Oskar Hertwig
Encyclopedia

Oscar Hertwig was a German
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

 zoologist and professor
Professor
A professor is a scholarly teacher; the precise meaning of the term varies by country. Literally, professor derives from Latin as a "person who professes" being usually an expert in arts or sciences; a teacher of high rank...

, who also wrote about the theory of evolution circa 1916, over 55 years after Charles Darwin
Charles Darwin
Charles Robert Darwin FRS was an English naturalist. He established that all species of life have descended over time from common ancestry, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection.He published his theory...

's book The Origin of Species
The Origin of Species
Charles Darwin's On the Origin of Species, published on 24 November 1859, is a work of scientific literature which is considered to be the foundation of evolutionary biology. Its full title was On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the...

. He was the elder brother of zoologist-professor Richard Hertwig
Richard Hertwig
Richard Wilhelm Karl Theodor Ritter von Hertwig , also Richard Hertwig or Richard von Hertwig, was a German zoologist and professor of 50 years, notable as the first to describe zygote formation as the fusing of spermatozoa inside the membrane of an egg cell during fertilization.Richard Hertwig was...

 (1850–1937).

The Hertwig brothers were the most eminent scholars of Ernst Haeckel
Ernst Haeckel
The "European War" became known as "The Great War", and it was not until 1920, in the book "The First World War 1914-1918" by Charles à Court Repington, that the term "First World War" was used as the official name for the conflict.-Research:...

 (and Carl Gegenbaur) from the University of Jena. They were independent of Haeckel's philosophical speculations but took his ideas in a positive way to widen their concepts in zoology
Zoology
Zoology |zoölogy]]), is the branch of biology that relates to the animal kingdom, including the structure, embryology, evolution, classification, habits, and distribution of all animals, both living and extinct...

. Initially, between 1879–1883, they performed embryological studies, especially on the theory of the coelom
Coelom
The coelom is a fluid-filled cavity formed within the mesoderm. Coeloms developed in triploblasts but were subsequently lost in several lineages. Loss of coelom is correlated with reduction in body size...

 (1881), the fluid-filled body cavity. These problems were based on the phylogenetic theorems of Haeckel, i.e. the biogenic theory (German = biogenetisches Grundgesetz), and the "gastraea theory".

Within 10 years, the two brothers moved apart to the north and south of Germany
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

.
Oscar Hertwig later became a professor of anatomy
Anatomy
Anatomy is a branch of biology and medicine that is the consideration of the structure of living things. It is a general term that includes human anatomy, animal anatomy , and plant anatomy...

 in 1888 in Berlin
Berlin
Berlin is the capital city of Germany and is one of the 16 states of Germany. With a population of 3.45 million people, Berlin is Germany's largest city. It is the second most populous city proper and the seventh most populous urban area in the European Union...

; however, Richard Hertwig had moved 3 years prior, becoming a professor of zoology in Munich
Munich
Munich The city's motto is "" . Before 2006, it was "Weltstadt mit Herz" . Its native name, , is derived from the Old High German Munichen, meaning "by the monks' place". The city's name derives from the monks of the Benedictine order who founded the city; hence the monk depicted on the city's coat...

 from 1885–1925, at Ludwig Maximilians Universität
Ludwig Maximilians University of Munich
The Ludwig Maximilian University of Munich , commonly known as the University of Munich or LMU, is a university in Munich, Germany...

, where he served the last 40 years of his 50-year career as a professor at 4 universities.

Richard's research focused on protist
Protist
Protists are a diverse group of eukaryotic microorganisms. Historically, protists were treated as the kingdom Protista, which includes mostly unicellular organisms that do not fit into the other kingdoms, but this group is contested in modern taxonomy...

s (the relationship between the nucleus and the plasm = "Kern-Plasma-Relation"), as well as on developmental physiological studies on sea urchins and frogs. He also wrote a leading Zoology textbook.

Oscar Hertwig was a leader in the field of comparative and causal animal-developmental history. He also wrote a leading textbook. He discovered fertilization of sea urchin
Sea urchin
Sea urchins or urchins are small, spiny, globular animals which, with their close kin, such as sand dollars, constitute the class Echinoidea of the echinoderm phylum. They inhabit all oceans. Their shell, or "test", is round and spiny, typically from across. Common colors include black and dull...

s, he recognized the role of the cell nucleus during inheritance and chromosome reduction during meiosis
Meiosis
Meiosis is a special type of cell division necessary for sexual reproduction. The cells produced by meiosis are gametes or spores. The animals' gametes are called sperm and egg cells....

: in 1876, he found that fertilization includes the penetration of a spermatozoon into an egg cell. While Oscar was well interested in developmental biology, he was opposed to chance as assumed in Charles Darwin
Charles Darwin
Charles Robert Darwin FRS was an English naturalist. He established that all species of life have descended over time from common ancestry, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection.He published his theory...

´s theory. His most important theoretical book was: "Das Werden der Organismen, eine Widerlegung der Darwinschen Zufallslehre" (Jena, 1916) (translation: "The Origin of Organisms - a Refutation of Darwin's Theory of Chance").

Hertwig was elected a member of the Royal Swedish Academy of Sciences
Royal Swedish Academy of Sciences
The Royal Swedish Academy of Sciences or Kungliga Vetenskapsakademien is one of the Royal Academies of Sweden. The Academy is an independent, non-governmental scientific organization which acts to promote the sciences, primarily the natural sciences and mathematics.The Academy was founded on 2...

 in 1903.

Oscar Hertwig is known as Oscar Hedwig in the book "Who discovered what when" by David Ellyard. A history of the discovery of fertilization for mammals including scientists like Hertwig and other workers is given by the book "The Mammalian Egg" by Austin.

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