordering the ventrals. M. l. lebetina usually has 146-163 ventral scales. The anal scale

Anal scale

In snakes, the anal scale is the scale just in front of and covering the cloacal opening. This scale can be either single or paired . When paired, the division is oblique. It is preceded by the ventral scales and followed by the subcaudal scales....

 is single.

The color pattern is less varied than one might expect from a species that is so widely distributed. The head is normally uniformly colored, although it can occasionally be marked with a dark V-shape. Dorsally, the ground color for the body can be gray, brown, beige, pinkish, olive or khaki. The pattern, if present, is darker, can be gray, bluish, rust or brown in color, and may consist of a middorsal row or double row of large spots. When two rows are present, the spots may alternate or oppose, which can produce anything from a saddled to a continuous zigzag pattern. The spots are usually brown, dark gray or black, but are sometimes red, brick, yellow or olive in color. Males are usually 3 and a half feet (1 m) long, while females are 5 feet (1.5 m) long.

Common names

There are many names for this viper. Here are some common names: Blunt-nosed viper, Lebetine viper, Levant viper, Levantine viper, Levantine adder, Kufi or Kufi viper (from Arabic), Gjurza (from Russian

Russian language

Russian is a Slavic language used primarily in Russia, Belarus, Uzbekistan, Kazakhstan, Tajikistan and Kyrgyzstan. It is an unofficial but widely spoken language in Ukraine, Moldova, Latvia, Turkmenistan and Estonia and, to a lesser extent, the other countries that were once constituent republics...

), coffin snake, Levante viper, mountain viper, Gunas (from Kashmiri

Kashmiri language

Kashmiri is a language from the Dardic sub-group and it is spoken primarily in the Kashmir Valley, in Jammu and Kashmir. There are approximately 5,554,496 speakers in Jammu and Kashmir, according to the Census of 2001. Most of the 105,000 speakers or so in Pakistan are émigrés from the Kashmir...

), Fina or Kontonoura (the second comes from the translation of Cypriot local dialect for "short-tailed").

Geographic range

The Blunt-nosed viper can be found in these places, Algeria

Algeria

Algeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab...

, Tunisia

Tunisia

Tunisia , officially the Tunisian RepublicThe long name of Tunisia in other languages used in the country is: , is the northernmost country in Africa. It is a Maghreb country and is bordered by Algeria to the west, Libya to the southeast, and the Mediterranean Sea to the north and east. Its area...

, Cyprus

Cyprus

Cyprus , officially the Republic of Cyprus , is a Eurasian island country, member of the European Union, in the Eastern Mediterranean, east of Greece, south of Turkey, west of Syria and north of Egypt. It is the third largest island in the Mediterranean Sea.The earliest known human activity on the...

, Turkey

Turkey

Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

, Syria

Syria

Syria , officially the Syrian Arab Republic , is a country in Western Asia, bordering Lebanon and the Mediterranean Sea to the West, Turkey to the north, Iraq to the east, Jordan to the south, and Israel to the southwest....

, Jordan

Jordan

Jordan , officially the Hashemite Kingdom of Jordan , Al-Mamlaka al-Urduniyya al-Hashemiyya) is a kingdom on the East Bank of the River Jordan. The country borders Saudi Arabia to the east and south-east, Iraq to the north-east, Syria to the north and the West Bank and Israel to the west, sharing...

, Israel

Israel

The State of Israel is a parliamentary republic located in the Middle East, along the eastern shore of the Mediterranean Sea...

, Lebanon

Lebanon

Lebanon , officially the Republic of LebanonRepublic of Lebanon is the most common term used by Lebanese government agencies. The term Lebanese Republic, a literal translation of the official Arabic and French names that is not used in today's world. Arabic is the most common language spoken among...

, Iraq

Iraq

Iraq ; officially the Republic of Iraq is a country in Western Asia spanning most of the northwestern end of the Zagros mountain range, the eastern part of the Syrian Desert and the northern part of the Arabian Desert....

, Iran

Iran

Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia...

, Russian Caucasia

Caucasus

The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

, Armenia

Armenia

Armenia , officially the Republic of Armenia , is a landlocked mountainous country in the Caucasus region of Eurasia...

, Georgia

Georgia (country)

Georgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...

, Azerbaijan

Azerbaijan

Azerbaijan , officially the Republic of Azerbaijan is the largest country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to...

, Turkmenistan

Turkmenistan

Turkmenistan , formerly also known as Turkmenia is one of the Turkic states in Central Asia. Until 1991, it was a constituent republic of the Soviet Union, the Turkmen Soviet Socialist Republic . Turkmenistan is one of the six independent Turkic states...

, Uzbekistan

Uzbekistan

Uzbekistan , officially the Republic of Uzbekistan is a doubly landlocked country in Central Asia and one of the six independent Turkic states. It shares borders with Kazakhstan to the west and to the north, Kyrgyzstan and Tajikistan to the east, and Afghanistan and Turkmenistan to the south....

, Kazakhistan, Tadzikhistan, Afghanistan

Afghanistan

Afghanistan , officially the Islamic Republic of Afghanistan, is a landlocked country located in the centre of Asia, forming South Asia, Central Asia and the Middle East. With a population of about 29 million, it has an area of , making it the 42nd most populous and 41st largest nation in the world...

, Pakistan

Pakistan

Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

 and Kashmir

Kashmir

Kashmir is the northwestern region of the Indian subcontinent. Until the mid-19th century, the term Kashmir geographically denoted only the valley between the Great Himalayas and the Pir Panjal mountain range...

, India

India

India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

.

At one point there was also a report of this species from Yemen

Yemen

The Republic of Yemen , commonly known as Yemen , is a country located in the Middle East, occupying the southwestern to southern end of the Arabian Peninsula. It is bordered by Saudi Arabia to the north, the Red Sea to the west, and Oman to the east....

 by Scortecci (1929).

The type locality originally given was "Oriente". Mertens and Müller (1928) suggested restricting the range to "Cypern" (Cyprus).

Conservation status

Lebetine vipers are endangered. The species is listed as strictly protected (Appendix II) under the Berne Convention

Convention on the Conservation of European Wildlife and Natural Habitats

The Bern Convention on the Conservation of European Wildlife and Natural Habitats 1979, also known as the Bern Convention , came into force on June 1, 1982....

.

Subspecies

Subspecies	Taxon author	Common name	Geographic range
M. l. cernovi Macrovipera lebetina cernovi Macrovipera lebetina cernovi is a venomous viper subspecies found in Iran, Turkmenistan and Afghanistan.-Geographic range:Northeast Iran, southern Turkmenistan parts of northern Afghanistan and Pakistan....	(Chikin & Szczerbak, 1992)		Northeast Iran Iran Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia... , southern Turkmenistan Turkmenistan Turkmenistan , formerly also known as Turkmenia is one of the Turkic states in Central Asia. Until 1991, it was a constituent republic of the Soviet Union, the Turkmen Soviet Socialist Republic . Turkmenistan is one of the six independent Turkic states... , parts of northern Afghanistan Afghanistan Afghanistan , officially the Islamic Republic of Afghanistan, is a landlocked country located in the centre of Asia, forming South Asia, Central Asia and the Middle East. With a population of about 29 million, it has an area of , making it the 42nd most populous and 41st largest nation in the world...  and Pakistan Pakistan Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...  (Kashmir Kashmir Kashmir is the northwestern region of the Indian subcontinent. Until the mid-19th century, the term Kashmir geographically denoted only the valley between the Great Himalayas and the Pir Panjal mountain range... ).
M. l. lebetina	(Linnaeus Carolus Linnaeus Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology... , 1758)	Cyprian blunt-nosed viper	Cyprus Cyprus Cyprus , officially the Republic of Cyprus , is a Eurasian island country, member of the European Union, in the Eastern Mediterranean, east of Greece, south of Turkey, west of Syria and north of Egypt. It is the third largest island in the Mediterranean Sea.The earliest known human activity on the...
M. l. obtusa Macrovipera lebetina obtusa Macrovipera lebetina obtusa is a venomous viper subspecies found between central Turkey and northern Pakistan .-Description:Distinguished by its higher scale counts—usually 170-175 ventrals and 25 midbody dorsal scales -- and relatively dark color pattern.-Geographic range:From central Turkey...	(Dwigubsky, 1832)	Levant blunt-nosed viper	Turkey Turkey Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe... , Syria Syria Syria , officially the Syrian Arab Republic , is a country in Western Asia, bordering Lebanon and the Mediterranean Sea to the West, Turkey to the north, Iraq to the east, Jordan to the south, and Israel to the southwest.... , Lebanon Lebanon Lebanon , officially the Republic of LebanonRepublic of Lebanon is the most common term used by Lebanese government agencies. The term Lebanese Republic, a literal translation of the official Arabic and French names that is not used in today's world. Arabic is the most common language spoken among... , Iraq Iraq Iraq ; officially the Republic of Iraq is a country in Western Asia spanning most of the northwestern end of the Zagros mountain range, the eastern part of the Syrian Desert and the northern part of the Arabian Desert.... , north Jordan Jordan Jordan , officially the Hashemite Kingdom of Jordan , Al-Mamlaka al-Urduniyya al-Hashemiyya) is a kingdom on the East Bank of the River Jordan. The country borders Saudi Arabia to the east and south-east, Iraq to the north-east, Syria to the north and the West Bank and Israel to the west, sharing... , Caucasus Caucasus The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...  (incl. Armenia Armenia Armenia , officially the Republic of Armenia , is a landlocked mountainous country in the Caucasus region of Eurasia... ), Azerbaijan Azerbaijan Azerbaijan , officially the Republic of Azerbaijan is the largest country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to... , Dagestan Dagestan The Republic of Dagestan is a federal subject of Russia, located in the North Caucasus region. Its capital and the largest city is Makhachkala, located at the center of Dagestan on the Caspian Sea... , Iran Iran Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia... , southern Afghanistan, Pakistan Pakistan Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...  (Kashmir Kashmir Kashmir is the northwestern region of the Indian subcontinent. Until the mid-19th century, the term Kashmir geographically denoted only the valley between the Great Himalayas and the Pir Panjal mountain range... ), and north India India India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...
M. l. transmediterranea Macrovipera lebetina transmediterranea Macrovipera lebetina transmediterranea is a venomous viper subspecies found only in Algeria and Tunis.-Description:This subspecies is not known to exceed 1 m in length. It is further distinguished by having only 25 midbody dorsal scale rows, a lower ventral scale count of 150-164 scales and more...	(Nilson & Andrén, 1988)		Algeria Algeria Algeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab... , Tunisia Tunisia Tunisia , officially the Tunisian RepublicThe long name of Tunisia in other languages used in the country is: , is the northernmost country in Africa. It is a Maghreb country and is bordered by Algeria to the west, Libya to the southeast, and the Mediterranean Sea to the north and east. Its area...
M. l. turanica Macrovipera lebetina turanica Macrovipera lebetina turanica is a venomous viper subspecies found in Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan, Afghanistan and Pakistan.-Description: The color pattern consists of a dark ground color with a lighter, orange dorsal zigzag pattern...	(Cernov, 1940)	Turan blunt-nosed viper	Eastern Turkmenistan Turkmenistan Turkmenistan , formerly also known as Turkmenia is one of the Turkic states in Central Asia. Until 1991, it was a constituent republic of the Soviet Union, the Turkmen Soviet Socialist Republic . Turkmenistan is one of the six independent Turkic states... , Uzbekistan Uzbekistan Uzbekistan , officially the Republic of Uzbekistan is a doubly landlocked country in Central Asia and one of the six independent Turkic states. It shares borders with Kazakhstan to the west and to the north, Kyrgyzstan and Tajikistan to the east, and Afghanistan and Turkmenistan to the south.... , Tajikistan Tajikistan Tajikistan , officially the Republic of Tajikistan , is a mountainous landlocked country in Central Asia. Afghanistan borders it to the south, Uzbekistan to the west, Kyrgyzstan to the north, and China to the east.... , southwestern Kazakhstan Kazakhstan Kazakhstan , officially the Republic of Kazakhstan, is a transcontinental country in Central Asia and Eastern Europe. Ranked as the ninth largest country in the world, it is also the world's largest landlocked country; its territory of is greater than Western Europe... , parts of northern Afghanistan Afghanistan Afghanistan , officially the Islamic Republic of Afghanistan, is a landlocked country located in the centre of Asia, forming South Asia, Central Asia and the Middle East. With a population of about 29 million, it has an area of , making it the 42nd most populous and 41st largest nation in the world...  and western Pakistan Pakistan Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

Taxonomy

This species is currently subject to review. It is likely that certain subspecies will soon be elevated to valid species status. The nominate subspecies was restricted to Cyprus in 1928 by Mertens and Müller and so does not actually occur in the Levant

Levant

The Levant or ) is the geographic region and culture zone of the "eastern Mediterranean littoral between Anatolia and Egypt" . The Levant includes most of modern Lebanon, Syria, Jordan, Israel, the Palestinian territories, and sometimes parts of Turkey and Iraq, and corresponds roughly to the...

 region.

The populations found in southern Afghanistan

Afghanistan

Afghanistan , officially the Islamic Republic of Afghanistan, is a landlocked country located in the centre of Asia, forming South Asia, Central Asia and the Middle East. With a population of about 29 million, it has an area of , making it the 42nd most populous and 41st largest nation in the world...

, Pakistan

Pakistan

Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

 (Kashmir

Kashmir

Kashmir is the northwestern region of the Indian subcontinent. Until the mid-19th century, the term Kashmir geographically denoted only the valley between the Great Himalayas and the Pir Panjal mountain range...

) and northern India

India

India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

 are sometimes referred to as a separate subspecies: M. l. peilei. These normally have semidivided supraoculars.

Vipera Euphratica was originally used to refer to the populations that occur in the Euphrates

Euphrates

The Euphrates is the longest and one of the most historically important rivers of Western Asia. Together with the Tigris, it is one of the two defining rivers of Mesopotamia...

river basin of Turkey, Syria and Iraq. It was synonymized with M. l. obtusa in several publications, including Joger (1984). However, Golay et al. (1993) include it in the synonymy of M. l. lebetina.

External links

• Macrovipera lebetina at Amphibians and Reptiles of Europe. Accessed 10 August 2006.
• Macrovipera at Herpbreeder. Accessed 10 August 2006.
• Macrovipera lebetina at Wildlife of Pakistan. Accessed 15 March 2007.
• Macrovipera lebetina at Checklist of Armenia's Amphibians and Reptiles, Tadevosyan's Herpetological Resources. Accessed 30 March 2007.