Tapejara is a genus

Genus

In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of Brazil

Brazil

Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

ian pterosaur

Pterosaur

Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

 from the Cretaceous

Cretaceous

The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...

 Period (Santana Formation

Santana Formation

The Santana Formation is a geologic Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau...

, dating to about 108 Ma ago). Tapejara crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head.

Species and classification

The type species and only one currently recognized as valid by most researchers, T. wellnhoferi, is the smallest species to have been assigned to Tapejara and does not preserve evidence of soft-tissue crest extensions. The specific name honours German paleontologist Peter Wellnhofer

Peter Wellnhofer

Peter Wellnhofer is a German paleontologist at the "Bayerische Staatssammlung fur Paläontologie" in Munich. He is best known for his work on the various fossil specimens of Archaeopteryx or "Urvogel", the first known bird...

. A second species, originally named Tapejara imperator ("emperor"), is much larger and possessed a crest made up of distinctively long prongs, projecting from the rounded snout crest and the back of the skull, which supported a large, possibly rounded sail-like crest of keratin. A third species, Tapejara navigans ("sailing"), was mid-sized and sported a similar crest to T. imperator, though narrower and more dome-shaped, that lacked the backwards-pointing bony support prong.

Several studies in 2007 showed that T. imperator and T. navigans are too different from T. wellnhoferi and therefore require their own genus names. The species T. imperator was given its own genus, Tupandactylus

Tupandactylus

Tupandactylus is a genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Crato Formation of Brazil. It is notable for its large cranial crest, composed partly of bone and partly of soft tissue...

, by Kellner and Campos. Unwin and Martill found that T. imperator and T. navigans belong in the same genus, and named them Ingridia imperator and I. navigans, respectively. The genus name honoured Wellnhofer's late wife Ingrid. Because Tupandactylus was named first, it retains priority over the name Ingridia. To complicate matters, both the name Tupandactylus and Ingridia used the former Tapejara imperator as their type species

Type species

In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

. The scientists who described Tupandactylus did not name a Tupandactylus navigans (but instead suggested it was synonymous to Tupandactylus imperator), and Tapejara navigans was not formally reclassified as a distinct species of Tupandactylus until 2011.

Paleobiology

Comparisons between the scleral ring

Sclerotic ring

Sclerotic rings are rings of bone found in the eyes of several groups of vertebrate animals, except for mammals and crocodilians. They can be made up of single bones or small bones together. They are believed to have a role in supporting the eye, especially in animals whose eyes are not spherical,...

s of Tapejara and modern birds and reptiles suggest that it may have been cathemeral

Cathemeral

A cathemeral organism is one that has sporadic and random intervals of activity during the day or night in which food is acquired, socializing with other organisms occurs, and any other activities necessary for livelihood are performed...

, active throughout the day at short intervals.