Tilia americana is a species of Tilia

Tilia

Tilia is a genus of about 30 species of trees native throughout most of the temperate Northern Hemisphere. The greatest species diversity is found in Asia, and the genus also occurs in Europe and eastern North America, but not western North America...

native to eastern North America

North America

North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

, from southeast Manitoba

Manitoba

Manitoba is a Canadian prairie province with an area of . The province has over 110,000 lakes and has a largely continental climate because of its flat topography. Agriculture, mostly concentrated in the fertile southern and western parts of the province, is vital to the province's economy; other...

 east to New Brunswick

New Brunswick

New Brunswick is one of Canada's three Maritime provinces and is the only province in the federation that is constitutionally bilingual . The provincial capital is Fredericton and Saint John is the most populous city. Greater Moncton is the largest Census Metropolitan Area...

, southwest to northeast Texas

Texas

Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

, and southeast to South Carolina

South Carolina

South Carolina is a state in the Deep South of the United States that borders Georgia to the south, North Carolina to the north, and the Atlantic Ocean to the east. Originally part of the Province of Carolina, the Province of South Carolina was one of the 13 colonies that declared independence...

, and west along the Niobrara River

Niobrara River

The Niobrara River is a tributary of the Missouri River, approximately long, running through the U.S. states of Wyoming and Nebraska. The river drains one of the most arid sections of the Great Plains, and has a low flow for a river of its length...

 to Cherry County, Nebraska

Cherry County, Nebraska

Cherry County is unusual in being split between two time zones, Mountain Time and Central Time. The lines between the two runs roughly north to south along the eastern third of the county. Cherry County is also the location of the Valentine National Wildlife Refuge, the Fort Niobrara National...

. Common names include American Linden and Basswood (also applied to other species of Tilia in the timber

Timber

Timber may refer to:* Timber, a term common in the United Kingdom and Australia for wood materials * Timber, Oregon, an unincorporated community in the U.S...

 trade).

Description

Tilia americana is a medium-sized to large deciduous

Deciduous

Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 tree

Tree

A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 reaching a height of 60 to 120 ft (exceptionally 129 ft) with a trunk diameter of 3-4 ft at maturity. The crown is domed, the branches spreading, often pendulous. The bark

Bark

Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines and shrubs. Bark refers to all the tissues outside of the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner...

 is gray to light brown, with narrow, well defined fissures. The root

Root

In vascular plants, the root is the organ of a plant that typically lies below the surface of the soil. This is not always the case, however, since a root can also be aerial or aerating . Furthermore, a stem normally occurring below ground is not exceptional either...

s are large, deep, and spreading. The twig

Twig

A twig is a small thin terminal branch of a woody plant. Twigs are critically important in identification of trees, shrubs and vines, especially in wintertime. The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away...

s are smooth, reddish-green, becoming light gray in their second year, finally dark brown or brownish gray, marked with dark wart-like excrescences. The winter buds are stout, ovate-acute, smooth, deep red, with two bud scales visible. The leaves

Leaf

A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are simple, alternately arranged, ovate to cordate, inequalateral at the base (the side nearest the branch the largest), 10-15 cm (can grow up to 25 cm) long and broad, with a long, slender petiole, a coarsely serrated margin and an acuminate apex. They open from the bud conduplicate, pale green, downy; when full grown are dark green, smooth, shining above, paler beneath, with tufts of rusty brown hairs in the axils of the primary veins; the small stipule

Stipule

In botany, stipule is a term coined by Linnaeus which refers to outgrowths borne on either side of the base of a leafstalk...

s fall soon after leaf opening. The fall color is yellow-green to yellow. Both the twigs and leaves contain mucilaginous

Mucilage

Mucilage is a thick, gluey substance produced by most plants and some microorganisms. It is a polar glycoprotein and an exopolysaccharide.It occurs in various parts of nearly all classes of plant, usually in relatively small percentages, and is frequently associated with other substances, such as...

 sap. The flower

Flower

A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are small, fragrant, yellowish-white, 10–14 mm diameter, arranged in drooping, cymose

Inflorescence

An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Strictly, it is the part of the shoot of seed plants where flowers are formed and which is accordingly modified...

 clusters of 6–20 with a whitish-green leaf-like bract

Bract

In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis, or cone scale. Bracts are often different from foliage leaves. They may be smaller, larger, or of a different color, shape, or texture...

 attached for half its length at the base of the cyme; they are perfect, regular, with five sepals and petals, numerous stamens, and a five-celled superior ovary. Flowering is in early to mid summer; pollination is by bee

Bee

Bees are flying insects closely related to wasps and ants, and are known for their role in pollination and for producing honey and beeswax. Bees are a monophyletic lineage within the superfamily Apoidea, presently classified by the unranked taxon name Anthophila...

s. The fruit

Fruit

In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is a small, globose, downy, hard and dry cream-colored nutlet with a diameter of 8-10 mm.

Taxonomy

The circumscription of the species is disputed; some authors treat it in a narrow sense, with Tilia caroliniana, Tilia heterophylla

Tilia heterophylla

Tilia heterophylla , is a species of Tilia native to mesic forests in eastern North America from central New York south to northernmost Florida and west to Missouri; it is most common in the Appalachian Mountains....

, and Tilia mexicana regarded as separate species, while others treat these as varieties or synonyms of T. americana. The distribution and description above treat the species in its narrow sense; taken in the broad sense, including plants with the leaves white below with dense tomentum (velvety hairs), the distribution extends southwards to Florida

Florida

Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

, and in Mexico

Mexico

The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 west to Sinaloa

Sinaloa

Sinaloa officially Estado Libre y Soberano de Sinaloa is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 18 municipalities and its capital city is Culiacán Rosales....

 and south to Oaxaca

Oaxaca

Oaxaca , , officially Estado Libre y Soberano de Oaxaca is one of the 31 states which, along with the Federal District, comprise the 32 federative entities of Mexico. It is divided into 571 municipalities; of which 418 are governed by the system of customs and traditions...

.

Ecology

This species is dominant in the Acer saccharum - Tilia americana association, most common in western Wisconsin

Wisconsin

Wisconsin is a U.S. state located in the north-central United States and is part of the Midwest. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michigan to the northeast, and Lake Superior to the north. Wisconsin's capital is...

 and central Minnesota

Minnesota

Minnesota is a U.S. state located in the Midwestern United States. The twelfth largest state of the U.S., it is the twenty-first most populous, with 5.3 million residents. Minnesota was carved out of the eastern half of the Minnesota Territory and admitted to the Union as the thirty-second state...

, but occurs as far east as New England

New England

New England is a region in the northeastern corner of the United States consisting of the six states of Maine, New Hampshire, Vermont, Massachusetts, Rhode Island, and Connecticut...

 and southern Quebec

Quebec

Quebec or is a province in east-central Canada. It is the only Canadian province with a predominantly French-speaking population and the only one whose sole official language is French at the provincial level....

 where the soils are mesic

Mesic habitat

In ecology, a mesic habitat is a type of habitat with a moderate or well-balanced supply of moisture, e.g., a mesic forest, a temperate hardwood forest, or dry-mesic prairie. Compared to a dry habitat, a mesic habitat is moister....

 with relatively high pH

Soil pH

The soil pH is a measure of the acidity or basicity in soils. pH is defined as the negative logarithm of the activity of hydrogen ions in solution. It ranges from 0 to 14, with 7 being neutral. A pH below 7 is acidic and above 7 is basic. Soil pH is considered a master variable in soils as it...

. It also has minor occurrence in many other forest cover types.

Its flowers provide abundant nectar for insects. The seeds are eaten by chipmunk

Chipmunk

Chipmunks are small striped squirrels native to North America and Asia. They are usually classed either as a single genus with three subgenera, or as three genera.-Etymology and taxonomy:...

s, mice

MICE

-Fiction:*Mice , alien species in The Hitchhiker's Guide to the Galaxy*The Mice -Acronyms:* "Meetings, Incentives, Conferencing, Exhibitions", facilities terminology for events...

, and squirrel

Squirrel

Squirrels belong to a large family of small or medium-sized rodents called the Sciuridae. The family includes tree squirrels, ground squirrels, chipmunks, marmots , flying squirrels, and prairie dogs. Squirrels are indigenous to the Americas, Eurasia, and Africa and have been introduced to Australia...

s. Rabbit

Rabbit

Rabbits are small mammals in the family Leporidae of the order Lagomorpha, found in several parts of the world...

s and vole

Vole

A vole is a small rodent resembling a mouse but with a stouter body, a shorter hairy tail, a slightly rounder head, smaller ears and eyes, and differently formed molars . There are approximately 155 species of voles. They are sometimes known as meadow mice or field mice in North America...

s eat the bark, sometimes girdling young trees. The leaves serve as food for caterpillar

Caterpillar

Caterpillars are the larval form of members of the order Lepidoptera . They are mostly herbivorous in food habit, although some species are insectivorous. Caterpillars are voracious feeders and many of them are considered to be pests in agriculture...

s of various Lepidoptera

Lepidoptera

Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

 (see Lepidoptera which feed on Tilia). The ribbed cocoon maker

Bucculatricidae

Bucculatricidae or is a family of moths. This small family has representatives in all parts of the world. Some authors place the group as a subfamily of the family Lyonetiidae....

 species Bucculatrix improvisa has not been found on other plants.

This species is particularly susceptible to adult Japanese beetle

Japanese beetle

The beetle species Popillia japonica is commonly known as the Japanese beetle. It is about long and wide, with iridescent copper-colored elytra and green thorax and head...

s (an invasive species

Invasive species

"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....

 in North America) that feed on its leaves.

Cultivation and uses

Tilia americana can be propagated by cuttings and grafting as well as by seed. The plants grow rapidly in a rich soil, but are subject to the attacks of many insect enemies.

The species is recommended as an ornamental tree when the mass of foliage or a deep shade is desired; no native tree surpasses it in this respect. It is often planted on the windward side of an orchard as a protection to young and delicate trees. It is cultivated at least as far north as Juneau, Alaska

Juneau, Alaska

The City and Borough of Juneau is a unified municipality located on the Gastineau Channel in the panhandle of the U.S. state of Alaska. It has been the capital of Alaska since 1906, when the government of the then-District of Alaska was moved from Sitka as dictated by the U.S. Congress in 1900...

.

The foliage and flowers are both edible, though many prefer only to eat the tender young leaves. It is a beneficial species for attracting pollinators as well. Bees produce excellent honey with a mildly spicy flavor from its blossoms.

Cultivars include 'Nova', 'Duros' (with an upright crown), the pyramidal 'Frontyard' and the conic-crowned 'Redmond'.

Wood

The wood is pale brown, sometimes nearly white or faintly tinged with red; light, soft with fine close grain; clear of knots but does not split easily. It is sold generally under the name basswood, but is sometimes confounded with tulip-wood and then called white-wood, and is largely used in the manufacture of wooden-ware, wagon boxes and furniture. It has a density of 0.4525. This makes it valuable in the manufacture of wooden-ware, cheap furniture, bodies of carriages; it is also especially adapted for wood-carving. The inner bark is very tough and fibrous, used in the past for making ropes.
It is a common wood for use in the production of solid body electric guitars, where it is considered an analogue for aspen and poplar, because it is light, strong and resonant, though it is usually used for guitars that will be painted an opaque color, because its lack of notable grain makes it an unattractive candidate for transparent finish.

Medicinal Uses

Although Tilia cordata

Tilia cordata

Tilia cordata is a species of Tilia native to much of Europe and western Asia, north to southern Great Britain , central Scandinavia, east to central Russia, and south to central Spain, Italy, Bulgaria, Turkey, and the Caucasus; in the south of its range it is restricted to...

is believed to be stronger, T. americana is also used medicinally. The dried flowers are mildly sweet and sticky, and the fruit is somewhat sweet and mucilaginous. Linden tea has a pleasing taste, due to the aromatic volatile oil found in the flowers. The flowers, leaves, wood, and charcoal (obtained from the wood) are used for medicinal purposes. Active ingredients in the linden flowers include flavonoid

Flavonoid

Flavonoids , are a class of plant secondary metabolites....

s (which act as antioxidant

Antioxidant

An antioxidant is a molecule capable of inhibiting the oxidation of other molecules. Oxidation is a chemical reaction that transfers electrons or hydrogen from a substance to an oxidizing agent. Oxidation reactions can produce free radicals. In turn, these radicals can start chain reactions. When...

s), volatile oils, and mucilaginous constituents (which soothe and reduce inflammation). The plant also contains tannin

Tannin

A tannin is an astringent, bitter plant polyphenolic compound that binds to and precipitates proteins and various other organic compounds including amino acids and alkaloids.The term tannin refers to the use of...

s that can act as an astringent

Astringent

An astringent substance is a chemical compound that tends to shrink or constrict body tissues, usually locally after topical medicinal application. The word "astringent" derives from Latin adstringere, meaning "to bind fast"...

.

Linden flowers are used in colds, cough, fever, infections, inflammation, high blood pressure, headache (particularly migraine), as a diuretic (increases urine production), antispasmodic (reduces smooth muscle spasm along the digestive tract), and sedative. The flowers were added to baths to quell hysteria, and steeped as a tea to relieve anxiety-related indigestion, irregular heartbeat, and vomiting. The leaves are used to promote sweating to reduce fevers. The wood is used for liver and gallbladder disorders and cellulitis (inflammation of the skin and surrounding soft tissue). That wood burned to charcoal is ingested to treat intestinal disorders and used topically to treat edema or infection, such as cellulitis or ulcers of the lower leg.