Urocoptidae is a family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of land snail

Land snail

A land snail is any of the many species of snail that live on land, as opposed to those that live in salt water and fresh water. Land snails are terrestrial gastropod mollusks that have shells, It is not always an easy matter to say which species are terrestrial, because some are more or less...

s, gastropods in the superfamily Urocoptoidea

Urocoptoidea

Urocoptoidea is a superfamily of land snails, gastropods in the informal group Sigmurethra. cfThe superfamily Urocoptoidea has been established by Uit de Weerd based on molecular phylogeny research in 2008.- Families :...

.

2005 taxonomy

The Urocoptidae family has been classified in the superfamily Orthalicoidea

Orthalicoidea

The Orthalicoidea are a superfamily of air-breathing land snails, terrestrial gastropod mollusks in the informal group Sigmurethra.- 2005 taxonomy :...

 within the informal group Sigmurethra

Sigmurethra

Sigmurethra is a taxonomic category of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This is an Informal Group which includes the majority of land snails and slugs....

, itself belonging to the clade

Clade

A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

 Stylommatophora

Stylommatophora

Stylommatophora is a taxon of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This taxon is currently considered to be a clade. It was previously regarded as an infraorder...

 within the clade Eupulmonata

Eupulmonata

The Eupulmonata is a taxonomic clade of air-breathing snails. The great majority of this group are land snails and slugs, but some are marine and some are saltmarsh snails that can tolerate salty conditions.-Linnean taxonomy:...

 (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005

Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)

The taxonomy of the Gastropoda as it was revised by Philippe Bouchet and Jean-Pierre Rocroi is currently the most up-to-date overall system for classifying gastropod mollusks...

).

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

• Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868
• Apomatinae Paul, 1982
• Brachypodellinae H. B. Baker, 1956
• Eucalodiinae P. Fischer & Crosse, 1873
• Holospirinae Pilsbry, 1946
• Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972
• Tetrentodontinae Bartsch, 1943

2008 taxonomy

Uit de Weerd (2008) has moved the Urocoptidae to newly established superfamily Urocoptoidea

Urocoptoidea

Urocoptoidea is a superfamily of land snails, gastropods in the informal group Sigmurethra. cfThe superfamily Urocoptoidea has been established by Uit de Weerd based on molecular phylogeny research in 2008.- Families :...

 based on molecular phylogeny

Molecular phylogeny

Molecular phylogenetics is the analysis of hereditary molecular differences, mainly in DNA sequences, to gain information on an organism's evolutionary relationships. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree...

 research.

A cladogram

Cladogram

A cladogram is a diagram used in cladistics which shows ancestral relations between organisms, to represent the evolutionary tree of life. Although traditionally such cladograms were generated largely on the basis of morphological characters, DNA and RNA sequencing data and computational...

 based on Bayesian analysis of sequences of 28S ribosomal RNA

28S ribosomal RNA

28S ribosomal RNA is the eukaryotic nuclear homologue of the prokaryotic 23S ribosomal RNA; this is the structural RNA for the large component of eukaryotic cytoplasmic ribosomes, and thus one of the basic components of all eukaryotic cells....

 showing phylogenic relations of Urocoptoidea. There are all up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color

Khaki (color)

The name of the color khaki coined in British India comes from the Hindustani language , meaning "dusty, dust covered or earth colored." It has been used by many armies around the world for uniforms, including camouflage...

 in the cladogram.

Genera

Genera in the family Urocoptidae include:

Urocoptinae

• Autocoptis Pilsbry, 1902
• Cylindrella L. Pfeiffer, 1840
• Urocoptis
  Urocoptis
  Urocoptis is a genus of air-breathing land snails, terrestrial pulmonate gastropod molluscs in the family Urocoptidae.Urocoptis is the type genus of the family Urocoptidae.- Further reading :...
  
  Beck, 1837 - type genus of the family Urocoptidae

Apominae

• Apoma Beck, 1837 - type genus of the subfamily Apomatinae
• Simplicervix Pilsbry, 1903

Brachypodellinae

• Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae

Eucalodiinae

• Anisospira Strebel, 1880
• Archegocoptis Pilsbry, 1903
• Dissotropis Bartsch, 1906
• Eucalodium Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae

Holospirinae

• Coelostemma Dall, 1895
• Epirobia Strebel & Pfeffer, 1880
• Hendersoniella Dall, 1905
• Holospira
  Holospira
  Holospira is a genus of air-breathing, land snails, terrestrial pulmonate gastropod molluscs in the family Urocoptidae.Holospira is the type genus of the subfamily 'Holospirinae Pilsbry, 1946'.- Etymology :...
  
  Martens, 1860 - type genus of the subfamily Holospirinae

Microceraminae

• Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae
• Macroceramus Guilding, 1828

Tetrentodontinae

• Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae
• Torrecoptis Bartsch, 1943

other genera include:

• Allocoptis Thompson & Franz, 1976
• Amphicosmia Pilsbry & Vanatta, 1898
• Anoma
• Arangia
• Bactrocoptis
• Badiofaux
• Bostrichocentrum
• Bostrychocentrum
• Callonia
• Centralia
• Cochlodinella
• Coelocentrum
• Geoscala
• Gongylostoma
• Gyraxis
• Haplocion
• Heterocoptis
• Idiostemma
• Liocallonia
• Metastoma
• Mychostoma
• Nesocoptis
• Nodulia
• Oligostylus
• Organocoptis
• Pfeiffericoptis
• Pineria
• Pleurostemma
• Poeycoptis
• Propilsbrya
• Pseudopineria
• Pycnoptychia
  Pycnoptychia
  Pycnoptychia is a genus of a land snails, terrestrial gastropod mollusks in the family Urocoptidae.- Species :Species in the genus Pycnoptychia include:* Pycnoptychia amicorum Torre & Bartsch, 2008...
  
  Pilsbry & Vanatta, 1898
• Sagracoptis
• Spiroceramus
• Spirostemma
• Strophina
• Tomelasmus
• Trilamellaxis
• Uncinicoptis

Further reading

• Bartsch P.
  Paul Bartsch
  Paul Bartsch was an American malacologist and carcinologist.Paul Bartsch emigrated to the U.S.A in 1880. He graduated from the University of Iowa with a B.S. in 1896, and M.S. in 1899, and PhD in 1905...
  
   1943. Notes on Mexican Urocoptid mollusks. Journal of the Washington Academy of Sciences 33(2): 54-59; 7 figs.
• Carlos de la Torre. 1912. New Cuban Urocoptis of the U. cinerea group. The Nautilus, volume 26, number 4, pages 55-58, plate 6.
• Pilsbry H. A. & Henderson J. B. jr. (February) 1913. Two new Cuban Urocoptidae. The Nautilus, volume 26, number 10, pages 109-110. Miguel L. Jaume & Alfredo de la Torre. 1976. Los Urocoptidae de Cuba (Mollusca-Pulmonata). Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp.
• Richardson C. L. 1991. Urocoptidae: catalogue of species. Tryonia 22: 1-245.
• Tryon G. W.
  George Washington Tryon
  George Washington Tryon, Jr. was an American malacologist who worked at the Academy of Natural Sciences in Philadelphia.- Biography :George Washington Tryon was the son of Edward K. Tryon and Adeline Savidt...
  
   (1903) Manual of Conchology. Second series: Pulmonata. Volume 15. Urocoptidae. Continued by H. A. Pilsbry.

External links

• photos
• photos
• blog Bram's Snail Site with photos