Sinosaurosphargis
Encyclopedia
Sinosaurosphargis is an extinct
Extinction
In biology and ecology, extinction is the end of an organism or of a group of organisms , normally a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point...

 genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of non-sauropterygian marine reptile known from the lower Middle Triassic
Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided. It spans the time between 245 ± 1.5 Ma and 228 ± 2 Ma...

 (Pelsonian substage, Anisian
Anisian
In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from 245 million years ago until 237 million years ago, approximately...

 stage) of Yangmazhai, Luoping County
Luoping County
Luoping County is located in Qujing Prefecture, Yunnan, China.The county of Luoping in eastern Yunnan is noted for its beautiful scenery in spring, when its fields of canola flowers are in full bloom, surrounding the area's mountains with a sea of golden flowers to spectacular effect.Located near...

 of Yunnan Province
Yunnan
Yunnan is a province of the People's Republic of China, located in the far southwest of the country spanning approximately and with a population of 45.7 million . The capital of the province is Kunming. The province borders Burma, Laos, and Vietnam.Yunnan is situated in a mountainous area, with...

, southwestern China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

. It is known from the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 IVPP
Institute of Vertebrate Paleontology and Paleoanthropology
The Institute of Vertebrate Paleontology and Paleoanthropology of China is a prominent research institution and collections repository for fossils, including many dinosaur and pterosaurand cat poo specimens...

 V 17040, which consists of nearly complete skeleton
Skeleton
The skeleton is the body part that forms the supporting structure of an organism. There are two different skeletal types: the exoskeleton, which is the stable outer shell of an organism, and the endoskeleton, which forms the support structure inside the body.In a figurative sense, skeleton can...

 and skull
Skull
The skull is a bony structure in the head of many animals that supports the structures of the face and forms a cavity for the brain.The skull is composed of two parts: the cranium and the mandible. A skull without a mandible is only a cranium. Animals that have skulls are called craniates...

, from the paratype
Paratype
Paratype is a technical term used in the scientific naming of species and other taxa of organisms. The exact meaning of the term paratype when it is used in zoology is not the same as the meaning when it is used in botany...

 IVPP V 16076, dorsal vertebra, rib
Rib
In vertebrate anatomy, ribs are the long curved bones which form the rib cage. In most vertebrates, ribs surround the chest, enabling the lungs to expand and thus facilitate breathing by expanding the chest cavity. They serve to protect the lungs, heart, and other internal organs of the thorax...

s, with osteoderm
Osteoderm
Osteoderms are bony deposits forming scales, plates or other structures in the dermal layers of the skin. Osteoderms are found in many groups of extant and extinct reptiles, including lizards, various groups of dinosaurs , crocodilians, phytosaurs, aetosaurs, placodonts, and hupehsuchians...

s and gastral rib fragments and from the referred material ZMNH M 8797, a right forelimb
Forelimb
A forelimb is an anterior limb on an animal's body. When referring to quadrupeds , the term foreleg is often instead used....

. It was found in the new Middle Triassic
Triassic
The Triassic is a geologic period and system that extends from about 250 to 200 Mya . As the first period of the Mesozoic Era, the Triassic follows the Permian and is followed by the Jurassic. Both the start and end of the Triassic are marked by major extinction events...

 Lagerstätte
Lagerstätte
A Lagerstätte is a sedimentary deposit that exhibits extraordinary fossil richness or completeness.Palaeontologists distinguish two kinds....

 of the Guanling Formation
Guanling Formation
The Guanling Formation is a Mesozoic geologic formation. Plesiosaur remains are among the fossils that have been recovered from its strata....

. It was first named by Chun Li, Olivier Rieppel, Xiao-Chun Wu, Li-Jun Zhao and Li-Ting Wang in 2011
2011 in paleontology
-Lobopods:-Arachnids:-Insects:-Newly named jawless vertebrates:-Newly named acanthodian:-Newly named cartilaginous fishes:-Newly named bony fishes:-Newly named lepospondyls:-Newly named temnospondyls:-Newly named lissamphibians:...

 and the type species is Sinosaurosphargis yunguiensis. The generic name comes from Sino, from China; Sauro, sauria
Sauria
Sauria is a clade of reptiles that includes all living diapsids, as well as their common ancestor and all its extinct descendants. The ancestral saurian was probably a small lizard-like creature living in the Permian Period...

n and sphargis, turtle-like (Sphargis is a junior synonym of Dermochelys). The specific name is derived from Yunnan
Yunnan
Yunnan is a province of the People's Republic of China, located in the far southwest of the country spanning approximately and with a population of 45.7 million . The capital of the province is Kunming. The province borders Burma, Laos, and Vietnam.Yunnan is situated in a mountainous area, with...

 and Guizhou
Guizhou
' is a province of the People's Republic of China located in the southwestern part of the country. Its provincial capital city is Guiyang.- History :...

 Provinces, where the fossils were found. It is closely related to the genus Saurosphargis and is similar in appearance to placodont
Placodont
Placodonts were a group of marine reptiles that lived during the Triassic period, becoming extinct at the end of the period. It is believed that they were part of Sauropterygia, the group that includes Plesiosaurs...

s.

Phylogeny

Cladogram
Cladogram
A cladogram is a diagram used in cladistics which shows ancestral relations between organisms, to represent the evolutionary tree of life. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational...

 after Li et al., 2011:
Without Ichthyopterygia
Ichthyopterygia
Ichthyopterygia was a designation introduced by Sir Richard Owen in 1840 to designate the Jurassic ichthyosaurs that were known at the time, but the term is now used more often for both true Ichthyosauria and their more primitive early and middle Triassic ancestors.Basal ichthyopterygians were...

:
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