Ophthalmosauridae is an extinct family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of thunnosaur ichthyosaur

Ichthyosaur

Ichthyosaurs were giant marine reptiles that resembled fish and dolphins...

s from the Middle Jurassic

Middle Jurassic

The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from 176-161 million years ago. In European lithostratigraphy, rocks of this Middle Jurassic age are called the Dogger....

 to the early Late Cretaceous

Late Cretaceous

The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 (Bajocian

Bajocian

In the geologic timescale, the Bajocian is an age or stage in the Middle Jurassic. It lasted from approximately 171.6 Ma to around 167.7 Ma . The Bajocian age succeeds the Aalenian age and precedes the Bathonian age....

 - Cenomanian

Cenomanian

The Cenomanian is, in the ICS' geological timescale the oldest or earliest age of the Late Cretaceous epoch or the lowest stage of the Upper Cretaceous series. An age is a unit of geochronology: it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding...

) of Asia

Asia

Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

, Australia

Australia

Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

, Europe

Europe

Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, North America

North America

North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 and South America

South America

South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

. Currently, the oldest known ophthalmosaurid is Mollesaurus

Mollesaurus

Mollesaurus is an extinct genus of large ophthalmosaurid ichthyosaur known from northwestern Patagonia of Argentina.-Description:Mollesaurus is known from the holotype MOZ 2282 V, articulated partial skeleton which preserved partial skull and most of the vertebral column...

from the early Bajocian of Argentina

Argentina

Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

. Named by George H. Baur, in 1887, it contains the basal taxa like Ophthalmosaurus

Ophthalmosaurus

Ophthalmosaurus is an ichthyosaur of the Middle to Late Jurassic period , named for its extremely large eyes. It had a graceful 6 metre long dolphin-shaped body, and its almost toothless jaw was well adapted for catching squid...

. Appleby (1956) named the taxon Ophthalmosauria which was followed by some authors, but if these two names are synonyms, Ophthalmosauridae has the priority over Ophthalmosauria.

Phylogeny

Ophthalmosauridae is a node-based taxon defined by Ryosuke Motani (1999) as "the last common ancestor of Brachypterygius extremus and Ophthalmosaurus icenicus and all of its descendants". The definition he proposed for Ophthalmosauria was exactly the same. In this case both defintions are synonyms. The cladogram

Cladogram

A cladogram is a diagram used in cladistics which shows ancestral relations between organisms, to represent the evolutionary tree of life. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational...

 below follows the topology from a 2010 analysis by Patrick S. Druckenmiller and Erin E. Maxwell.
Valentin Fischer, Edwige Masure, Maxim S. Arkhangelsky and Pascal Godefroit (2011) described a new genus of Ophthalmosauridae. They redefined it as "the last common ancestor of Arthropterygius chrisorum and Ophthalmosaurus icenicus and all of its descendants". In this case Ophthalmosauria isn't a junior synonym of Ophthalmosauridae. The cladogram

Cladogram

A cladogram is a diagram used in cladistics which shows ancestral relations between organisms, to represent the evolutionary tree of life. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational...

 below follows Fischer et al. 2011.
*Note: Placement of Ophthalmosauria by definition.