Picris echioides
Encyclopedia
Picris echioides, bristly ox-tongue, is a stiff annual
Annual plant
An annual plant is a plant that usually germinates, flowers, and dies in a year or season. True annuals will only live longer than a year if they are prevented from setting seed...

 or biennial
Biennial plant
A biennial plant is a flowering plant that takes two years to complete its biological lifecycle. In the first year the plant grows leaves, stems, and roots , then it enters a period of dormancy over the colder months. Usually the stem remains very short and the leaves are low to the ground, forming...

 herb
Herb
Except in botanical usage, an herb is "any plant with leaves, seeds, or flowers used for flavoring, food, medicine, or perfume" or "a part of such a plant as used in cooking"...

 native to Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 and North Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

. It was traditionally used as an antihelminthic treatment .

Description

P. echioides may grow up to 90 cm (35 in) tall, with a thick, furrowed stem and spreading branches. The leaves are 10–20 cm (3.9–7.9 in) long, oblanceolate with a short petiole
Petiole (botany)
In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...

. The leaves, branches and stem are all covered in thick bristle
Bristle
A bristle is a stiff hair or feather. Also used are synthetic materials such as nylon in items such as brooms and sweepers. Bristles are often used to make brushes for cleaning uses, as they are strongly abrasive; common examples include the toothbrush and toilet brush...

s. The inflorescence
Inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Strictly, it is the part of the shoot of seed plants where flowers are formed and which is accordingly modified...

s are 2–3.5 cm (0.78740157480315–1.4 in) wide and subtended by between 3 and 5 large ovate-cordate involucral bracts . These large bracts are the defining feature of the genus Helminthotheca
Helminthotheca
The genus Helminthotheca is an genus, which belongs to the family Asteraceae. Its members are often treated as species of other genera:-List of species:* Helminthotheca aculeata → Picris aculeata...

, to which P. echioides may be assigned .

A number of infraspecific taxa are recognised, varying in their leaf shape .

Distribution

Picris echioides is native to the Mediterranean Basin
Mediterranean Basin
In biogeography, the Mediterranean Basin refers to the lands around the Mediterranean Sea that have a Mediterranean climate, with mild, rainy winters and hot, dry summers, which supports characteristic Mediterranean forests, woodlands, and scrub vegetation...

, but has become widely naturalised
Naturalisation (biology)
In biology, naturalisation is any process by which a non-native organism spreads into the wild and its reproduction is sufficient to maintain its population. Such populations are said to be naturalised....

 outside that range. In the British Isles
British Isles
The British Isles are a group of islands off the northwest coast of continental Europe that include the islands of Great Britain and Ireland and over six thousand smaller isles. There are two sovereign states located on the islands: the United Kingdom of Great Britain and Northern Ireland and...

, it is widely distributed in the south and east, but more patchily distributed to the north and west . In Northern Ireland
Northern Ireland
Northern Ireland is one of the four countries of the United Kingdom. Situated in the north-east of the island of Ireland, it shares a border with the Republic of Ireland to the south and west...

, P. echioides is only found on the north side of Belfast Lough
Belfast Lough
Belfast Lough is a large, natural intertidal sea lough at the mouth of the River Lagan on the east coast of Northern Ireland. The inner part of the lough comprises a series of mudflats and lagoons. The outer lough is restricted to mainly rocky shores with some small sandy bays...

 .

It has been introduced to North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

, where it can now be found from Nova Scotia
Nova Scotia
Nova Scotia is one of Canada's three Maritime provinces and is the most populous province in Atlantic Canada. The name of the province is Latin for "New Scotland," but "Nova Scotia" is the recognized, English-language name of the province. The provincial capital is Halifax. Nova Scotia is the...

 to British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

 and California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 .

Etymology

The specific epithet echioides comes from the resemblance of the leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 of this species to those of Echium vulgare .
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