Silene spaldingii
Encyclopedia
Silene spaldingii is a rare species of flowering plant in the pink family
Caryophyllaceae
The Caryophyllaceae, commonly called the pink family or carnation family, is a family of flowering plants. It is included in the dicotyledon order Caryophyllales in the APG III system, alongside 33 other families, including Amaranthaceae, Cactaceae and Polygonaceae...

 known by the common names Spalding's catchfly and Spalding's campion. It is native to eastern Washington, and eastern Oregon
Oregon
Oregon is a state in the Pacific Northwest region of the United States. It is located on the Pacific coast, with Washington to the north, California to the south, Nevada on the southeast and Idaho to the east. The Columbia and Snake rivers delineate much of Oregon's northern and eastern...

 in the United States, as well as northern Idaho
Idaho
Idaho is a state in the Rocky Mountain area of the United States. The state's largest city and capital is Boise. Residents are called "Idahoans". Idaho was admitted to the Union on July 3, 1890, as the 43rd state....

 and northern Montana
Montana
Montana is a state in the Western United States. The western third of Montana contains numerous mountain ranges. Smaller, "island ranges" are found in the central third of the state, for a total of 77 named ranges of the Rocky Mountains. This geographical fact is reflected in the state's name,...

, where its distribution extends just into British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

, Canada. It is limited to the last remaining stretches of pristine prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

 grassland
Grassland
Grasslands are areas where the vegetation is dominated by grasses and other herbaceous plants . However, sedge and rush families can also be found. Grasslands occur naturally on all continents except Antarctica...

 in this region. It is threatened by the degradation and loss of its remaining habitat. It is federally listed as a threatened species in the United States and it is designated endangered by Canada's COSEWIC.

This is a perennial herb producing many stems and shoots from a thick taproot
Taproot
A taproot is an enlarged, somewhat straight to tapering plant root that grows vertically downward. It forms a center from which other roots sprout laterally.Plants with taproots are difficult to transplant...

 and woody, branching caudex
Caudex
A caudex is a form of stem morphology appearing as a thickened, short, perennial stem that is either underground or near ground level . It may be swollen for the purpose of water storage, especially in xerophytes...

. The stems grow erect 20 to 60 centimeters in maximum height. They are somewhat hairy and sticky in texture. Lance-shaped leaves occur in pairs along the stems, each blade up to 7 centimeters long. The inflorescence
Inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Strictly, it is the part of the shoot of seed plants where flowers are formed and which is accordingly modified...

 is an open cyme of flowers with greenish white petals. The base of the flower is enclosed in a tubular 10-veined calyx of sepal
Sepal
A sepal is a part of the flower of angiosperms . Collectively the sepals form the calyx, which is the outermost whorl of parts that form a flower. Usually green, sepals have the typical function of protecting the petals when the flower is in bud...

s. Blooming occurs in June and July. Flowers are pollinated by the bumblebee
Bumblebee
A bumble bee is any member of the bee genus Bombus, in the family Apidae. There are over 250 known species, existing primarily in the Northern Hemisphere although they are common in New Zealand and in the Australian state of Tasmania.Bumble bees are social insects that are characterised by black...

 Bombus fervidus
Bombus fervidus
Bombus fervidus is a species of bumblebee native to North America....

.

This plant is present in several regions in the northwestern United States and far southern British Columbia, such as the Palouse
Palouse
The Palouse is a region of the northwestern United States, encompassing parts of southeastern Washington, north central Idaho and, in some definitions, extending south into northeast Oregon. It is a major agricultural area, primarily producing wheat and legumes...

, the Channeled Scablands
Channeled scablands
The Channeled Scablands are a unique geological erosion feature in the U.S. state of Washington. They were created by the cataclysmic Missoula Floods that swept periodically across eastern Washington and down the Columbia River Plateau during the Pleistocene epoch. Geologist J Harlen Bretz coined...

, and the Blue Mountains ecoregion
Blue Mountains (ecoregion)
The Blue Mountains ecoregion is a Level III ecoregion designated by the United States Environmental Protection Agency in the U.S. state of Oregon, with small areas over the state border in Idaho and southeastern Washington...

, including the Zumwalt Prairie
Zumwalt Prairie
Zumwalt Prairie is a grassland area located in Wallowa County in northeast Oregon, United States. Measuring , much of the land is used for agriculture, with some portions protected as the Zumwalt Prairie Preserve owned by The Nature Conservancy...

. The latter contains the largest population, which has over 10,000 individuals. A large percentage of the populations are located in Montana. Recent surveys have led to the discovery of additional subpopulations and populations.

This species distribution has been reduced to patchy, geographically isolated fragments mainly due to the loss of its habitat to agriculture. This is an ongoing threat. Other threats include grazing
Grazing
Grazing generally describes a type of feeding, in which a herbivore feeds on plants , and also on other multicellular autotrophs...

 by livestock
Livestock
Livestock refers to one or more domesticated animals raised in an agricultural setting to produce commodities such as food, fiber and labor. The term "livestock" as used in this article does not include poultry or farmed fish; however the inclusion of these, especially poultry, within the meaning...

 and wildlife, and the invasion
Invasive species
"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....

 of introduced species
Introduced species
An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...

 of plants, two types of disturbance the plant does not tolerate. Other threats include the lack of a normal fire regime
Fire regime
A fire regime is the pattern, frequency and intensity of the bushfires and wildfires that prevails in an area. It is an integral part of fire ecology, and renewal for certain types of ecosystems. If fires are too frequent, plants may be killed before they have matured, or before they have set...

, insect damage, drought
Drought
A drought is an extended period of months or years when a region notes a deficiency in its water supply. Generally, this occurs when a region receives consistently below average precipitation. It can have a substantial impact on the ecosystem and agriculture of the affected region...

, and climate change. Gravel
Gravel
Gravel is composed of unconsolidated rock fragments that have a general particle size range and include size classes from granule- to boulder-sized fragments. Gravel can be sub-categorized into granule and cobble...

 mining, herbicide
Herbicide
Herbicides, also commonly known as weedkillers, are pesticides used to kill unwanted plants. Selective herbicides kill specific targets while leaving the desired crop relatively unharmed. Some of these act by interfering with the growth of the weed and are often synthetic "imitations" of plant...

s, and off-road vehicle
Off-road vehicle
An off-road vehicle is considered to be any type of vehicle which is capable of driving on and off paved or gravel surface. It is generally characterized by having large tires with deep, open treads, a flexible suspension, or even caterpillar tracks...

s are threats in some areas. Populations are generally small, making them vulnerable. For example, a population consisting of a single plant was eliminated when it was buried during road construction in Idaho.
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