Tree breeding
Encyclopedia
Tree breeding is the application of genetic principles to the genetic improvement and management of forest trees.
In contrast to the selective breeding of livestock, arable crops, and horticultural flowers over the last few centuries, the breeding of trees, with the exception of fruit trees, is a relatively recent occurrence.
A typical forest tree breeding program starts with selection of superior phenotypes (plus trees) in a natural or planted forest. This application of mass selection improves the mean performance of the forest. Offspring is obtained from selected trees and grown in test plantations that act as genetic trials. Based on such tests the best genotypes among the parents can be selected. Selected trees are typically multiplied by either seeds or grafting
and seed orchard
s are established when the preferred output is improved seed. Alternatively, the best genotypes can be directly propagated by cuttings or in-vitro methods and used directly in clonal plantations. The first system is frequently used in pines and other conifers, while the second is typical in some broadleaves (poplars, eucalypts and others). The objectives of a tree breeding program range from yield improvement and adaptation to particular conditions, to pest-resistance, wood characters etc. Currently, tree breeding is gaining new interest under the threat of Global Warming, and it is taking advantage of the fast development in plant genetics and genomics.
type, testing and selection methods, testing population size and time) and deployment methods of the genetically improved material (seed orchard
s and clonal forestry: genetic contribution, size). Computer simulators are frequently used: stochastic – based on defined and random algorithms; and deterministic – based on defined algorithms.
Selection strategies have been compared for annual progress in long-term breeding at a given annual cost considering genetic gain, gene diversity, cost components, and time components. For Norway spruce it seems favourable to clone full sib families and then select based on clonal performance while for Scots pine a two stage strategy seems best, first phenotypic pre-selection and then progeny-testing the selections.
In contrast to the selective breeding of livestock, arable crops, and horticultural flowers over the last few centuries, the breeding of trees, with the exception of fruit trees, is a relatively recent occurrence.
A typical forest tree breeding program starts with selection of superior phenotypes (plus trees) in a natural or planted forest. This application of mass selection improves the mean performance of the forest. Offspring is obtained from selected trees and grown in test plantations that act as genetic trials. Based on such tests the best genotypes among the parents can be selected. Selected trees are typically multiplied by either seeds or grafting
Grafting
Grafting is a horticultural technique whereby tissues from one plant are inserted into those of another so that the two sets of vascular tissues may join together. This vascular joining is called inosculation...
and seed orchard
Seed orchard
A seed orchard is an intensively-managed plantation of specifically arranged trees for the mass production of genetically improved seeds to create plants, or seeds for the establishment of new forests.-General:...
s are established when the preferred output is improved seed. Alternatively, the best genotypes can be directly propagated by cuttings or in-vitro methods and used directly in clonal plantations. The first system is frequently used in pines and other conifers, while the second is typical in some broadleaves (poplars, eucalypts and others). The objectives of a tree breeding program range from yield improvement and adaptation to particular conditions, to pest-resistance, wood characters etc. Currently, tree breeding is gaining new interest under the threat of Global Warming, and it is taking advantage of the fast development in plant genetics and genomics.
Optimisation
Tree breeders make efforts to get their operation efficient by optimising tree breeding. Scientists develop tools aimed at improvement of the efficiency of tree breeding programmes. Optimising can mean adapting strategies and methods to certain species, group of populations, structure of genetic variation and mode of inheritance of the important traits to obtain the highest benefit per unit of time. Optimising is usually carried out at the following levels: breeding strategy (appropriate intensity of breeding, breeding population structure and size, plan for maintenance of genetic diversity), breeding methods (matingMating
In biology, mating is the pairing of opposite-sex or hermaphroditic organisms for copulation. In social animals, it also includes the raising of their offspring. Copulation is the union of the sex organs of two sexually reproducing animals for insemination and subsequent internal fertilization...
type, testing and selection methods, testing population size and time) and deployment methods of the genetically improved material (seed orchard
Seed orchard
A seed orchard is an intensively-managed plantation of specifically arranged trees for the mass production of genetically improved seeds to create plants, or seeds for the establishment of new forests.-General:...
s and clonal forestry: genetic contribution, size). Computer simulators are frequently used: stochastic – based on defined and random algorithms; and deterministic – based on defined algorithms.
Selection strategies have been compared for annual progress in long-term breeding at a given annual cost considering genetic gain, gene diversity, cost components, and time components. For Norway spruce it seems favourable to clone full sib families and then select based on clonal performance while for Scots pine a two stage strategy seems best, first phenotypic pre-selection and then progeny-testing the selections.
Selected bibliography
- White, T.L., Adams, W.T. and Neale, D.B. 2007. Forest Genetics, CABI. ISBN 978-0851993485
- 2007 Gösta Eriksson, Inger Ekberg and David Clapham. An Introduction to Forest Genetics. ISBN 91-576-7190-7 http://www2.vbsg.slu.se/forgen/
- 1984. Applied Forest Tree Improvement. Bruce Zobel, John Talbert, North Carolina State University, John Wiley and Sons, Inc, ISBN 0-471-09682-2