Typhlobelus
Encyclopedia
Typhlobelus is a genus
of catfish
es (order
Siluriformes) of the family
Trichomycteridae
. It includes four species
, T. guacamaya, T. lundbergi, T. macromycterus, and T. ternetzi.
River basin of Venezuela
. T. lundbergi inhabits the lower Orinoco River, between Ciudad Bolivar
and Los Castillos, Venezuela. T. macromycterus is known from the Tocantins River
near Tucuruí
, Pará State, Brazil
. T. ternetzi lives in the upper Rio Negro basin, Brazil.
, the loss or extreme reduction of pectoral fins, a reduced lateral line
, and the reduction or complete loss of eyes in some species. These fish are markedly miniaturized, yet retain a relatively well-ossified skeleton comparable in both bone differentiation and degree of calcification to that observed in larger trichomycterids.
T. guacamaya is distinguished from all congeners by the presence of three branchiostegal rays
(vs. four in T. ternetzi and T. lundbergi, five in T. macromycterus), posterior naris
absent (vs. present, nares bilaterally paired), and the lack of pleural ribs (vs. one pair of pleural ribs associated with the first free vertebra). T. lundbergi is distinguished from all congeners by the presence of four laterosensory pores
on the head (vs. three). Both T. guacamaya and T. lundbergi are distinguished from ternetzi and T. macromycterus by the absence of eyes (vs. eyes present and vestigial); between T. ternetzi and T. lundbergi, the distinction may be between the presence or absence of eyes or eyespots. T. macromycterus has one or two odontode
s on the opercle
and five branchiostegal rays, while T. ternetzi and T. lundbergi have no odontodes and four branchiostegal rays.
These fish have greatly elongate, slender bodies. These fish have three pairs of barbels
, all similar in length and general appearance. The mouth is ventral. Typhlobelus has a long duck-billed rostrum
that protrudes anteriorly well beyond the bases of the maxillary barbels.
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of catfish
Catfish
Catfishes are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the heaviest and longest, the Mekong giant catfish from Southeast Asia and the second longest, the wels catfish of Eurasia, to detritivores...
es (order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...
Siluriformes) of the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...
Trichomycteridae
Trichomycteridae
Trichomycteridae is a family of catfishes commonly known as the pencil or parasitic catfishes. This family includes the infamous candiru fish, feared by some people for its alleged habit of entering into the urethra of humans....
. It includes four species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...
, T. guacamaya, T. lundbergi, T. macromycterus, and T. ternetzi.
Distribution
T. guacamaya originates from the Cuao River in the OrinocoOrinoco
The Orinoco is one of the longest rivers in South America at . Its drainage basin, sometimes called the Orinoquia, covers , with 76.3% of it in Venezuela and the remainder in Colombia...
River basin of Venezuela
Venezuela
Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...
. T. lundbergi inhabits the lower Orinoco River, between Ciudad Bolivar
Ciudad Bolívar
Ciudad Bolívar is the capital of Venezuela's southeastern Bolivar State. It was founded with the name Angostura in 1764, renamed in 1846, and, as of 2010, had an estimated population of 350,691....
and Los Castillos, Venezuela. T. macromycterus is known from the Tocantins River
Tocantins River
The Tocantins is a river in Brazil, the central fluvial artery of the country. In the Tupi language, its name means "toucan's beak" . It runs from south to north for about 2,640 km. It is not really a branch of the Amazon River, although usually so considered, since its waters flow into the...
near Tucuruí
Tucuruí
Tucuruí is a town and municipality in the state of Pará in the Northern region of Brazil.The city is served by Tucuruí Airport.-References:...
, Pará State, Brazil
Brazil
Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...
. T. ternetzi lives in the upper Rio Negro basin, Brazil.
Description
Species of Typhlobelus share the extreme reduction of pigmentation, loss of the dorsal finDorsal fin
A dorsal fin is a fin located on the backs of various unrelated marine and freshwater vertebrates, including most fishes, marine mammals , and the ichthyosaurs...
, the loss or extreme reduction of pectoral fins, a reduced lateral line
Lateral line
The lateral line is a sense organ in aquatic organisms , used to detect movement and vibration in the surrounding water. Lateral lines are usually visible as faint lines running lengthwise down each side, from the vicinity of the gill covers to the base of the tail...
, and the reduction or complete loss of eyes in some species. These fish are markedly miniaturized, yet retain a relatively well-ossified skeleton comparable in both bone differentiation and degree of calcification to that observed in larger trichomycterids.
T. guacamaya is distinguished from all congeners by the presence of three branchiostegal rays
Ichthyology terms
Ichthyology uses several terms that are unique to the science.Contents: A B C D E F G H I J K L M N O P Q R S T U V W X Y Z- A :* Abdomen: belly* Abdominal: pertaining to the belly* Actinosts: a series of bones at the base of the pectoral rays....
(vs. four in T. ternetzi and T. lundbergi, five in T. macromycterus), posterior naris
Nostril
A nostril is one of the two channels of the nose, from the point where they bifurcate to the external opening. In birds and mammals, they contain branched bones or cartilages called turbinates, whose function is to warm air on inhalation and remove moisture on exhalation...
absent (vs. present, nares bilaterally paired), and the lack of pleural ribs (vs. one pair of pleural ribs associated with the first free vertebra). T. lundbergi is distinguished from all congeners by the presence of four laterosensory pores
Lateral line
The lateral line is a sense organ in aquatic organisms , used to detect movement and vibration in the surrounding water. Lateral lines are usually visible as faint lines running lengthwise down each side, from the vicinity of the gill covers to the base of the tail...
on the head (vs. three). Both T. guacamaya and T. lundbergi are distinguished from ternetzi and T. macromycterus by the absence of eyes (vs. eyes present and vestigial); between T. ternetzi and T. lundbergi, the distinction may be between the presence or absence of eyes or eyespots. T. macromycterus has one or two odontode
Odontode
Odontodes, or dermal teeth, are hard structures found on the external surfaces of animals or near internal openings. They comprise a soft pulp surrounded by dentine and covered by a mineralized substance such as enamel, a structure similar to that of teeth. They generally do not have the same...
s on the opercle
Ichthyology terms
Ichthyology uses several terms that are unique to the science.Contents: A B C D E F G H I J K L M N O P Q R S T U V W X Y Z- A :* Abdomen: belly* Abdominal: pertaining to the belly* Actinosts: a series of bones at the base of the pectoral rays....
and five branchiostegal rays, while T. ternetzi and T. lundbergi have no odontodes and four branchiostegal rays.
These fish have greatly elongate, slender bodies. These fish have three pairs of barbels
Barbel (anatomy)
A barbel on a fish is a slender, whiskerlike tactile organ near the mouth. Fish that have barbels include the catfish, the carp, the goatfish, sturgeon, the zebrafish and some species of shark...
, all similar in length and general appearance. The mouth is ventral. Typhlobelus has a long duck-billed rostrum
Rostrum (anatomy)
The term rostrum is used for a number of unrelated structures in different groups of animals:*In crustaceans, the rostrum is the forward extension of the carapace in front of the eyes....
that protrudes anteriorly well beyond the bases of the maxillary barbels.