Pagurus
Encyclopedia
Pagurus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of hermit crab
Hermit crab
Hermit crabs are decapod crustaceans of the superfamily Paguroidea. Most of the 1100 species possess an asymmetrical abdomen which is concealed in an empty gastropod shell that is carried around by the hermit crab.-Description:...

s in the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Paguridae
Paguridae
Paguridae is a family of hermit crabs of the order Decapoda. It contains 542 species in over 70 genera:*Acanthopagurus de Saint Laurent, 1968*Agaricochirus McLaughlin, 1981*Alainopaguroides McLaughlin, 1997...

. Like other hermit crabs, their abdomen
Abdomen
In vertebrates such as mammals the abdomen constitutes the part of the body between the thorax and pelvis. The region enclosed by the abdomen is termed the abdominal cavity...

 is not calcified and they use snail shells
Gastropod shell
The gastropod shell is a shell which is part of the body of a gastropod or snail, one kind of mollusc. The gastropod shell is an external skeleton or exoskeleton, which serves not only for muscle attachment, but also for protection from predators and from mechanical damage...

 as portable homes. These marine
Marine (ocean)
Marine is an umbrella term. As an adjective it is usually applicable to things relating to the sea or ocean, such as marine biology, marine ecology and marine geology...

 decapod
Decapoda
The decapods or Decapoda are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crayfish, crabs, lobsters, prawns and shrimp. Most decapods are scavengers. It is estimated that the order contains nearly 15,000 species in around 2,700 genera, with...

 crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s are omnivorous
Omnivore
Omnivores are species that eat both plants and animals as their primary food source...

, but mostly prey on small animals and scavenge
Scavenger
Scavenging is both a carnivorous and herbivorous feeding behavior in which individual scavengers search out dead animal and dead plant biomass on which to feed. The eating of carrion from the same species is referred to as cannibalism. Scavengers play an important role in the ecosystem by...

 carrion
Carrion
Carrion refers to the carcass of a dead animal. Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters include vultures, hawks, eagles, hyenas, Virginia Opossum, Tasmanian Devils, coyotes, Komodo dragons, and burying beetles...

. Trigonocheirus and Pagurixus used to be considered subgenera
Subgenus
In biology, a subgenus is a taxonomic rank directly below genus.In zoology, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the generic name and the specific epithet: e.g. the Tiger Cowry of the Indo-Pacific, Cypraea tigris Linnaeus, which...

 of Pagurus, but the former is nowadays included in Orthopagurus, while the latter has been separated as a distinct genus.

Species

Some 170 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 are presently placed in Pagurus; many others have been placed here at one time or another but are now assigned to other genera of Paguroidea
Hermit crab
Hermit crabs are decapod crustaceans of the superfamily Paguroidea. Most of the 1100 species possess an asymmetrical abdomen which is concealed in an empty gastropod shell that is carried around by the hermit crab.-Description:...

. Usually, a few new species are discovered each year. The following list is current :
  • Pagurus acadianus Benedict, 1901
  • Pagurus alabamensis Rathbun, 1935
  • Pagurus alaini Komai, 1998
  • Pagurus alatus Fabricius, 1775
  • Pagurus albidianthus de Saint Laurent & McLaughlin, 2000
  • Pagurus albus (Benedict, 1892)
  • Pagurus alcocki (Balss, 1911)
  • Pagurus aleuticus (Benedict, 1892)
  • Pagurus anachoretoides Forest, 1966
  • Pagurus anachoretus Risso, 1827
  • Pagurus angustus (Stimpson, 1858)
  • Pagurus annexus McLaughlin & Haig, 1993
  • Pagurus annulipes (Stimpson, 1860)
  • Pagurus arcuatus Squires, 1964
  • Pagurus arenisaxatilis Harvey & McLaughlin, 1991
  • Pagurus armatus
    Pagurus armatus
    Pagurus armatus, the armed hermit crab or black-eyed hermit crab, is a species of hermit crab found in the eastern Pacific Ocean of the United States and British Columbia, Canada.-Description:...

    (Dana, 1851)
  • Pagurus benedicti (Bouvier, 1892)
  • Pagurus beringanus (Benedict, 1892)
  • Pagurus bernhardus
    Pagurus bernhardus
    Pagurus bernhardus is the common marine hermit crab of Europe's Atlantic coasts. It is sometimes referred to as the common hermit crab or soldier crab. It is about long, and is found in both rocky and sandy areas, from the Arctic waters of Iceland, Svalbard and Russia as far south as southern...

    (Linnaeus, 1758)
  • Pagurus boletifer
  • Pagurus boriaustraliensis Morgan, 1990
  • Pagurus bouvieri (Faxon, 1895)
  • Pagurus brachiomastus (Thallwitz, 1892)
  • Pagurus brandti (Benedict, 1892)
  • Pagurus brevidactylus (Stimpson, 1859)
  • Pagurus brucei (Rathbun, 1926)
  • Pagurus bullisi Wass, 1963
  • Pagurus capillatus (Benedict, 1892)
  • Pagurus capsularis McLaughlin, 1997
  • Pagurus carneus (Pocock, 1889)
  • Pagurus carolinensis McLaughlin, 1975
  • Pagurus carpoforminatus (Alcock, 1905)
  • Pagurus caurinus Hart, 1971
  • Pagurus cavicarpus (Paulson, 1875)
  • Pagurus chevreuxi (Bouvier, 1896)
  • Pagurus compressipes (Mieers, 1884)
  • Pagurus comptus White, 1847
  • Pagurus conformis (De Haan, 1849)
  • Pagurus confragosus (Benedict, 1892)
  • Pagurus confusus Komai & Yu, 1999
  • Pagurus constans (Stimpson, 1858)
  • Pagurus cornutus (Benedict, 1892)
  • Pagurus criniticornis (Dana, 1852)
  • Pagurus cuanensis Bell, 1845
  • Pagurus curacaoensis (Benedict, 1892)
  • Pagurus dalli (Benedict, 1892)
  • Pagurus dartevellei (Forest, 1958)
  • Pagurus decimbranchiae Komai & Osawa, 2001
  • Pagurus defensus (Benedict, 1892)
  • Pagurus delsolari Haig, 1974
  • Pagurus dissimilis (A. Milne-Edwards & Bouvier, 1893)
  • Pagurus edwardsii (Dana, 1852)
  • Pagurus emmersoni McLaughlin & Forest, 1999
  • Pagurus erythrogrammus Komai, 2003
  • Pagurus excavatus (Herbst, 1791)
  • Pagurus exiguus (Melin, 1939)
  • Pagurus exilis (Benedict, 1892)
  • Pagurus filholi (De Man, 1887)
  • Pagurus fimbriatus Forest, 1966
  • Pagurus forbesii Bell, 1845
  • Pagurus forceps H. Milne-Edwards, 1836
  • Pagurus fungiformis Komai & Rahayu, 2004
  • Pagurus gladius (Benedict, 1892)
  • Pagurus gordonae (Forest, 1956)
  • Pagurus gracilipes (Stimpson, 1858)
  • Pagurus granosimanus (Stimpson, 1859)
  • Pagurus gymnodactylus Lemaitre, 1982
  • Pagurus hartae (McLaughlin & Jensen, 1996)
  • Pagurus heblingi Nucci & Melo, 2003
  • Pagurus hedleyi (Grant & McCulloch, 1906)
  • Pagurus hemphilli (Benedict, 1892)
  • Pagurus hirsutiusculus
    Pagurus hirsutiusculus
    Pagurus hirsutiusculus is a species of hermit crab, commonly called the hairy hermit crab. It lives from the Bering Strait south to California and Japan, from the intertidal zone to a depth of .-Description:...

    (Dana, 1851)
  • Pagurus holmi Ng & McLaughlin, 2009
  • Pagurus ikedai Lemaitre & Watabe, 2005
  • Pagurus imafukui McLaughlin & Konishi, 1994
  • Pagurus imaii (Yokoya, 1939)
  • Pagurus imarpe Haig, 1974
  • Pagurus impressus (Benedict, 1892)
  • Pagurus indicus Sarojini & Nagabhushanam, 1972
  • Pagurus insulae Asakura, 1991
  • Pagurus investigatoris (Alcock, 1905)
  • Pagurus iridocarpus de Saint Laurent & McLaughlin, 2000
  • Pagurus irregularis (A. Milne-Edwards & Bouvier, 1892)
  • Pagurus japonicus (Stimpson, 1858)
  • Pagurus kaiensis McLaughlin, 1997
  • Pagurus kennerlyi (Stimpson, 1854)
  • Pagurus kulkarnii Sankolli, 1962
  • Pagurus lanuginosus De Haan, 1849
  • Pagurus laurentae Forest, 1978
  • Pagurus lepidus (Bouvier, 1898)
  • Pagurus leptonyx Forest & de Saint Laurent, 1968
  • Pagurus limatulus Fausto Filho, 1970
  • Pagurus liochele (Barnard, 1947)
  • Pagurus longicarpus
    Pagurus longicarpus
    Pagurus longicarpus, the long-clawed hermit crab, is a small common subtidal decapod ranging out to deep found living in shells from periwinkles, oyster drills, and eastern mud snails. It is found from Nova Scotia to Florida and Texas....

    Say, 1817
  • Pagurus longimanus Wass, 1963
  • Pagurus lophochela Komai, 1999
  • Pagurus luticola Komai & Chan, 2006
  • Pagurus macardlei (Alcock, 1905)
  • Pagurus maclaughlinae Garcia-Gomez, 1982
  • Pagurus maculosus Komai & Imafuku, 1996
  • Pagurus malloryi Schweitzer & Feldmann, 2001
  • Pagurus marshi Benedict, 1901
  • Pagurus mbizi (Forest, 1955)
  • Pagurus meloi Lemaitre & Cruz Castano, 2004
  • Pagurus mertensii Brandt, 1851
  • Pagurus middendorffii Brandt, 1851
  • Pagurus minutus Hess, 1865
  • Pagurus moluccensis Haig & Ball, 1988
  • Pagurus nanodes Haig & Harvey, 1991
  • Pagurus nesiotes Haig & McLaughlin, 1991
  • Pagurus nigrivittatus Komai, 2003
  • Pagurus nigrofascia Komai, 1996
  • Pagurus nipponensis (Yokoya, 1933)
  • Pagurus nisari Siddiqui & Komai, 2008
  • Pagurus novizealandiae (Dana, 1852)
  • Pagurus ochotensis Brandt, 1851
  • Pagurus parvispina Komai, 1997
  • Pagurus parvus (Benedict, 1892)
  • Pagurus pectinatus (Stimpson, 1858)
  • Pagurus pergranulatus (Henderson, 1896)
  • Pagurus perlatus H. Milne-Edwards, 1848
  • Pagurus pilosipes (Stimpson, 1858)
  • Pagurus pilsbryi Roberts, 1962
  • Pagurus pitagsaleei McLaughlin, 2002
  • Pagurus politus (Smith, 1882)
  • Pagurus pollicaris Say, 1817
  • Pagurus prideaux
    Pagurus prideaux
    Pagurus prideaux is a species of hermit crab in the family Paguridae. It is found in shallow waters off the northwest coast of Europe and usually lives symbiotically with the sea anemone Adamsia palliata.-Description:...

    Leach, 1815
  • Pagurus protuberocarpus McLaughlin, 1982
  • Pagurus provenzanoi Forest & de Saint Laurent, 1968
  • Pagurus proximus Komai, 2000
  • Pagurus pubescens Krøyer, 1838
  • Pagurus pubescentulus (A. Milne-Edwards & Bouvier, 1892)
  • Pagurus pulchellus (A. Milne-Edwards & Bouvier, 1892)
  • Pagurus pycnacanthus (Forest, 1955)
  • Pagurus quaylei Hart, 1971
  • Pagurus quinquelineatus Komai, 2003
  • Pagurus rathbuni (Benedict, 1892)
  • Pagurus redondoensis Wicksten, 1982
  • Pagurus retrorsimanus Wicksten & McLaughlin, 1998
  • Pagurus rhabdotus Haig & Harvey, 1991
  • Pagurus rotundimanus Wass, 1963
  • Pagurus ruber (A. Milne-Edwards & Bouvier, 1892)
  • Pagurus rubrior Komai, 2003
  • Pagurus samoensis (Ortmann, 1892)
  • Pagurus samuelis
    Pagurus samuelis
    Pagurus samuelis, the blueband hermit crab, is a species of hermit crab from the west coast of North America, and the most common hermit crab in California. It is a small species, with distinctive blue bands on its legs...

    (Stimpson, 1857)
  • Pagurus setosus (Benedict, 1892)
  • Pagurus similimanus (Balss, 1921)
  • Pagurus similis (Ortmann, 1892)
  • Pagurus simulans Komai, 2000
  • Pagurus sinuatus
    Pagurus sinuatus
    Pagurus sinuatus is a large species of hermit crab found in Australia and the Kermadec Islands. It is red or orange in colour with coloured bands on the legs and patches on the body.-Description:...

    (Stimpson, 1858)
  • Pagurus smithi (Benedict, 1892)
  • Pagurus souriei (Forest, 1952)
  • Pagurus spighti McLaughlin & Haig, 1993
  • Pagurus spilocarpus Haig, 1977
  • Pagurus spina Komai, 1994
  • Pagurus spinulentus (Henderson, 1888)
  • Pagurus stevensae Hart, 1971
  • Pagurus sticticus McLaughlin, 2008
  • Pagurus stimpsoni (A. Milne-Edwards & Bouvier, 1893)
  • Pagurus tanneri (Benedict, 1892)
  • Pagurus townsendi (Benedict, 1892)
  • Pagurus traversi (Filhol, 1885)
  • Pagurus triangularis (Chevreux & Bouvier, 1892)
  • Pagurus trichocerus Forest & de Saint Laurent, 1968
  • Pagurus trigonocheirus (Stimpson, 1858)
  • Pagurus tristanensis (Henderson, 1888)
  • Pagurus tuberculosus Harvey, 1998
  • Pagurus undosus (Benedict, 1892)
  • Pagurus venturensis Coffin, 1957
  • Pagurus vetaultae Harvey & McLaughlin, 1991
  • Pagurus villosus Nicolet, 1849
  • Pagurus virgulatus Haig & Harvey, 1991

The following are all nomina nuda
Nomen nudum
The phrase nomen nudum is a Latin term, meaning "naked name", used in taxonomy...

. They have never been validly described as new species, but only mentioned under these names:
  • Pagurus bunomanus Glassell, 1937
  • Pagurus crenatus Hope, 1851
  • Pagurus cuanensis Thompson, 1844
  • Pagurus cultratus White, 1847
  • Pagurus hirtimanus White, 1847
  • Pagurus hyndmani Thompson, 1844
  • Pagurus laevis Thompson, 1844
  • Pagurus ulidiae Thompson, 1843
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