Microecosystem
Encyclopedia
Microecosystems can exist in locations which are precisely defined by critical environmental
Natural environment
The natural environment encompasses all living and non-living things occurring naturally on Earth or some region thereof. It is an environment that encompasses the interaction of all living species....

 factors within small or tiny spaces.

Such factors may include temperature
Temperature
Temperature is a physical property of matter that quantitatively expresses the common notions of hot and cold. Objects of low temperature are cold, while various degrees of higher temperatures are referred to as warm or hot...

, pH
PH
In chemistry, pH is a measure of the acidity or basicity of an aqueous solution. Pure water is said to be neutral, with a pH close to 7.0 at . Solutions with a pH less than 7 are said to be acidic and solutions with a pH greater than 7 are basic or alkaline...

, chemical milieu, nutrient
Nutrient
A nutrient is a chemical that an organism needs to live and grow or a substance used in an organism's metabolism which must be taken in from its environment. They are used to build and repair tissues, regulate body processes and are converted to and used as energy...

 supply, presence of symbionts or solid substrates, gaseous atmosphere (aerobic or anaerobic) etc.

Pond microecosystems

These microecosystems with limited water
Water
Water is a chemical substance with the chemical formula H2O. A water molecule contains one oxygen and two hydrogen atoms connected by covalent bonds. Water is a liquid at ambient conditions, but it often co-exists on Earth with its solid state, ice, and gaseous state . Water also exists in a...

 volume are often only of temporary duration and hence colonized by organisms which possess a drought-resistant spore
Spore
In biology, a spore is a reproductive structure that is adapted for dispersal and surviving for extended periods of time in unfavorable conditions. Spores form part of the life cycles of many bacteria, plants, algae, fungi and some protozoa. According to scientist Dr...

 stage in the lifecycle, or by organisms which do not need to live in water continuously. The ecosystem conditions appliying at a typical pond edge can be quite different from those further from shore. Extremely space-limited water ecosystems can be found in, for example, the water collected in bromeliad leaf bases and the "pitchers" of Nepenthes
Nepenthes
The Nepenthes , popularly known as tropical pitcher plants or monkey cups, are a genus of carnivorous plants in the monotypic family Nepenthaceae. The genus comprises roughly 130 species, numerous natural and many cultivated hybrids...

.

Animal gut microecosystems

These include the buccal region (especially cavities in the gingiva
Gingiva
The gingiva , or gums, consists of the mucosal tissue that lies over the mandible and maxilla inside the mouth.-General description:...

), rumen
Rumen
The rumen, also known as a paunch, forms the larger part of the reticulorumen, which is the first chamber in the alimentary canal of ruminant animals. It serves as the primary site for microbial fermentation of ingested feed...

, caecum etc. of mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

ian herbivores or even invertebrate
Invertebrate
An invertebrate is an animal without a backbone. The group includes 97% of all animal species – all animals except those in the chordate subphylum Vertebrata .Invertebrates form a paraphyletic group...

 digestive tracts. In the case of mammalian gastrointestinal microecology, microorganisms such as protozoa, bacteria, as well as curious incompletely defined organisms (such as certain large structurally-complex Selenomonad
Selenomonad
The genus Selenomonas constitutes a group of motile crescent-shaped bacteria within the Veillonellaceae family and includes species living in the gastrointestinal tracts of animals, in particular, the Ruminants....

s, Quinella ovalis "Quin's Oval", Magnoovum eadii "Eadie's Oval", Oscillospira etc.) can exist in the rumen as incredibly complex, highly enriched mixed populations, (see Moir and Masson images ). This type of microecosystem can adjust rapidly to changes in the nutrition or health of the host animal (usually a ruminant such as cow, sheep, goat etc.); see Hungate's "The Rumen and its microbes 1966). Even within a small closed system such as the rumen there may exist a range of ecological conditions: Many organisms live freely in the rumen fluid whereas others require the substrate and metabolic products supplied by the stomach wall tissue with its folds and interstices. Interesting questions are also posed concerning the transfer of the strict anaerobe organisms in the gut microflora/microfauna to the next host generation. Here, mutual licking and coprophagia
Coprophagia
Coprophagia or coprophagy is the consumption of feces, from the Greek κόπρος copros and φαγεῖν phagein . Many animal species practice coprophagia as a matter of course; other species do not normally consume feces but may do so under unusual conditions...

 certainly play important roles.

Soil microecosystems

A typical soil microecosystem may be restricted to less than a millimeter in its total depth range owing to steep variation in humidity and/or atmospheric gas composition. The soil grain size and physical and chemical properties of the substrate may also play important roles. Because of the predominant solid phase in these systems they are notoriously difficult to study microscopically without simultaneously disrupting the fine spatial distribution of their components.

Terrestrial hot-spring microecosystems

These are defined by gradients of water temperature, nutrients, dissolved gases, salt
Salt
In chemistry, salts are ionic compounds that result from the neutralization reaction of an acid and a base. They are composed of cations and anions so that the product is electrically neutral...

 concentrations etc. Along the path of terrestrial water flow the resulting temperature gradient continuum alone may provide many different minute microecosystems, starting with thermophilic bacteria such as Archaea
Archaea
The Archaea are a group of single-celled microorganisms. A single individual or species from this domain is called an archaeon...

 "Archaebacteria" (100+°C), followed by conventional thermophiles (60-100°C), cyanobacteria (blue-green algae) such as the motile filaments of Oscillatoria (30-60°C), protozoa
Protozoa
Protozoa are a diverse group of single-cells eukaryotic organisms, many of which are motile. Throughout history, protozoa have been defined as single-cell protists with animal-like behavior, e.g., movement...

 such as Amoeba, rotifers, then green algae
Green algae
The green algae are the large group of algae from which the embryophytes emerged. As such, they form a paraphyletic group, although the group including both green algae and embryophytes is monophyletic...

 (0-30°C) etc.
Of course other factors than temperature also play important roles. Hot springs can provide classic and straightforward ecosystems for microecology studies as well as providing a haven for hitherto undescribed organisms.

Deep-sea microecosystems

The best known contain rare specialized organisms, found only in the immediate vicinity (sometimes within centimeters) of underwater volcanic vents (or "smokers"). These ecosystems require extremely advanced diving and collection techniques for their scientific exploration.

Closed microecosystem

One that is sealed and completely independent of outside factors, except for temperature and light. A good example would be a plant contained in a sealed jar and submerged under water. No new factors would be able to enter this ecosystem.
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