Peteinosaurus
Encyclopedia
Peteinosaurus was a prehistoric reptile genus belonging to the Pterosauria. It lived in the late Triassic
period in the middle Norian
(about 221-210 million years ago).
is Peteinosaurus zambellii. The genus name is derived from Greek peteinos, "winged" and sauros, "lizard", the latter being used to indicate any saurian. The specific name, zambellii, honours Rocco Zambelli, the curator of the Bergamo
natural history museum.
, Italy
. The first fossil, the holotype
MCSNB 2886, is fragmentary and disarticulated. The second, the articulated paratype
MCSNB 3359, lacks any diagnostic features of Peteinosaurus and thus might be a different species. This paratype has a long tail (20 cm) made more stiff by long extensions of the vertebrae; this feature is common among pterosaurs of the Triassic. The third exemplar is MCSNB 3496, another fragmentary skeleton. All specimina are those of subadults and of none has the skull been preserved.
Like most pterosaurs, Peteinosaurus had bones that were strong but very light. Peteinosaurus is trimorphodontic, with three types of conical teeth. An insectivorous lifestyle has been attributed to Peteinosaurus. The fifth toe of Peteinosaurus was long and clawless. Its joint allowed it to flex in a different plane than the other phalanges, an adaptation whose function is unknown.
whose wingspan exceeded twenty feet. Its wings were also proportionally smaller than those of later pterosaurs, as its wing length was only twice the length of the hindlimb. All other known pterosaurs have wingspans at least three times the length of their hindlimbs. It also had single cusped teeth that lacked the specialized heterodonty present in the other Italian Triassic pterosaur genus, Eudimorphodon
.
All these factors converge to hint that Peteinosaurus belongs to a group that possibly represents the most basal known pterosaurs: the Dimorphodontidae
, to which it was assigned in 1988 by Robert L. Carroll
. The only other known member of that group is the later genus Dimorphodon
, which lent its name to the family including both genera. Later cladistic analyses however, have not shown a close connection between the two forms. Nevertheless the possible basal position of Peteinosaurus has been affirmed by Fabio Marco Dalla Vecchia who suggested that Preondactylus
, according to David Unwin the most basal pterosaur, might be a subjective junior synonym of Peteinosaurus. A 2010 cladistic analysis by Brian Andres and colleagues placed Peteinosaurus in Lonchognatha which includes Eudimorphodon and Austriadactylus
as more basal.
Triassic
The Triassic is a geologic period and system that extends from about 250 to 200 Mya . As the first period of the Mesozoic Era, the Triassic follows the Permian and is followed by the Jurassic. Both the start and end of the Triassic are marked by major extinction events...
period in the middle Norian
Norian
The Norian is a division of the Triassic geological period. It has the rank of an age or stage . The Norian lasted from 216.5 ± 2.0 to 203.6 ± 1.5 million years ago. It was preceded by the Carnian and succeeded by the Rhaetian.-Stratigraphic definitions:The Norian was named after the Noric Alps in...
(about 221-210 million years ago).
Name
The genus has been described by the German paleontologist Rupert Wild in 1978. The type speciesType species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...
is Peteinosaurus zambellii. The genus name is derived from Greek peteinos, "winged" and sauros, "lizard", the latter being used to indicate any saurian. The specific name, zambellii, honours Rocco Zambelli, the curator of the Bergamo
Bergamo
Bergamo is a town and comune in Lombardy, Italy, about 40 km northeast of Milan. The comune is home to over 120,000 inhabitants. It is served by the Orio al Serio Airport, which also serves the Province of Bergamo, and to a lesser extent the metropolitan area of Milan...
natural history museum.
Specimens
Three fossils have been found near CeneCene
Cene is a comune in the Province of Bergamo in the Italian region of Lombardy, located about 60 km northeast of Milan and about 15 km northeast of Bergamo...
, Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...
. The first fossil, the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
MCSNB 2886, is fragmentary and disarticulated. The second, the articulated paratype
Paratype
Paratype is a technical term used in the scientific naming of species and other taxa of organisms. The exact meaning of the term paratype when it is used in zoology is not the same as the meaning when it is used in botany...
MCSNB 3359, lacks any diagnostic features of Peteinosaurus and thus might be a different species. This paratype has a long tail (20 cm) made more stiff by long extensions of the vertebrae; this feature is common among pterosaurs of the Triassic. The third exemplar is MCSNB 3496, another fragmentary skeleton. All specimina are those of subadults and of none has the skull been preserved.
Like most pterosaurs, Peteinosaurus had bones that were strong but very light. Peteinosaurus is trimorphodontic, with three types of conical teeth. An insectivorous lifestyle has been attributed to Peteinosaurus. The fifth toe of Peteinosaurus was long and clawless. Its joint allowed it to flex in a different plane than the other phalanges, an adaptation whose function is unknown.
Phylogeny and classification
Peteinosaurus is one of the oldest-known pterosaurs, and at a mere sixty centimetres, had a tiny wingspan when compared to some later genera, such as PteranodonPteranodon
Pteranodon , from the Late Cretaceous geological period of North America in present day Kansas, Alabama, Nebraska, Wyoming, and South Dakota, was one of the largest pterosaur genera and had a maximum wingspan of over...
whose wingspan exceeded twenty feet. Its wings were also proportionally smaller than those of later pterosaurs, as its wing length was only twice the length of the hindlimb. All other known pterosaurs have wingspans at least three times the length of their hindlimbs. It also had single cusped teeth that lacked the specialized heterodonty present in the other Italian Triassic pterosaur genus, Eudimorphodon
Eudimorphodon
Eudimorphodon was a pterosaur that was discovered in 1973 by Mario Pandolfi near Bergamo, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic , making Eudimorphodon the oldest pterosaur then known...
.
All these factors converge to hint that Peteinosaurus belongs to a group that possibly represents the most basal known pterosaurs: the Dimorphodontidae
Dimorphodontidae
Dimorphodontidae is a family of early "rhamphorhynchoid" pterosaurs named after Dimorphodon, that lived in the Late Triassic to Early Jurassic....
, to which it was assigned in 1988 by Robert L. Carroll
Robert L. Carroll
Robert Lynn Carroll is a vertebrate paleontologist who specialises in Paleozoic and Mesozoic amphibians and reptiles.Carroll was an only child and grew up on a farm near Lansing, Michigan...
. The only other known member of that group is the later genus Dimorphodon
Dimorphodon
Dimorphodon was a genus of medium-sized pterosaur from the early Jurassic Period. It was named by paleontologist Richard Owen in 1859. Dimorphodon means "two-form tooth", derived from Greek δι/di meaning 'two', μορφη/morphe meaning 'shape' and οδων/odon meaning 'tooth', referring to the fact that...
, which lent its name to the family including both genera. Later cladistic analyses however, have not shown a close connection between the two forms. Nevertheless the possible basal position of Peteinosaurus has been affirmed by Fabio Marco Dalla Vecchia who suggested that Preondactylus
Preondactylus
Preondactylus is a genus of long-tailed pterosaur from the Late Triassic that inhabited what is now Italy. It was discovered by Nando Buffarini in 1982 near Udine in the Preone valley of the Italian Alps.-Anatomy:...
, according to David Unwin the most basal pterosaur, might be a subjective junior synonym of Peteinosaurus. A 2010 cladistic analysis by Brian Andres and colleagues placed Peteinosaurus in Lonchognatha which includes Eudimorphodon and Austriadactylus
Austriadactylus
Austriadactylus is a genus of "rhamphorhynchoid" pterosaur. The fossil remains were unearthed in Late Triassic rocks of Austria.The genus was named in 2002 by Fabio Marco Dalla Vecchia e.a.. The type species is Austriadactylus cristatus. The genus name is derived from Latin Austria and Greek...
as more basal.
Further reading
- Haines, T., and P. Chambers. The Complete Guide to Prehistoric Life. Richmond Hill, Ontario, Canada: Firefly Books Ltd., 2006. Pg. 72. ISBN 1554071259.