Arvicolinae
Encyclopedia
The Arvicolinae are a subfamily of rodent
Rodent
Rodentia is an order of mammals also known as rodents, characterised by two continuously growing incisors in the upper and lower jaws which must be kept short by gnawing....

s that includes the vole
Vole
A vole is a small rodent resembling a mouse but with a stouter body, a shorter hairy tail, a slightly rounder head, smaller ears and eyes, and differently formed molars . There are approximately 155 species of voles. They are sometimes known as meadow mice or field mice in North America...

s, lemming
Lemming
Lemmings are small rodents, usually found in or near the Arctic, in tundra biomes. They are subniveal animals, and together with voles and muskrats, they make up the subfamily Arvicolinae , which forms part of the largest mammal radiation by far, the superfamily Muroidea, which also includes rats,...

s, and muskrat
Muskrat
The muskrat , the only species in genus Ondatra, is a medium-sized semi-aquatic rodent native to North America, and introduced in parts of Europe, Asia, and South America. The muskrat is found in wetlands and is a very successful animal over a wide range of climates and habitats...

s. They are most closely related to the other subfamilies in the Cricetidae
Cricetidae
The Cricetidae are a family of rodents in the large and complex superfamily Muroidea. It includes true hamsters, voles, lemmings, and New World rats and mice...

 (comprising the hamster
Hamster
Hamsters are rodents belonging to the subfamily Cricetinae. The subfamily contains about 25 species, classified in six or seven genera....

s and New World rats and mice
New World rats and mice
The New World rats and mice are a group of related rodents found in North and South America. They are extremely diverse in appearance and ecology, ranging in from the tiny Baiomys to the large Kunsia...

). Some authorities place the subfamily Arvicolinae in the family Muridae
Muridae
Muridae is the largest family of mammals. It contains over 600 species found naturally throughout Eurasia, Africa, and Australia. They have been introduced worldwide. The group includes true mice and rats, gerbils, and relatives....

 along with all other members of the superfamily Muroidea
Muroidea
Muroidea is a large superfamily of rodents. It includes hamsters, gerbils, true mice and rats, and many other relatives. They occupy a vast variety of habitats on every continent except Antarctica. Some authorities have placed all members of this group into a single family, Muridae, due to...

. Some refer to the subfamily as the Microtinae or rank the taxon as a full family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

, the Arvicolidae .

The Arvicolinae are the most populous group of Rodentia in the Northern Hemisphere. They often are found in fossil occlusions of bones cached by past predators such as owls and other birds of prey.

Fossils of this group are often used for biostratigraphic
Biostratigraphy
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. Usually the aim is correlation, demonstrating that a particular horizon in one geological section represents the same period...

 dating of archeological sites in North America and Europe.

Description

The most convenient distinguishing feature of the Arvicolinae is the nature of their molar
Molar (tooth)
Molars are the rearmost and most complicated kind of tooth in most mammals. In many mammals they grind food; hence the Latin name mola, "millstone"....

 teeth, which have prismatic cusps in the shape of alternating triangles. These molars are an adaptation to a herbivorous
Herbivore
Herbivores are organisms that are anatomically and physiologically adapted to eat plant-based foods. Herbivory is a form of consumption in which an organism principally eats autotrophs such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in...

 diet in which the major food plants include a large proportion of abrasive materials such as phytolith
Phytolith
Some plants can take up silica in the soil, whereupon it is deposited within different intracellular and extracellular structures of the plant. After these plants decay, silica is redeposited in the soil in the form of phytoliths , which are rigid, microscopic structures of varying sizes and shapes...

s; the teeth get worn down by abrasion throughout the adult life of the animal and they grow continuously in compensation.

Arvicolinae are Holarctic
Holarctic
The Holarctic ecozone refers to the habitats found throughout the northern continents of the world as a whole. This region is divided into the Palearctic, consisting of Northern Africa and all of Eurasia, with the exception of Southeast Asia and the Indian subcontinent, and the Nearctic,...

 in distribution and represent one of only a few major muroid
Muroidea
Muroidea is a large superfamily of rodents. It includes hamsters, gerbils, true mice and rats, and many other relatives. They occupy a vast variety of habitats on every continent except Antarctica. Some authorities have placed all members of this group into a single family, Muridae, due to...

 radiations to reach the New World
New World
The New World is one of the names used for the Western Hemisphere, specifically America and sometimes Oceania . The term originated in the late 15th century, when America had been recently discovered by European explorers, expanding the geographical horizon of the people of the European middle...

 via Beringia. (The others are the three subfamilies of New World rats and mice
New World rats and mice
The New World rats and mice are a group of related rodents found in North and South America. They are extremely diverse in appearance and ecology, ranging in from the tiny Baiomys to the large Kunsia...

.) Arvicolines do very well in the subnival zone beneath the winter snowpack and persist throughout winter without needing to hibernate
Hibernation
Hibernation is a state of inactivity and metabolic depression in animals, characterized by lower body temperature, slower breathing, and lower metabolic rate. Hibernating animals conserve food, especially during winter when food supplies are limited, tapping energy reserves, body fat, at a slow rate...

. They are also characterized by extreme fluctuations in population size.

Most arvicolines are small, furry, short tailed vole
Vole
A vole is a small rodent resembling a mouse but with a stouter body, a shorter hairy tail, a slightly rounder head, smaller ears and eyes, and differently formed molars . There are approximately 155 species of voles. They are sometimes known as meadow mice or field mice in North America...

s or lemming
Lemming
Lemmings are small rodents, usually found in or near the Arctic, in tundra biomes. They are subniveal animals, and together with voles and muskrats, they make up the subfamily Arvicolinae , which forms part of the largest mammal radiation by far, the superfamily Muroidea, which also includes rats,...

s, but some such as Ellobius
Ellobius
Ellobius is a genus of rodent in the family Cricetidae.It contains the following species:* Alai Mole Vole * Southern Mole Vole * Transcaucasian Mole Vole...

 and Hyperacrius
Hyperacrius
Hyperacrius is a genus of rodent in the family Cricetidae.It contains the following species:* True's Vole * Murree Vole -References:...

 are well adapted to a fossorial
Fossorial
A fossorial organism is one that is adapted to digging and life underground such as the badger, the naked mole rat, and the mole salamanders Ambystomatidae...

 lifestyle. Others, such as Ondatra, Neofiber, and Arvicola
Arvicola
The water voles are large voles in the genus Arvicola. They are found in both aquatic and dry habitat through Europe and much of northern Asia...

 have evolved a larger body size and associated with an aquatic
Aquatic animal
An aquatic animal is an animal, either vertebrate or invertebrate, which lives in water for most or all of its life. It may breathe air or extract its oxygen from that dissolved in water through specialised organs called gills, or directly through its skin. Natural environments and the animals that...

 lifestyle.

Phylogeny

The phylogeny of Arvicolinae has been studied using morphological and molecular characters. Markers for the molecular phylogeny of arvicolines included the mitochondrial DNA
Mitochondrial DNA
Mitochondrial DNA is the DNA located in organelles called mitochondria, structures within eukaryotic cells that convert the chemical energy from food into a form that cells can use, adenosine triphosphate...

 cytochrome b (cyb) gene
and the exon 10 of the growth hormone receptor
Growth hormone receptor
Growth hormone receptor is a protein that in humans is encoded by the GHR gene. GHR orthologs have been identified in most mammals.-Interactions:...

 (ghr) nuclear gene . The comparison of the cyb and ghr phylogenetic results seems to indicate that nuclear genes are useful for resolving relationships of recently evolved animals. As compared to mitochondrial genes, nuclear gene
Nuclear gene
A Nuclear gene is a gene located in the cell nucleus of a eukaryote. The term is used to distinguish nuclear genes from the genes in the mitochondrion, and in case of plants, also the chloroplast, which host their own genetic system and can produce proteins from scratch...

s display several informative sites in third codon positions that evolve rapidly enough to accumulate synapomorphies, but slow enough to avoid evolutionary noise.
Of note, mitochondrial pseudogenes translocated within the nuclear genome complicate the assessment of the mitochondrial DNA orthology, but they can also be used as phylogenetic markers .
Sequencing complete mitochondrial genomes of voles may help to distinguish between authentical genes and pseudogenes.

The complementary phylogenetic analysis of morphological and molecular characters
suggests that :
  • Ellobius, Prometheomys and Lagurus are among the most basal arvicolines ;
  • Dicrostonyx, Phenacomys and Arborimus may form a clade ;
  • Core arvicolines include three subclades:
    • Lemmini: Synaptomys, Lemmus, Myopus ;
    • Clethrionomyini: Eothenomys, Myodes ;
    • Arvicolini: Arvicola, Chionomys, Stenocranius and Microtus.
  • Microtus
    Microtus
    The genus Microtus is a group of voles found in North America, Europe and northern Asia. The genus name refers to the small ears of these animals. There are 62 species in the genus....

     sensu lato contains Alexandromys, ‘Neodon’, Mynomes, Lasiopodomys, Terricola, and Microtus sensu stricto ;
  • Ondatra and Dinaromys positions are uncertain, probably compromised by the convergent evolution
    Convergent evolution
    Convergent evolution describes the acquisition of the same biological trait in unrelated lineages.The wing is a classic example of convergent evolution in action. Although their last common ancestor did not have wings, both birds and bats do, and are capable of powered flight. The wings are...

     of morphological characters.


Some authorities have placed the zokor
Zokor
Zokors are Asiatic burrowing rodents resembling mole rats. They include two genera, Myospalax and Eospalax. Zokors are native to much of China, Kazhakstan, and Siberian Russia....

s within the Arvicolinae, but they have been shown to be unrelated.

Classification

Subfamily Arvicolinae - vole
Vole
A vole is a small rodent resembling a mouse but with a stouter body, a shorter hairy tail, a slightly rounder head, smaller ears and eyes, and differently formed molars . There are approximately 155 species of voles. They are sometimes known as meadow mice or field mice in North America...

s, lemming
Lemming
Lemmings are small rodents, usually found in or near the Arctic, in tundra biomes. They are subniveal animals, and together with voles and muskrats, they make up the subfamily Arvicolinae , which forms part of the largest mammal radiation by far, the superfamily Muroidea, which also includes rats,...

s, muskrat
Muskrat
The muskrat , the only species in genus Ondatra, is a medium-sized semi-aquatic rodent native to North America, and introduced in parts of Europe, Asia, and South America. The muskrat is found in wetlands and is a very successful animal over a wide range of climates and habitats...

s

The subfamily Arvicolinae contains ten tribes, seven of which are classified as voles, one as lemmings, and two as muskrats.
  • Tribe Arvicolini
    Arvicolini
    Arvicolini is a tribe of voles in the subfamily Arvicolinae.-List of species:Tribe Arvicolini*Genus Arvicola - water voles**European Water Vole, Arvicola amphibius ...

    • Genus Arvicola
      Arvicola
      The water voles are large voles in the genus Arvicola. They are found in both aquatic and dry habitat through Europe and much of northern Asia...

       - water voles
      • European (or Northern) Water Vole, Arvicola amphibius (Arvicola terrestris)
      • Southwestern (or Southern) Water Vole
        Southwestern Water Vole
        The Southwestern Water Vole or Southern Water Vole, Arvicola sapidus is a large amphibious vole found in most of France and south-westwards through Spain and Portugal. Although historically considered to be a member of the same species as the Eurasian Water Vole, Musser and Carleton considered it...

        , Arvicola sapidus
      • Montane Water Vole
        Montane Water Vole
        The Montane Water Vole or Arvicola scherman is a species of vole found throughout Europe, from the mountains of northern Spain through central Europe and eastwards through to central Romania. Initially regarded as a species, it was reassigned as a subspecies of terrestris before being again...

        , Arvicola scherman
    • Genus Blanfordimys
      Blanfordimys
      Blanfordimys is a genus of rodent in the family Cricetidae.It contains the following species:* Afghan Vole * Bucharian Vole -References:...

      • Afghan Vole
        Afghan Vole
        The Afghan Vole is a species of rodent in the family Cricetidae. It is found in south-central Asia.-Characteristics:...

        , Blanfordimys afghanus
      • Bucharian Vole
        Bucharian Vole
        The Bucharian Vole is a species of rodent in the family Cricetidae. It is found in possibly Afghanistan Tajikistan and Uzbekistan. Its natural habitat is temperate desert.-References:...

        , Blanfordimys bucharicus
    • Genus Chionomys
      Chionomys
      Chionomys is a genus of rodent in the family Cricetidae.It contains the following species:* Caucasian Snow Vole * European Snow Vole * Robert's Snow Vole -References:...

       - snow voles
      • Caucasian Snow Vole
        Caucasian Snow Vole
        The Caucasian Snow Vole is a species of rodent in the family Cricetidae.It is found in Georgia, Russian Federation, and Turkey.Its natural habitats are temperate forests and temperate grassland.-References:...

        , Chionomys gud
      • European Snow Vole
        European Snow Vole
        The European Snow Vole or Snow Vole is a species of rodent in the family Cricetidae.-Description:...

        , Chionomys nivalis
      • Robert's Snow Vole
        Robert's Snow Vole
        Robert's Snow Vole is a species of rodent in the family Cricetidae.It is found in Georgia, Russian Federation, and Turkey.Its natural habitats are temperate forests and temperate grassland.-References:...

        , Chionomys roberti
    • Genus Lasiopodomys
      Lasiopodomys
      Lasiopodomys is a genus of rodent in the family Cricetidae.It contains the following species:* Brandt's Vole * Plateau Vole * Mandarin Vole -References:...

      • Brandt's Vole
        Brandt's Vole
        Brandt's vole, , also known as the steppe vole, is a species of rodent in the family Cricetidae.It is found in China, Mongolia and Russia.-Morphology:...

        , Lasiopodomys brandtii
      • Plateau Vole
        Plateau Vole
        The Plateau Vole is a species of rodent in the family Cricetidae.It is found only in China.Its natural habitat is temperate grassland.-References:...

        , Lasiopodomys fuscus
      • Mandarin Vole
        Mandarin Vole
        The Mandarin Vole is a species of vole found in central China as well as the southern and central Korean Peninsula. An adult Mandarin Vole is typically of about 115 millimeters in length, in addition to about 28 mm of tail...

        , Lasiopodomys mandarinus
    • Genus Lemmiscus
      • Sagebrush Vole
        Sagebrush Vole
        The Sagebrush Vole is a tiny vole found in western North America. This is the only member of genus Lemmiscus....

        , Lemmiscus curtatus
    • Genus Microtus
      Microtus
      The genus Microtus is a group of voles found in North America, Europe and northern Asia. The genus name refers to the small ears of these animals. There are 62 species in the genus....

       - voles
      • Insular Vole
        Insular Vole
        The Insular Vole or St. Matthew Island Vole is a species of rodent in the family Cricetidae. It occurs only on St. Matthew Island and the adjacent Hall Island, in Alaska. On these Bering Sea islands, Insular Voles live in damp lowland areas, on the lower slopes of mountains, and on rye...

        , Microtus abbreviatus
      • California Vole
        California Vole
        The California vole is a type of vole which lives throughout much of California and part of southwestern Oregon. It is also known as the California meadow mouse. It averages in length although this length varies greatly between subspecies.-Description:The California vole is a medium sized vole,...

        , Microtus californicus
      • Rock Vole
        Rock Vole
        The Rock Vole is a medium-sized vole found in eastern North America. It is also called the Yellow-nosed Vole....

        , Microtus chrotorrhinus
      • Long-tailed Vole
        Long-tailed Vole
        The Long-tailed Vole is a small vole found in western North America. The Coronation Island Vole, once considered to be a separate species, is now believed to be a subspecies....

        , Microtus longicaudus
      • Mexican Vole
        Mexican Vole
        The Mexican Vole is a species of rodent in the family Cricetidae. It is found in Mexico and the United States.-References:...

        , Microtus mexicanus
      • Singing Vole
        Singing Vole
        The singing vole, Microtus miurus, is a medium-sized vole found in northwestern North America, including Alaska and northwestern Canada.-Physical characteristics:...

        , Microtus miurus
      • Water Vole, Microtus richardsoni
      • Zempoaltépec Vole
        Zempoaltépec Vole
        The Zempoaltépec Vole is a species of rodent in the family Cricetidae.It is found only in Mexico.-References:...

        , Microtus umbrosus
      • Taiga Vole
        Taiga Vole
        The Taiga Vole, Microtus xanthognathus, is a large vole found in northwestern North America, including Alaska and northwestern Canada. It is also sometimes called the Yellow-cheeked Vole or Chestnut-cheeked Vole....

        , Microtus xanthognathus
      • Subgenus Microtus
        • Field Vole
          Field Vole
          The field vole or short-tailed vole, Microtus agrestis, is a grey-brown vole, around 110mm in length, with only a short tail. They are one of the most common mammals in Europe, ranging from the Atlantic coast to Lake Baikal. They are found in moist grassy habitats, such as woodland, marsh, or...

          , Microtus agrestis
        • Anatolian Vole
          Anatolian Vole
          The Anatolian Vole or Microtus anatolicus, is a species of vole found in Turkey, specifically Konya Province.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. Pp. 894-1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder...

          , Microtus anatolicus
        • Common Vole
          Common Vole
          The Common Vole, Microtus arvalis, is a European mammal.- Distribution and habitat :The common vole is hardly restricted in means of distribution and habitat and inhabits large areas of Eurasia. As Microtus arvalis followed human civilisation, primary and secondary habitats can be distinguished...

          , Microtus arvalis
        • Cabrera's Vole
          Cabrera's Vole
          Cabrera's Vole is a species of rodent in the family Cricetidae.It is found in Portugal, and Spain.-References:* Amori, G. 1996. . Downloaded on 19 July 2007....

          , Microtus cabrerae
        • Doğramaci's Vole
          Dogramaci's Vole
          Doğramaci's Vole is a species of rodent in the family Cricetidae. It is found only in central Turkey, and is similar to the Social Vole although different in terms of cranial proportions and karyotype.-References:...

          , Microtus dogramacii
        • Günther's Vole
          Günther's Vole
          Günther's Vole is a species of rodent in the family Cricetidae.It is found in Bulgaria, Greece, Iran, Iraq, Israel, Lebanon, Macedonia, Serbia and Montenegro, Syria, Turkey, and Libya. In Libya, its natural habitats are temperate grassland, subtropical or tropical high-altitude grassland, and...

          , Microtus guentheri
        • Tien Shan Vole
          Tien Shan Vole
          The Tien Shan Vole is a species of rodent in the family Cricetidae.It is found in Afghanistan, Kazakhstan, Kyrgyzstan, Tajikistan, and Turkmenistan.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

          , Microtus ilaeus
        • Persian Vole
          Persian Vole
          The Persian Vole is a species of rodent in the family Cricetidae.It is only found in Iran.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

          , Microtus irani
        • Southern Vole
          Southern Vole
          The Southern Vole is a species of rodent in the family Cricetidae.It is found in Albania, Bulgaria, Finland, Greece, Iran, Svalbard , the Republic of Macedonia, Romania, Russia, Serbia and Montenegro, Slovakia, Turkey, Ukraine and Norway.-References:*Musser, G. G. and M. D. Carleton. 2005....

          , Microtus levis
        • Paradox Vole
          Paradox Vole
          The Paradox Vole is a species of rodent in the family Cricetidae found in southern Turkmenistan.-References:* D.E. Wilson & D.M. Reeder, 2005: Mammal Species of the World: A Taxonomic and Geographic Reference. Third Edition. The Johns Hopkins University Press, Baltimore....

          , Microtus paradoxus
        • Qazvin Vole
          Qazvin Vole
          The Qazvin Vole is a species of rodent in the family Cricetidae. It is found in Iran and is normally given as either part of the Microtus or Sumeriomys subgenus. It is a close relative of M. guentheri distinguished by different pelage coloration and a more complex occlusal...

          , Microtus qazvinensis
        • Schidlovsky's Vole
          Schidlovsky's Vole
          Schidlovsky's Vole is a species of rodent in the family Cricetidae. It is normally found in north-western Armenia, and was long considered a subspecies of the Social Vole until relisted as a species by Golenishchev in 2002....

          , Microtus schidlovskii
        • Social Vole
          Social Vole
          The Social Vole is a species of rodent in the family Cricetidae.It is found in China, Iran, Kazakhstan, Syria, Turkey, and Ukraine.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

          , Microtus socialis
        • European Pine Vole
          European Pine Vole
          The European Pine Vole is a species of rodent in the family Cricetidae.It is found in Albania, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Estonia, France, Germany, Greece, Hungary, Italy, Liechtenstein, Luxembourg, Macedonia, Moldova, the Netherlands, Poland, Romania, Russia,...

          , Microtus subterraneus
        • Transcaspian Vole
          Transcaspian Vole
          The Transcaspian Vole is a species of rodent in the family Cricetidae.It is found in Afghanistan, Iran, and Turkmenistan.-References:...

          , Microtus transcaspicus
      • Subgenus Terricola
        Terricola (subgenus)
        Terricola is a subgenus of voles in the genus Microtus. The authority for this species is Fatio in 1867, but it is commonly given as Flemming in 1828...

        • Bavarian Pine Vole
          Bavarian pine vole
          The Bavarian pine vole is a vole from the Austrian, Italian, and Bavarian Alps of Europe. It lived in moist meadows at elevations of 600-1,000 metres...

          , Microtus bavaricus
        • Calabria Pine Vole
          Calabria Pine Vole
          The Calabria Pine Vole is a vole found in southern and central Italy initially described by Lehmann as a subspecies of M. savii. Genetic tests in the Calabrian region found that, although similar, the X chromosome is larger than that of samples of M...

          , Microtus brachycercus
        • Daghestan Pine Vole
          Daghestan Pine Vole
          The Daghestan Pine Vole is a species of rodent in the family Cricetidae.It is found in Russia, Georgia, Armenia, and Azerbaijan.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

          , Microtus daghestanicus
        • Mediterranean Pine Vole
          Mediterranean Pine Vole
          The Mediterranean Pine Vole is a species of rodent in the family Cricetidae.It is found in France, Portugal, and Spain.-References:* Amori, G. 1996. . Downloaded on 19 July 2007....

          , Microtus duodecimcostatus
        • Felten's Vole
          Felten's Vole
          Felten's Vole is a species of rodent in the family Cricetidae.It is found in Albania, Greece, Macedonia, and Serbia and Montenegro.-References:...

          , Microtus felteni
        • Liechtenstein's Pine Vole
          Liechtenstein's Pine Vole
          Liechtenstein's Pine Vole is a species of rodent in the family Cricetidae. It is found in central and eastern Europe, from northern Italy through to Austria, Slovenia and Croatia.-References:...

          , Microtus liechtensteini
        • Lusitanian Pine Vole
          Lusitanian Pine Vole
          The Lusitanian Pine Vole is a species of rodent in the family Cricetidae.It is found in France, Portugal, and Spain.-References:...

          , Microtus lusitanicus
        • Major's Pine Vole
          Major's Pine Vole
          Major's Pine Vole is a species of rodent in the family Cricetidae found in Turkey....

          , Microtus majori
        • Alpine Pine Vole
          Alpine Pine Vole
          The Alpine Pine Vole is a species of rodent in the family Cricetidae.It is found in Austria, France, Italy, Serbia and Montenegro, and Switzerland.-References:* Amori, G. 1996. . Downloaded on 19 July 2007....

          , Microtus multiplex
        • Savi's Pine Vole
          Savi's Pine Vole
          Savi's Pine Vole is a species of rodent in the family Cricetidae.It is found in France and Italy.-References: Database entry includes a brief justification of why this species is of least concern....

          , Microtus savii
        • Tatra Pine Vole
          Tatra Pine Vole
          The Tatra Pine Vole, Microtus tatricus, also called the Tatra vole or Tatra ground vole, is a species of vole endemic to the Carpathian mountain range in Slovakia, Poland, Ukraine and Romania. Two subspecies have been recognised. M. t. tatricus occupies the western part of the range in Slovakia and...

          , Microtus tatricus
        • Thomas's Pine Vole
          Thomas's Pine Vole
          Thomas's Pine Vole is a species of rodent in the family Cricetidae.It is found in Bosnia and Herzegovina, Greece, Macedonia, Serbia, Montenegro, and possibly Albania.-References:...

          , Microtus thomasi
      • Subgenus Mynomes
        Mynomes
        Mynomes is a subgenus of voles in the genus Microtus. Species in this subgenus are:*Beach Vole *Gray-tailed Vole *Montane Vole *Creeping Vole...

        • Beach Vole
          Beach Vole
          The Beach Vole is a species of rodent in the family Cricetidae. It is found only in the United States, and its natural habitat is temperate grassland....

          , Microtus breweri
        • Gray-tailed Vole
          Gray-tailed Vole
          The Gray-tailed vole is a common small mammal that inhabits grasslands in the Willamette Valley of western Oregon . Gray tailed voles are polygamous: females are territorial, males have large home ranges that overlap those of several females, and juvenile emigration is male biased...

          , Microtus canicaudus
        • Montane Vole
          Montane Vole
          The Montane Vole is a species of rodent in the family Cricetidae.It is found in the western United States and British Columbia in Canada.-References:...

          , Microtus montanus
        • Creeping Vole
          Creeping Vole
          The Creeping Vole is a species of rodent in the family Cricetidae.It is found in British Columbia in Canada and in California, Oregon and Washington in the United States.-References:...

          , Microtus oregoni
        • Meadow Vole
          Meadow Vole
          The Meadow Vole , sometimes called the Field Mouse or Meadow Mouse, is a North American vole found across Canada, Alaska and the northern United States. Its range extends further south along the Atlantic coast. One subspecies, the Florida Salt Marsh Vole , is found in Florida, and is classified as...

          , Microtus pennsylvanicus
        • Townsend's Vole
          Townsend's Vole
          Townsend's Vole is a species of rodent in the family Cricetidae.It is found in British Columbia in Canada and in Washington and Oregon in the United States.Its natural habitat is temperate grassland.-References:...

          , Microtus townsendii
      • Subgenus Alexandromys
        Alexandromys
        Alexandromys is a subgenus of voles in the genus Microtus. Species in this subgenus are:*Clarke's Vole *Evorsk Vole *Reed Vole *Gerbe's Vole...

        • Clarke's Vole
          Clarke's Vole
          Clarke's Vole is a species of rodent in the Cricetidae family.It is found only in China.-References:* Baillie, J. 1996. . Downloaded on 20 July 2007....

          , Microtus clarkei
        • Evorsk Vole
          Evorsk Vole
          The Evorsk Vole is a species of rodent in the family Cricetidae.It is found only in Russia. They are stout rodents with short ears, legs and tails. These animals live in grassy areas where they eat green vegetation such as grasses and sedges in summer and grains, seeds, roots and bark at other...

          , Microtus evoronensis
        • Reed Vole
          Reed Vole
          The Reed Vole is a species of vole. It is found in northern and central Eurasia, including northern China and the Korean Peninsula. The species is somewhat larger and longer-tailed than most other voles. An adult Reed Vole has a body 12-18 centimeters long, and a tail of 4.7-5.7...

          , Microtus fortis
        • Gerbe's Vole
          Gerbe's Vole
          Gerbe's Vole is a species of rodent in the family Cricetidae.It is found in France and Spain.-References:* Amori, G. 2000. . Downloaded on 19 July 2007....

          , Microtus gerbei
        • Taiwan Vole
          Taiwan Vole
          The Taiwan Vole is a species of rodent in the family Cricetidae.It is found only in Taiwan.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

          , Microtus kikuchii
        • Lacustrine Vole
          Lacustrine Vole
          The Lacustrine Vole is a species of rodent in the family Cricetidae.It is found in China and Mongolia.-References:...

          , Microtus limnophilus
        • Maximowicz's Vole
          Maximowicz's Vole
          Maximowicz's Vole is a species of rodent in the family Cricetidae.It is found in China, Mongolia, and Russia.-References:...

          , Microtus maximowiczii
        • Middendorf's Vole
          Middendorf's Vole
          Middendorf's Vole is a species of rodent in the family Cricetidae.It is found only in Russia.-References:...

          , Microtus middendorffi
        • Mongolian Vole
          Mongolian Vole
          The Mongolian Vole is a species of rodent in the family Cricetidae.It is found in China, Mongolia, and Russia.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

          , Microtus mongolicus
        • Japanese Grass Vole
          Japanese Grass Vole
          The Japanese Grass Vole is a species of rodent in the family Cricetidae.It is found only in Japan.-References:* Baillie, J. 1996. . Downloaded on 9 July 2007....

          , Microtus montebelli
        • Muisk Vole
          Muisk Vole
          The Muisk Vole is a species of rodent in the family Cricetidae.It is found only in Russia.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

          , Microtus mujanensis
        • Tundra Vole (Root Vole)
          Tundra Vole
          The Tundra Vole or Root Vole, Microtus oeconomus, is a medium-sized vole found in Northern and Central Europe, Asia, and northwestern North America, including Alaska and northwestern Canada...

          , Microtus oeconomus
        • Sakhalin Vole
          Sakhalin Vole
          The Sakhalin Vole is a species of rodent in the family Cricetidae.It is found only in Russia.-References:...

          , Microtus sachalinensis
      • Subgenus Stenocranius
        • Narrow-headed Vole
          Narrow-Headed Vole
          The Narrow-headed Vole is a species of rodent in the family Cricetidae.It is found in China, Mongolia, and Russia. It is the only species in the subgenus Stenocranius-References:...

          , Microtus gregalis
      • Subgenus Pitymys
        Pitymys
        Pitymys is a subgenus of voles in the genus Microtus. Species in this subgenus are:*Guatemalan Vole *Tarabundí Vole *Woodland Vole *Jalapan Pine Vole...

        • Guatemalan Vole
          Guatemalan Vole
          The Guatemalan vole is a species of rodent in the family Cricetidae.It is found in Guatemala and Chiapas, Mexico in montane pine-oak forest and meadow at elevations between 2600 to 3100 m. It is terrestrial and probably diurnal or crepuscular....

          , Microtus guatemalensis
        • Tarabundí Vole
          Tarabundí Vole
          The Tarabundí Vole is a species of rodent in the family Cricetidae.It is found only in Mexico.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

          , Microtus oaxacensis
        • Woodland Vole
          Woodland Vole
          The Woodland Vole, Microtus pinetorum, is a small vole found in eastern North America. It is also known as the Pine Vole.-Description:...

          , Microtus pinetorum
        • Jalapan Pine Vole
          Jalapan Pine Vole
          The Jalapan Pine Vole is a species of rodent in the family Cricetidae.It is found only in Mexico.-References:...

          , Microtus quasiater
      • Subgenus Pedomys
        • Prairie Vole
          Prairie Vole
          The Prairie Vole is a small vole found in central North America. The vole has long, coarse grayish-brown fur on the upper portion of the body and yellowish fur on the lower portion of the body...

          , Microtus ochrogaster
      • Subgenus Hyrcanicola
        • Schelkovnikov's Pine Vole
          Schelkovnikov's Pine Vole
          Schelkovnikov's Pine Vole is a species of rodent in the family Cricetidae. It is found in Azerbaijan and Iran. It has recently been considered the sole species in the subgenus Hyrcanicola.-References:...

          , Microtus schelkovnikovi
    • Genus Neodon
      Neodon
      Neodon is a genus of rodent in the family Cricetidae. Species within the Neodon genus are classified as relics of the Pleistocene epoch because the occlusal patterns resemble the extinct Allophaiomys.The Neodon genus contains the following species:...

       - mountain voles
      • Juniper Vole
        Juniper Vole
        The Juniper Vole is a species of rodent in the family Cricetidae.It is found in Afghanistan, China, Pakistan, Russia, and Tajikistan.-References:...

        , Neodon juldaschi
      • Chinese Scrub Vole
        Chinese Scrub Vole
        The Chinese Scrub Vole is a species of rodent in the family Cricetidae.It is found in China and Myanmar.-References:...

        , Neodon irene
      • Sikkim Vole
        Sikkim Vole
        The Sikkim Mountain Vole is a species of rodent in the family Cricetidae. It is found in Bhutan, China, India, and Nepal.-References:...

        , Neodon sikimensis
      • Forrest's Vole
        Forrest's Vole
        Forrest's Mountain Vole is a species of rodent in the family Cricetidae. It is found within china in the north west of Yunnan province. The initial study by Hinton in 1923 identified it as morphologically close to the Chinese Scrub Vole but with a larger body size and longer and darker Pelage....

        , Neodon forresti
    • Genus Phaiomys
      • Blyth's Vole
        Blyth's Vole
        Blyth's Vole is a species of rodent in the family Cricetidae. It is the only species in the genus Phaiomys.It is found in China and India.-References:...

        , Phaiomys leucurus
    • Genus Proedromys
      Proedromys
      Proedromys is a genus of rodent in the family Cricetidae from China. It is part of the tribe Arvicolini within the subfamily Arvicolinae, which contains the voles, lemmings, and relatives.It contains the following species:...

      • Duke of Bedford's Vole
        Duke of Bedford's Vole
        The Duke of Bedford's Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:...

        , Proedromys bedfordi
      • Proedromys liangshanensis
    • Genus Volemys
      Volemys
      Volemys is a small genus of rodents in the family Cricetidae.It contains the following species:* Szechuan Vole * Marie's Vole Both members of this genus are endemic to China.-References:...

      • Szechuan Vole
        Szechuan Vole
        The Szechuan Vole is a species of rodent in the family Cricetidae.It is found only in China. It is one of two species in the genus Volemys along with Marie's Vole .-References:...

        , Volemys millicens
      • Marie's Vole
        Marie's Vole
        Marie's Vole is a species of rodent in the family Cricetidae.It is found only in China. Volemys musseri is one of two species in the genus Volemys along with the Szechuan Vole .-References:...

        , Volemys musseri
  • Tribe Dicrostonychini
    Dicrostonychini
    Dicrostonychini is a tribe of lemmings in the subfamily Arvicolinae. It contains only one extant genus, as well as one extinct genus....

     - collared lemmings
    • Genus Dicrostonyx
      • St. Lawrence Island Collared Lemming, Dicrostonyx exsul (Dicrostonyx groenlandicus exsul)
      • Northern Collared Lemming
        Northern Collared Lemming
        The Northern Collared Lemming , sometimes called the Peary Land Collared Lemming in Canada, is a small North American lemming. At one time, it was considered to be a subspecies of the Arctic Lemming...

        , Dicrostonyx groenlandicus
      • Ungava Collared Lemming
        Ungava Collared Lemming
        The Ungava Collared Lemming or Labrador Collared Lemming, Dicrostonyx hudsonius is a small North American lemming.They have short chunky bodies covered with brownish-grey fur with a thin dark stripe along their back and a yellow line along their sides. They have small ears, short legs and a very...

        , Dicrostonyx hudsonius
      • Victoria Collared Lemming, Dicrostonyx kilangmiutak (Dicrostonyx groenlandicus kilangmiutak)
      • Nelson's Collared Lemming
        Nelson's Collared Lemming
        Nelson's Collared Lemming is a species of rodent in the family Cricetidae.It is found in western and southwestern Alaska in the United States.-References:...

        , Dicrostonyx nelsoni (Dicrostonyx groenlandicus nelsoni)
      • Ogilvie Mountains Collared Lemming
        Ogilvie Mountains Collared Lemming
        The Ogilvie Mountains Collared Lemming is a species of rodent in the family Cricetidae.It is found only in Canada.Its natural habitat is tundra....

        , Dicrostonyx nunatakensis
      • Richardson's Collared Lemming
        Richardson's Collared Lemming
        The Richardson's Collared Lemming, Dicrostonyx richardsoni is a small North American lemming. At one time, they were considered to be a subspecies of the Arctic Lemming, Dicrostonyx torquatus...

        , Dicrostonyx richardsoni
      • Bering Collared Lemming, Dicrostonyx rubricatus (Dicrostonyx groenlandicus rubricatus)
      • Arctic Lemming
        Arctic Lemming
        The Arctic Lemming is a species of rodent in the family Cricetidae.It is found only in The Arctic Biomes in Russia and Canada.-References:...

        , Dicrostonyx torquatus
      • Unalaska Collared Lemming
        Unalaska Collared Lemming
        The Unalaska Collared Lemming is a species of rodent in the family Cricetidae.This species is found on two islands, Umnak and Unalaska, in the Aleutian Archipelago of Alaska in the United States....

        , Dicrostonyx unalascensis
      • Wrangel Lemming
        Wrangel Lemming
        The Wrangel Lemming is a species of rodent in the family Cricetidae.It is found only in Russia.Its natural habitat is tundra.It is threatened by habitat loss.-References:...

        , Dicrostonyx vinogradovi
  • Tribe Ellobiusini - mole voles
    • Genus Ellobius
      Ellobius
      Ellobius is a genus of rodent in the family Cricetidae.It contains the following species:* Alai Mole Vole * Southern Mole Vole * Transcaucasian Mole Vole...

       - mole voles
      • Alai Mole Vole
        Alai Mole Vole
        The Alai Mole Vole is a species of rodent in the family Cricetidae. It is known only from Kyrgyzstan, where it has been found in temperate grassland in the Alai Mountains. Little else is known about the vole....

        , Ellobius alaicus
      • Southern Mole Vole
        Southern Mole Vole
        The Southern Mole Vole is a species of rodent in the family Cricetidae.It is found in Afghanistan, Iran, Pakistan, and Turkmenistan.-References:...

        , Ellobius fuscocapillus
      • Transcaucasian Mole Vole
        Transcaucasian Mole Vole
        The Transcaucasian Mole Vole is a species of rodent in the family Cricetidae.It is found in Armenia, Azerbaijan, Georgia, Iran, and Turkey.-References:...

        , Ellobius lutescens
      • Northern Mole Vole
        Northern Mole Vole
        The northern mole vole is a species of rodent in the family Cricetidae.It is distributed over large parts of Eastern Europe and Asia.-Distribution:...

        , Ellobius talpinus
      • Zaisan Mole Vole
        Zaisan Mole Vole
        The Zaisan Mole Vole is a species of rodent in the family Cricetidae.It is found in China, Kazakhstan, Mongolia, Turkmenistan, and Uzbekistan.-References:...

        , Ellobius tancrei
  • Tribe Lagurini
    • Genus Eolagurus
      Eolagurus
      Eolagurus is a genus of rodent in the family Cricetidae.It contains the following species:* Yellow Steppe Lemming * Przewalski's Steppe Lemming -References:...

      • Yellow Steppe Lemming
        Yellow Steppe Lemming
        The Yellow Steppe Lemming is a species of rodent in the family Cricetidae.It is found in China, Kazakhstan, and Mongolia.Its natural habitat is temperate desert.-References:...

        , Eolagurus luteus
      • Przewalski's Steppe Lemming
        Przewalski's Steppe Lemming
        Przewalski's Steppe Lemming is a species of rodent in the family Cricetidae.It is found in China and Mongolia.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

        , Eolagurus przewalskii
    • Genus Lagurus
      • Steppe Lemming
        Steppe Lemming
        The steppe lemming or steppe vole is a small, plump, light grey rodent, similar in appearance to the Norway lemming but not in the same genus. The steppe lemming eats shoots and leaves and is more active at night though it is not strictly nocturnal. In the wild it is found in Russia and Ukraine...

        , Lagurus lagurus
  • Tribe Lemmini - lemmings
    • Genus Lemmus - true lemmings
      • Amur Lemming
        Amur Lemming
        The Amur Lemming is a species of lemming found near the Amur River on the side of Siberia. The Amur Lemming is on the IUCN Red List.-External links:IUCN Red List information:...

        , Lemmus amurensis
      • Norway lemming
        Norway lemming
        The Norway lemming , Lemmus lemmus, is a common species of lemming found in northern Scandinavia and adjacent areas of Russia. It is the only vertebrate species endemic to the region. The Norway lemming dwells in tundra and fells, and prefers to live near water. Adults feed primarily on...

        , Lemmus lemmus
      • Siberian Brown Lemming
        Siberian Brown Lemming
        The Siberian Brown Lemming is a species of rodent in the family Cricetidae.It is found in Canada, Russian Federation, and United States.It does not hibernate during winter, because it lives in burrows. It is prey to several animals including the snowy owl and the Arctic fox.-References:*Musser,...

        , Lemmus sibiricus
      • North American Brown Lemming
        North American Brown Lemming
        The North American brown lemming, Lemmus trimucronatus, is a small North American lemming. Originally called the Siberian Brown Lemming Lemmus sibiricus it was later decided that they formed two distinct species....

        , Lemmus trimucronatus
      • Wrangel Island Lemming
        Wrangel Island Lemming
        The Wrangel Island Lemming or Lemmus portenkoi, is a species of lemming endemic to Wrangel Island in Russia. It was included as a subspecies of L. sibiricus by Jarrell and Fredga in 1993, while Chernyavskii regarded it as a separate species.-References:*D.E. Wilson & D.M. Reeder, 2005: Mammal...

        , Lemmus portenkoi
    • Genus Myopus
      • Wood Lemming
        Wood Lemming
        The Wood Lemming is a species of rodent in the family Cricetidae. It belongs into the Arvicolinae subfamily of rodents therefore is a relative of the voles, lemmings, and muskrats....

        , Myopus schisticolor
    • Genus Synaptomys - bog lemmings
      • Northern Bog Lemming
        Northern Bog Lemming
        The Northern Bog Lemming, Synaptomys borealis is a small North American lemming. This is one of two species in genus Synaptomys, the other being the Southern Bog Lemming....

        , Synaptomys borealis
      • Southern Bog Lemming
        Southern Bog Lemming
        The Southern Bog Lemming, Synaptomys cooperi is a small North American lemming. Its range overlaps with the other species in genus Synaptomys, the Northern Bog Lemming, in southeastern Canada but extends further south....

        , Synaptomys cooperi
  • Tribe Myodini
    Myodini
    Myodini is a tribe of forest voles in the subfamily Arvicolinae. Species in this tribe are:Genus Alticola - voles from Central Asia*Subgenus Alticola**White-tailed Mountain Vole, Alticola albicauda**Silver Mountain Vole, Alticola argentatus...

    • Genus Alticola
      Alticola
      Alticola is a genus of rodent in the family Cricetidae.It contains the following species:*Subgenus Alticola** White-tailed Mountain Vole ** Silver Mountain Vole...

       - voles from Central Asia
      Central Asia
      Central Asia is a core region of the Asian continent from the Caspian Sea in the west, China in the east, Afghanistan in the south, and Russia in the north...

      • Subgenus Alticola
        • White-tailed Mountain Vole
          White-tailed Mountain Vole
          The White-tailed Mountain Vole is a species of vole in the family Cricetidae. It is found in India and Pakistan....

          , Alticola albicauda
        • Silver Mountain Vole
          Silver Mountain Vole
          The Silver Mountain Vole is a species of rodent in the family Cricetidae.It is found in the Tien Shan and Pamir mountains of Kazakhstan, Kyrgyzstan, Tajikistan, Afghanistan, China, and Pakistan....

          , Alticola argentatus
        • Gobi Altai Mountain Vole
          Gobi Altai Mountain Vole
          The Gobi Altai Mountain Vole is a species of rodent in the family Cricetidae. It can be found in China, Mongolia, and the Russian Federation....

          , Alticola barakshin
        • Central Kashmir Vole
          Central Kashmir Vole
          The Central Kashmir Vole is a species of rodent in the family Cricetidae.It is found only in India and Pakistan....

          , Alticola montosa
        • Royle's Mountain Vole
          Royle's Mountain Vole
          Royle's Mountain Vole is a species of rodent in the family Cricetidae.It is found in China and India....

          , Alticola roylei
        • Mongolian Silver Vole
          Mongolian Silver Vole
          The Mongolian Silver Vole is a species of rodent in the family Cricetidae.It is found only in Mongolia....

          , Alticola semicanus
        • Stoliczka's Mountain Vole
          Stoliczka's Mountain Vole
          Stolička's or Stoliczka's Mountain Vole is a species of rodent in the family Cricetidae.It is found in China, Pakistan, India, and Nepal....

          , Alticola stoliczkanus
        • Tuva Silver Vole
          Tuva Silver Vole
          The Tuva Silver Vole is a species of rodent in the family Cricetidae.It is found in Mongolia and Russian Federation....

          , Alticola tuvinicus
      • Subgenus Aschizomys
        • Lemming Vole
          Lemming Vole
          The Lemming Vole is a species of rodent in the family Cricetidae. It is found only in Russian Federation....

          , Alticola lemminus
        • Large-eared Vole
          Large-eared Vole
          The Large-eared Vole is a species of rodent in the family Cricetidae.It is found in Mongolia and Russian Federation....

          , Alticola macrotis
        • Lake Baikal Mountain Vole
          Lake Baikal Mountain Vole
          The Lake Baikal or Olkhon Mountain Vole is a species of rodent in the family Cricetidae. It is found principally on the Olkhon and Ogoi islands on Lake Baikal. While it was originally described as a species, Pavlinov and Rossolimo reassigned it as a subspecies of A. tuvinicus in 1987 before...

          , Alticola olchonensis
      • Subgenus Platycranius
        • Flat-headed Vole
          Flat-headed Vole
          The Flat-headed Vole is a species of rodent in the family Cricetidae.It is found in China, Kazakhstan, Mongolia, and Russian Federation....

          , Alticola strelzowi
    • Genus Caryomys
      • Ganzu Vole
        Ganzu Vole
        The Ganzu Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

        , Caryomys eva
      • Kolan Vole
        Kolan Vole
        The Kolan Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

        , Caryomys inez
    • Genus Eothenomys
      Eothenomys
      Eothenomys is a genus of rodent in the family Cricetidae.It contains the following species:* Kachin Red-backed Vole * Pratt's Vole * Southwest China Vole...

       - voles from East Asia
      East Asia
      East Asia or Eastern Asia is a subregion of Asia that can be defined in either geographical or cultural terms...

      • Kachin Red-backed Vole
        Kachin Red-backed Vole
        The Kachin Red-backed Vole is a species of rodent in the family Cricetidae. It is found in Kachin State in northern Burma. Although described as a species by Thomas in 1921, it was subsequently included as a subspecies of either E. melanogaster or E. miletus. Comparisons of a 1941 sampling showed...

        , Eothenomys cachinus
      • Pratt's Vole
        Pratt's Vole
        Pratt's Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

        , Eothenomys chinensis
      • Southwest China Vole
        Southwest China Vole
        The Southwest China Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:* Baillie, J. 1996. . Downloaded on 19 July 2007....

        , Eothenomys custos
      • Pére David's Vole
        Père David's Vole
        Père David's Vole is a species of rodent in the family Cricetidae.It is found in China, Myanmar, Taiwan, and Thailand.-References:...

        , Eothenomys melanogaster
      • Yunnan Red-backed Vole
        Yunnan Red-backed Vole
        The Yunnan Red-backed Vole is a species of rodent in the family Cricetidae. It is found in Southwest China, specifically Yunnan Province. It is the largest member of the Eothenomys genus, with a higher cranium, a soft, thick and long coat with tawny brown to reddish brown coloring and grey...

        , Eothenomys miletus
      • Chaotung Vole
        Chaotung Vole
        The Chaotung Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

        , Eothenomys olitor
      • Yulungshan Vole
        Yulungshan Vole
        The Yulungshan Vole is a species of rodent in the family Cricetidae, found only in China.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

        , Eothenomys proditor
      • Ward's Red-backed Vole
        Ward's Red-backed Vole
        The Ward's Red-backed Vole is a species of rodent in the family Cricetidae. It is found in China, specifically the north-western area of Yunnan Province. It is sometimes considered a subspecies of Eothenomys chinensis, but most zoologists consider it to be a separate species due to the noticeably...

        , Eothenomys wardi
    • Genus Hyperacrius
      Hyperacrius
      Hyperacrius is a genus of rodent in the family Cricetidae.It contains the following species:* True's Vole * Murree Vole -References:...

       - voles from Pakistan
      Pakistan
      Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

      • True's Vole
        True's Vole
        True's Vole is a species of rodent in the family Cricetidae.It is found in India and Pakistan.-References:...

        , Hyperacrius fertilis
      • Murree Vole
        Murree Vole
        The Murree Vole is a species of rodent in the family Cricetidae. It is endemic to Pakistan.-Distribution:It is confined to the Himalayas including Murree Hills, Kaghan Valley and Swat Valley.-References:...

        , Hyperacrius wynnei
    • Genus Myodes - red-backed voles
      • Anderson's Red-backed Vole, Myodes andersoni
      • Western Red-backed Vole
        Western Red-Backed Vole
        The Western Red-Backed Vole is a species of vole in the family Cricetidae. It is found in California and Oregon in the United States and lives mainly in coniferous forest...

        , Myodes californicus
      • Tien Shan Red-backed Vole
        Tien Shan Red-Backed Vole
        The Tien Shan Red-backed Vole is a species of rodent in the family Cricetidae.It is found in China and Kyrgyzstan.-References:...

        , Myodes centralis
      • Southern Red-backed Vole
        Southern Red-backed Vole
        The Southern Red-backed Vole or Gapper's Red-backed Vole is a small slender vole found in Canada and the northern United States...

        , Myodes gapperi
      • Bank Vole
        Bank Vole
        The bank vole is a small vole with red-brown fur and some grey patches, with a tail about half as long as its body. A rodent, it lives in woodland areas and is around in length. The bank vole is found in western Europe and northern Asia...

        , Myodes glareolus
      • Imaizumi's Red-backed Vole
        Imaizumi's Red-backed Vole
        Imaizumi's Red-backed Vole Myodes imaizumii is a species of vole in the family Cricetidae. It is found in Japan and was initially designated as a subspecies of the Japanese Red-Backed Vole, with studies of cranial and molar patterns supporting this. However, genetic tests in the late 1990s provide...

        , Myodes imaizumii
      • Korean Red-backed Vole, Myodes regulus
      • Hokkaido Red-backed Vole
        Hokkaido Red-backed Vole
        The Hokkaido Red-backed Vole is a species of rodent in the family Cricetidae.It is found only in Japan.Its natural habitat is temperate forests.-References:...

        , Myodes rex
      • Grey Red-backed Vole
        Grey Red-Backed Vole
        The Grey Red-backed Vole is a species of vole. An adult Grey Red-backed Vole weighs 20-50 grams, and has a body length of 110-142 millimeters, with a tail of 39–55 mm in length. This species ranges across northern Eurasia, including northern China, the northern Korean Peninsula, and...

        , Myodes rufocanus
      • Northern Red-backed Vole
        Northern Red-backed Vole
        The northern red-backed vole, Myodes rutilus, is a small slender vole found in Alaska, northern Canada, Scandinavia and northern Russia.-Description:...

        , Myodes rutilus
      • Shansei Vole
        Shansei Vole
        The Shansei Vole is a species of rodent in the family Cricetidae.It is found only in China.-References:*Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns...

        , Myodes shanseius
      • Smith's Vole
        Smith's Vole
        Smith's vole is a species of rodent in the family Cricetidae. It is also known as Smith's red-backed vole and is found only in Japan....

        , Myodes smithii
  • Tribe Neofibrini
    • Genus Neofiber
      • Round-tailed Muskrat
        Round-Tailed Muskrat
        The Round-tailed Muskrat is a species of rodent in the family Cricetidae, sometimes called the Florida water rat. It is the only species in the genus Neofiber....

        , Neofiber alleni
  • Tribe Ondatrini - muskrat
    • Genus Ondatra
      • Muskrat
        Muskrat
        The muskrat , the only species in genus Ondatra, is a medium-sized semi-aquatic rodent native to North America, and introduced in parts of Europe, Asia, and South America. The muskrat is found in wetlands and is a very successful animal over a wide range of climates and habitats...

        , Ondatra zibethicus
  • Tribe Pliomyini
    • Genus Dinaromys
      • Balkan Snow Vole
        Balkan Snow Vole
        The Balkan Snow Vole, Dinaromys bogdanovi, also known as Martino's Snow Vole is the only member of the genus Dinaromys. Eight subspecies of this vole have been recognized from southern parts of Europe. The genus name means "Dinaric mouse", referring to the Dinaric Alps...

        , Dinaromys bogdanovi
  • Tribe Prometheomyini
    • Genus Prometheomys
      • Long-clawed Mole Vole
        Long-Clawed Mole Vole
        The Long-clawed Mole Vole is a species of rodent in the family Cricetidae. It is the only species in the genus Prometheomys.It is found in Georgia and Turkey.-References:...

        , Prometheomys schaposchnikowi
  • incertae sedis
    • Genus Arborimus - tree voles
      • White-footed Vole
        White-footed Vole
        The White-footed Vole is a species of rodent in the family Cricetidae. It is found only in the United States. Its natural habitat is temperate forests.-Source:...

        , Arborimus albipes
      • Red Tree Vole
        Red Tree Vole
        The Red Tree Vole is a species of rodent in the family Cricetidae.It is found only in the United States. Red tree voles are found in the upper branches of arboreal conifers throughout Northern California and Western Oregon. Their preference and food source is thought to be Douglas Fir and Redwood...

        , Arborimus longicaudus
      • Sonoma Tree Vole, Arborimus pomo
    • Genus Phenacomys - heather voles
      • Western Heather Vole
        Western Heather Vole
        The Western Heather Vole, Phenacomys intermedius, is a small vole found in western North America. Until recently, the Eastern Heather Vole, Phenacomys ungava, was considered to be a subspecies....

        , Phenacomys intermedius
      • Eastern Heather Vole
        Eastern Heather Vole
        The Eastern Heather Vole, Phenacomys ungava, is a small North American vole. Until recently, this species was considered to belong to the same species as the Western Heather Vole, Phenacomys intermedius. It is also called the Ungava Vole....

        , Phenacomys ungava

Fossil species

  • Tribe Dicrostonychini
    Dicrostonychini
    Dicrostonychini is a tribe of lemmings in the subfamily Arvicolinae. It contains only one extant genus, as well as one extinct genus....

     - collared lemmings
    • Genus Predicrostonyx
      Predicrostonyx
      Predicrostonyx is a genus of extinct rodent in the family Cricetidae.It contains only the species, Hopkins's Lemming, Predicrostonyx hopkinsi, which is considered one of the earliest examples of collared lemmings.-References:...

       †
      • Hopkins's Lemming, Predicrostonyx hopkinsi †
  • Tribe Clethrionomyini
    • Genus Altaiomys
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