Nymphalinae is a subfamily of brush-footed butterflies (family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Nymphalidae). Sometimes, the Limenitidinae

Limenitidinae

Limenitidinae is a subfamily of butterflies that includes the admirals and relatives. The common names of many species and genera reference military ranks or – namely the Adoliadini – titles of nobility , in reference to these butterflies' large size, bold pattern and dashing flight...

 (and Biblidinae

Biblidinae

Biblidinae is a subfamily of nymphalid butterflies that includes the tropical brushfoots. This subfamily was sometimes merged within the Limenitidinae, but they are now recognized as quite distinct lineages. In older literature, this subfamily is sometimes called Eurytelinae.As of 2008, there are...

) are included here as further tribe

Tribe (biology)

In biology, a tribe is a taxonomic rank between family and genus. It is sometimes subdivided into subtribes.Some examples include the tribes: Canini, Acalypheae, Hominini, Bombini, and Antidesmeae.-See also:* Biological classification* Rank...

(s), while the Melitaeini

Melitaeini

Melitaeini are a group of brush-footed butterflies. Usually classified as a tribe of the Nymphalinae, they are sometimes raised to subfamily status as Melitaeinae...

 are occasionally considered a distinct subfamily.

Systematics

The 6 usually-recognized tribes of Nymphalinae are here listed in the presumed phylogenetic sequence.

• Coeini
  Coeini
  Coeini is a tribe of brush-footed butterflies.- List of genera :* Baeotus Hemming, 1939* Colobura Billberg, 1820* Historis Hübner, 1819* Pycina Doubleday, 1849* Smyrna Hübner, 1823...
  
   (6-7 genera)
• Nymphalini
  Nymphalini
  Nymphalini is a tribe of nymphalid brush-footed butterflies. Common names include admirals, anglewings, commas, and tortoiseshells, but neither of these is specific to one particular genus....
  
   – anglewings, tortoiseshells and relatives (c.15 genera, 2 fossil)
• Kallimini
  Kallimini
  Kallimini is a tribe of brush-footed butterflies.- List of genera :* Catacroptera Karsch, 1894 – pirates* Doleschallia C. & R...
  
   (c.5 genera)
• Victorinini
  Victorinini
  Victorinini is a tribe of brush-footed butterflies. The four butterfly genera contained in this tribe are Anartia, Metamorpha, Napeocles, and Siproeta....
  
   (4 genera, formerly in Kallimini)
• Junoniini
  Junoniini
  Junoniini is a tribe of nymphalid brush-footed butterflies.- Genera :* Junonia Hübner, 1819 – buckeyes, commodores, "pansies"* Precis Hübner, 1819 – "pansies"* Protogoniomorpha Wallengren, 1857...
  
   (c.5 genera)
• Melitaeini
  Melitaeini
  Melitaeini are a group of brush-footed butterflies. Usually classified as a tribe of the Nymphalinae, they are sometimes raised to subfamily status as Melitaeinae...
  
   – fritillaries (c.25 genera)

Genera incertae sedis

Incertae sedis

, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...

are:

• Rhinopalpa – The Wizard
• Kallimoides Shirôzu & Nakanishi, 1984

• Vanessula Dewitz, 1887

In addition to these, Crenidomimas is sometimes placed here but may actually belong in the Limenitidinae

Limenitidinae

Limenitidinae is a subfamily of butterflies that includes the admirals and relatives. The common names of many species and genera reference military ranks or – namely the Adoliadini – titles of nobility , in reference to these butterflies' large size, bold pattern and dashing flight...

 genus Euryphura

Euryphura

Euryphura is a butterfly genus in the subfamily Limenitidinae.-Species:Listed alphabetically within species groups.*The prophyrion species-group**Euryphura porphyrion **Euryphura togoensis Suffert, 1904...

.

The Late Eocene fossil

Fossil

Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

 genus Lithodryas

Lithodryas

Lithodryas is a prehistoric genus of butterfly in the family Lycaenidae. It was introduced as a replacement for Samuel Hubbard Scudder's genus Lithopsyche which is invalid as a homonym. The type species was found in Tertiary deposits at Florissant....

might belong here, but it is generally assigned to the Lycaenidae

Lycaenidae

The Lycaenidae are the second-largest family of butterflies, with about 6000 species worldwide, whose members are also called gossamer-winged butterflies...

.