Mahajangasuchidae is an extinct family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of neosuchia

Neosuchia

Neosuchia is an unranked clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to Crocodylus niloticus than to Notosuchus terrestris...

n crocodyliforms. It currently contains two genera, Mahajangasuchus

Mahajangasuchus

Mahajangasuchus is an extinct genus of crocodyliform which had blunt, conical teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar.Sereno et al....

and Kaprosuchus

Kaprosuchus

Kaprosuchus is an extinct genus of mahajangasuchid crocodyliform. It is known from a single nearly complete skull collected from the Upper Cretaceous Echkar Formation of Niger. The name means "boar crocodile" from the Greek kapros and souchos in reference to its unusually large caniniform teeth...

, both of which lived during the Late Cretaceous

Late Cretaceous

The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 in Gondwana

Gondwana

In paleogeography, Gondwana , originally Gondwanaland, was the southernmost of two supercontinents that later became parts of the Pangaea supercontinent. It existed from approximately 510 to 180 million years ago . Gondwana is believed to have sutured between ca. 570 and 510 Mya,...

. It is defined as the most inclusive clade containing Mahajangasuchus insignis but not Notosuchus terrestris

Notosuchus

Notosuchus is an extinct genus of South American notosuchian crocodylomorph. It was terrestrial, living approximately 85 million years ago in the Coniacian or Santonian stages of the Late Cretaceous. Remains have been found in the Bajo de la Carpa Formation in Patagonia, Argentina. First named in...

, Simosuchus clarki

Simosuchus

Simosuchus is an extinct genus of notosuchian crocodylomorphs from the Late Cretaceous of Madagascar. It is named for its unusually short skull. Fully grown individuals were about in length. The type species is S. clarki, found from the Maevarano Formation in Mahajanga Province.The teeth of S...

, Araripesuchus gomesii

Araripesuchus

Araripesuchus is a genus of extinct crocodyliform whose species existed during the Cretaceous period of the late Mesozoic era some 125 to 65 million years ago. They are generally considered to be notosuchians , characterized by their varied teeth types and distinct skull elements...

, Baurusuchus pachecoi

Baurusuchus

Baurusuchus is an extinct genus of baurusuchid mesoeucrocodylian from the Late Cretaceous of Brazil. It was a terrestrial predator and scavenger, about 3.5 to 4 meters long. Baurusuchus lived during the Turonian to Santonian stages of the Late Cretaceous Period, in Adamantina Formation, Brazil...

, Peirosaurus torminni

Peirosaurus

Peirosaurus is an extinct genus of peirosaurid crocodylomorph. It is the type genus of the family Peirosauridae. Fossils of the type species P. tormini, first described in 1955, have been found from the Marília Formation in Uberaba, Brazil and date back to the late Maastrichtian stage of the Late...

, Goniopholis crassidens

Goniopholis

Goniopholis is an extinct genus of crocodyliform that lived in North America, Europe and Asia during the Late Jurassic and Early Cretaceous. Being semi-aquatic it is very similar to modern crocodiles...

,
Pholidosaurus schaumbergensis

Pholidosaurus

Pholidosaurus is an extinct genus of neosuchian crocodylomorph. It is the type genus of the family Pholidosauridae. Fossils have been found from Germany and England. The genus is known to have existed during the Berriasian stage of the Early Cretaceous...

, or Crocodylus niloticus. Phyogenetically, Mahajangasuchidae is placed just outside pholidosaurids

Pholidosauridae

Pholidosauridae is an extinct family of mesoeucrocodylian crocodylomorphs. Fossils have been found in the United States, Canada, China, Kyrgyzstan, Africa and Spain...

 and more derived neosuchians.

Defining characters of the family include fused nasals

Nasal bone

The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face, and form, by their junction, "the bridge" of the nose.Each has two surfaces and four borders....

, a jaw articulation below the posterior maxilla

Maxilla

The maxilla is a fusion of two bones along the palatal fissure that form the upper jaw. This is similar to the mandible , which is also a fusion of two halves at the mental symphysis. Sometimes The maxilla (plural: maxillae) is a fusion of two bones along the palatal fissure that form the upper...

ry tooth row, a deep mandibular symphysis that is oriented anterodorsally, and the formation of a hornlike posterodorsal process from the squamosal

Squamosal

The squamosal is a bone of the head of higher vertebrates. It is the principal component of the cheek region in the skull, lying below the temporal series and otic notch and bounded anteriorly by postorbital. Posteriorly, the squamosal articulates with the posterior elements of the palatal complex,...

 and parietal

Parietal

Parietal may refer to:*Parietal placentation*Parietal lobe of the brain*Parietal bone of the skull*Parietal scales of a snake lie in the general region of the parietal bone*Parietal cell in the stomach*Parietal pleura...

 (which is much more pronounced in Kaprosuchus).

Cladogram

Cladogram

A cladogram is a diagram used in cladistics which shows ancestral relations between organisms, to represent the evolutionary tree of life. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational...

 showing the phylogenetic relationship of Mahajangasuchidae within Neosuchia after Sereno and Larsson, 2009:

External links

• ZooBank