Banksia ser. Cyrtostylis
Encyclopedia
Banksia ser. Cyrtostylis is a valid botanic name for a taxonomic
Taxonomy
Taxonomy is the science of identifying and naming species, and arranging them into a classification. The field of taxonomy, sometimes referred to as "biological taxonomy", revolves around the description and use of taxonomic units, known as taxa...

 series
Series (botany)
Series is a low-level taxonomic rank below that of section but above that of species.In botany, a series is a subdivision of a genus...

 within the plant
Plant
Plants are living organisms belonging to the kingdom Plantae. Precise definitions of the kingdom vary, but as the term is used here, plants include familiar organisms such as trees, flowers, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The group is also called green plants or...

 genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

. First published at section
Section (botany)
In botany, a section is a taxonomic rank below the genus, but above the species. The subgenus, if present, is higher than the section, and the rank of series, if present, is below the section. Sections are typically used to help organise very large genera, which may have hundreds of species...

al rank by George Bentham
George Bentham
George Bentham CMG FRS was an English botanist, characterized by Duane Isely as "the premier systematic botanist of the nineteenth century".- Formative years :...

 in 1870, it was demoted to a series by Alex George
Alex George
Alexander Segger George is a Western Australian botanist. He is the authority on the plant genera Banksia and Dryandra...

 in 1981. The name has had three circumscriptions.

According to Bentham

Bentham published B. sect. Cyrtostylis in Flora Australiensis
Flora Australiensis
Flora Australiensis: a description of the plants of the Australian Territory, more commonly referred to as Flora Australiensis, and also known by its standard abbreviation Fl. Austral., is a seven-volume flora of Australia published between 1863 and 1878 by George Bentham, with the assistance of...

, as part of his taxonomic arrangement of Banksia
Bentham's taxonomic arrangement of Banksia
George Bentham's taxonomic arrangement of Banksia was published in 1870, in Volume 5 of Bentham's Flora Australiensis. A substantial improvement on the previous arrangement, it would stand for over a century. It was eventually replaced by Alex George's 1981 arrangement, published in his classic...

, defining it as those taxa with This definition essentially defined Cyrtostylis as a section for those species that did not possess the characters of the other sections; thus it was highly heterogeneous.

George Bentham's placement and circumscription of B. sect. Cyrtostylis may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. sect. Oncostylis
Banksia sect. Oncostylis
Banksia sect. Oncostylis is one of four sections of subgenus Banksia subg. Banksia. It contains those Banksia species with hooked pistils. All of the species in Oncostylis also exhibit a top-down sequence of flower anthesis, except for Banksia nutans which is bottom-up.Banksia sect...

 (13 species, 2 varieties)
B. sect. Cyrtostylis
B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

B. Solandri
Banksia solandri
Banksia solandri, commonly known as Stirling Range Banksia, is a species of large shrub in the plant genus Banksia. It occurs only within the Stirling Range in southwest Western Australia.-Description:...

B. Goodii
Banksia goodii
Banksia goodii, commonly known as Good's Banksia, is an endangered shrub of Southwest Western Australia.Good's Banksia grows as a low shrub, either prostrate or with stems up to twenty centimetres high. The leaves are dark green with a prominent yellow midrib, and are held erect. They may be up...

B. petiolaris
Banksia petiolaris
Banksia petiolaris is a species of shrub of the genus Banksia in the Proteaceae family. It is a prostrate banksia, a group of several closely related species all with horizontal stems and thick, leathery upright leaves. No varieties have been subsequently described. It bears yellow inflorescences...

B. repens
Banksia repens
Banksia repens, the Creeping Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Ragged in the east....

B. prostrata (now B. gardneri
Banksia gardneri
The Prostrate Banksia is a species of prostrate shrub in the plant genus Banksia. It occurs along the south coast of Western Australia.-Scientific name:...

)
B. grandis
Banksia grandis
Banksia grandis, commonly known as Bull Banksia, Giant Banksia or Mangite, is a common and distinctive tree in South West Western Australia....

B. quercifolia
Banksia quercifolia
The Oak-leaved Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia from D'Entrecasteaux National Park in the west to Mount Manypeaks in the east.-Ecology:...

B. quercifolia var. integrifolia (now B. oreophila
Banksia oreophila
The Western Mountain Banksia or Mountain Banksia is a species of shrub in the plant genus Banksia. It occurs on the slopes and hilltops of the Stirling and Barren Ranges in southwest Western Australia.-Taxonomy:...

)
B. Baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

B. sect. Eubanksia
Banksia sect. Eubanksia
Banksia sect. Eubanksia is an obsolete section of Banksia. There have been two circumscriptions, one of which is synonymous with the recently abandoned B. subg. Banksia sensu Alex George, the other having no modern equivalent....

 (3 species)
B. sect. Orthostylis (19 species)
B. sect. Isostylis (1 species, 1 variety)

Bentham's circumscription of Cyrtostylis remained current for over 100 years, only being superseded in 1981 by George's revision.

According to George 1981

In 1981, George published a taxonomic revision of Banksia
George's taxonomic arrangement of Banksia
Alex George's taxonomic arrangement of Banksia was the first modern-day arrangement for that genus. First published in 1981 in the classic monograph The genus Banksia L.f. , it superseded the arrangement of George Bentham, which had stood for over a hundred years. It was overturned in 1996 by Kevin...

 in The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. (Proteaceae)
The genus Banksia L.f. is a 1981 monograph by Alex George on the taxonomy of the plant genus Banksia. Published by the Western Australian Herbarium as Nuytsia 3, it presented George's taxonomic arrangement of Banksia, the first major taxonomic revision of the genus since George Bentham published...

. He demoted Cyrtostylis to series rank, placing it within Banksia subgenus Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

, section Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

. It has remained at this rank since then, so its current name is Banksia ser. Cyrtostylis (Benth.) A.S.George.

George defined Cyrtostylis as those members of B. sect. Banksia with slender flowers, a small pollen-presenter
Pollen-presenter
A pollen-presenter is an area on the tip of the pistil in flowers of plants of the family Proteaceae. In this family, the anthers are difficult of access for potential pollination vectors such as bees, birds and nectariferous mammals....

, and beaked follicles
Follicle (fruit)
In botany, a follicle is a dry unilocular many-seeded fruit formed from one carpel and dehiscing by the ventral suture in order to release seeds, such as in larkspur, magnolia, banksia, peony and milkweed....

. Its type species was given as B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

 (Southern Plains Banksia).

George's 1981 placement and circumscription of B. ser. Cyrtostylis may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. sect. Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (9 species, 5 subspecies)
B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

 (2 species)
B. ser. Quercinae
Banksia ser. Quercinae
Banksia ser. Quercinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (3 species)
B. ser. Orthostylis (8 species, now B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

)
B. ser. Crocinae
Banksia ser. Crocinae
Banksia ser. Crocinae is a taxonomic series in the genus Banksia. It consists of four closely related species, all of which are endemic to Western Australia; namely B. prionotes , B. burdettii , B. hookeriana and B. victoriae...

 (4 species)
B. ser. Cyrtostylis
B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

B. pilostylis
Banksia pilostylis
The Marsh Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia around Esperance. Seeds do not require any treatment, and take 18 to 49 days to germinate.-External links:...

B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

B. lindleyana
Banksia lindleyana
Banksia lindleyana, commonly known as the Porcupine Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as a small shrub to 1 m high with long narrow serrated leaves, and bright yellow oval or round inflorescences...

B. ashbyi
Banksia ashbyi
The Ashby's Banksia is a species of shrub in the plant genus Banksia. It occurs in heath and spinifex country along the coast of Western Australia between Geraldton and Exmouth.-Description:...

B. benthamiana
Banksia benthamiana
Bentham's Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in scattered populations in the Wheatbelt region of Western Australia, one near Mullewa and the other near Dalwallinu.-Description:...

B. audax
Banksia audax
Banksia audax is a species of shrub in the plant genus Banksia. It occurs over a large area in the central south of Western Australia.-Description:...

B. laevigata
Banksia laevigata
Banksia laevigata is a species of shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland from Southern Cross south to the Fitzgerald River National Park. It is composed of two closely related subspecies, B. laevigata subsp. laevigata and B. laevigata...

B. laevigata subsp. laevigata
Banksia laevigata subsp. laevigata
The Tennis Ball Banksia is a subspecies of small woody shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland. It and the closely related B. laevigata subsp...

B. laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea is a subspecies of Banksia laevigata. It is native to the Southwest Botanical Province of Western Australia....

B. lullfitzii
Banksia lullfitzii
Banksia lullfitzii is a species of shrub in the plant genus Banksia. A many-branched, spreading bush with golden-orange flowers, it occurs in scattered populations over a large area of the eastern goldfields of Western Australia...

B. elderana (now spelled B. elderiana
Banksia elderiana
The Swordfish Banksia , commonly known as the swordfish banksia or palm banksia, is a species of shrub in the plant genus Banksia. It occurs in two disjunct areas in the Goldfields-Esperance region of Western Australia...

)
B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

 (6 species, 3 varieties)
B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

 (3 species)
B. ser. Coccineae (1 species)
B. sect. Oncostylis
Banksia sect. Oncostylis
Banksia sect. Oncostylis is one of four sections of subgenus Banksia subg. Banksia. It contains those Banksia species with hooked pistils. All of the species in Oncostylis also exhibit a top-down sequence of flower anthesis, except for Banksia nutans which is bottom-up.Banksia sect...

 (3 series, 21 species, 4 subspecies, 10 varieties)
B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

 (2 species)

A significant addition to the series occurred in 1988, when George published a new species, B. epica
Banksia epica
Banksia epica is a shrub that grows on the south coast of Western Australia. A spreading bush with wedge-shaped serrated leaves and large creamy-yellow flower spikes, it grows up to 3½ metres high. It is known only from two isolated populations in the remote south east of the state, near the...

, placing it between B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

 (Cut-leaf Banksia) and B. media.

According to Thiele and Ladiges

In 1996, Kevin Thiele
Kevin Thiele
Kevin R. Thiele is curator of the Western Australian Herbarium. His research interests include the systematics of the plant families Proteaceae, Rhamnaceae and Violaceae, and the conservation ecology of grassy woodland ecosystems...

 and Pauline Ladiges undertook a cladistic
Cladistics
Cladistics is a method of classifying species of organisms into groups called clades, which consist of an ancestor organism and all its descendants . For example, birds, dinosaurs, crocodiles, and all descendants of their most recent common ancestor form a clade...

 analysis of morphological
Morphology (biology)
In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

 characters of Banksia, which yielded a phylogeny somewhat at odds with George's taxonomic arrangement. They found B. ser. Cyrtostylis to be "widely polyphyletic". Six of the fourteen taxa in George's B. ser. Cyrtostylis occurred singly in locations throughout the cladogram; these were transferred to other series or left incertae sedis
Incertae sedis
, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...

. The remaining eight taxa formed a clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

, which further resolved into two subclades:

Thiele and Ladiges preferred to give series rank to the subclades, rather than the entire clade, so they transferred the taxa of the second clade into B. ser. Ochraceae
Banksia ser. Ochraceae
Banksia ser. Ochraceae is a valid botanic name for a taxonomic series within the plant genus Banksia. It was published by Kevin Thiele in 1996, but discarded by Alex George in 1999.-Cladistics:...

, retaining only the taxa of the first clade in B. ser. Cyrtostylis. No formal definition was given for the new circumscription, although Thiele and Ladiges attached no fewer than seven synapomorphies to the clade, three of which were reversals, the other four parallelisms.

The placement and circumscription of B. ser. Cyrtostylis in Thiele and Ladiges' arrangement of Banksia
Thiele and Ladiges' taxonomic arrangement of Banksia
Kevin Thiele and Pauline Ladiges' taxonomic arrangement of Banksia, published in 1996, was a novel taxonomic arrangement that was intended to align the taxonomy of Banksia more closely with the phylogeny that they had inferred from their cladistic analysis of the genus...

 may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

 (3 species)
B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

 (incertae sedis)
B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

 (4 species)
B. ser. Lindleyanae (1 species)
B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

 (2 subseries, 12 species)
B. baueri
Banksia baueri
The Woolly Banksia is a species of shrub in the plant genus Banksia. It occurs in southwest Western Australia north and east of Albany. It has a distinctively large and hairy looking inflorescence which can be 300 mm or more long and up to 200 mm in diameter.It is placed alone in series...

 (incertae sedis)
B. lullfitzii
Banksia lullfitzii
Banksia lullfitzii is a species of shrub in the plant genus Banksia. A many-branched, spreading bush with golden-orange flowers, it occurs in scattered populations over a large area of the eastern goldfields of Western Australia...

 (incertae sedis)
B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

 (incertae sedis)
B. ashbyi
Banksia ashbyi
The Ashby's Banksia is a species of shrub in the plant genus Banksia. It occurs in heath and spinifex country along the coast of Western Australia between Geraldton and Exmouth.-Description:...

 (incertae sedis)
B. coccinea
Banksia coccinea
Banksia coccinea, commonly known as the Scarlet Banksia, Waratah Banksia or Albany Banksia, is an erect shrub or small tree in the plant genus Banksia...

 (incertae sedis)
B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

 (8 species)
B. ser. Cyrtostylis
B. pilostylis
Banksia pilostylis
The Marsh Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia around Esperance. Seeds do not require any treatment, and take 18 to 49 days to germinate.-External links:...

B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

B. epica
Banksia epica
Banksia epica is a shrub that grows on the south coast of Western Australia. A spreading bush with wedge-shaped serrated leaves and large creamy-yellow flower spikes, it grows up to 3½ metres high. It is known only from two isolated populations in the remote south east of the state, near the...

B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

B. ser. Ochraceae
Banksia ser. Ochraceae
Banksia ser. Ochraceae is a valid botanic name for a taxonomic series within the plant genus Banksia. It was published by Kevin Thiele in 1996, but discarded by Alex George in 1999.-Cladistics:...

 (3 species, 2 subspecies)
B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

 (2 species)
B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (2 series, 11 species, 4 subspecies)
B. ser. Spicigerae
Banksia ser. Spicigerae
Banksia ser. Spicigerae is a taxonomic series in the genus Banksia. It consists of the seven species in section Oncostylis that have cylindrical inflorescences. These range in form from small shrubs to tall trees. The leaves grow in either an alternate or whorled pattern, with various shape forms...

 (3 series, 7 species, 6 varieties)
B. ser. Quercinae
Banksia ser. Quercinae
Banksia ser. Quercinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (2 species)
B. ser. Dryandroideae
Banksia ser. Dryandroideae
Banksia ser. Dryandroideae is a valid botanic name for a taxonomic series in the plant genus Banksia. First published by Carl Meissner in 1856, the name has had two circumscriptions. As presently circumscribed it is monotypic, containing only B. dryandroides.-According to Meissner:B. ser...

 (1 species)
B. ser. Abietinae
Banksia ser. Abietinae
Banksia ser. Abietinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (4 subseries, 15 species, 8 varieties)

According to George 1999

Thiele and Ladiges' arrangement remained current only until 1999, when it was overturned by George. George's 1999 arrangement
George's taxonomic arrangement of Banksia
Alex George's taxonomic arrangement of Banksia was the first modern-day arrangement for that genus. First published in 1981 in the classic monograph The genus Banksia L.f. , it superseded the arrangement of George Bentham, which had stood for over a hundred years. It was overturned in 1996 by Kevin...

 was largely a reversion to his 1981 arrangement. Despite George describing B. ser. Cyrtostylis as "a rather heterogeneous series", its 1981 circumscription was maintained, the only changes being the inclusion of B. epica, the relocation of B. lindleyana to the end of the series, and the relocation of B. laevigata to sit between B. elderiana
Banksia elderiana
The Swordfish Banksia , commonly known as the swordfish banksia or palm banksia, is a species of shrub in the plant genus Banksia. It occurs in two disjunct areas in the Goldfields-Esperance region of Western Australia...

 (Swordfish Banksia) and B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

 (Elegant Banksia). George's 1999 placement and circumscription of B. ser. Cyrtostylis may be summarised as follows:
Banksia
Banksia
Banksia is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes and fruiting "cones" and heads. When it comes to size, banksias range from prostrate woody shrubs to trees up...

B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

B. sect. Banksia
Banksia sect. Banksia
Banksia sect. Banksia is one of four sections of Banksia subgenus Banksia. It contains those species of subgenus Banksia with straight or sometimes curved but not hooked styles. These species all have cylindrical inflorescences and usually exhibit a bottom-up sequence of flower anthesis...

B. ser. Salicinae
Banksia ser. Salicinae
Banksia ser. Salicinae is a valid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (11 species, 7 subspecies)
B. ser. Grandes
Banksia ser. Grandes
Banksia ser. Grandes is a taxonomic series in the genus Banksia. It consists of two closely related species in section Banksia, both endemic to Western Australia. These are B. grandis and B. solandri ....

 (2 species)
B. ser. Banksia
Banksia ser. Banksia
Banksia ser. Banksia is avalid botanic name for a series of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-According to Bentham:Banksia ser. Banksia originated in 1870 as...

 (8 species)
B. ser. Crocinae
Banksia ser. Crocinae
Banksia ser. Crocinae is a taxonomic series in the genus Banksia. It consists of four closely related species, all of which are endemic to Western Australia; namely B. prionotes , B. burdettii , B. hookeriana and B. victoriae...

 (4 species)
B. ser. Prostratae
Banksia ser. Prostratae
Banksia ser. Prostratae is a taxonomic series in the genus Banksia. It consists of six closely related species in section Banksia, all endemic to Western Australia, with a prostrate habit.Banksia ser...

 (6 species, 3 varieties)
B. ser. Cyrtostylis
B. media
Banksia media
The Southern Plains Banksia , also known as Golden Stalk Banksia, is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia between Albany and Israelite Bay, where it is a common plant....

B. praemorsa
Banksia praemorsa
The Cut-leaf Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in a few isolated populations on the south coast of Western Australia between Albany and Cape Riche. Seeds do not require any treatment, and take 30 to 49 days to germinate.-References:...

B. epica
Banksia epica
Banksia epica is a shrub that grows on the south coast of Western Australia. A spreading bush with wedge-shaped serrated leaves and large creamy-yellow flower spikes, it grows up to 3½ metres high. It is known only from two isolated populations in the remote south east of the state, near the...

B. pilostylis
Banksia pilostylis
The Marsh Banksia is a species of shrub in the plant genus Banksia. It occurs on the south coast of Western Australia around Esperance. Seeds do not require any treatment, and take 18 to 49 days to germinate.-External links:...

B. attenuata
Banksia attenuata
Banksia attenuata, commonly known as the candlestick banksia or slender banksia, is a species of plant in the proteaceae family. Commonly a tree, it reaches 10 m high, but is often a shrub in dryer areas 0.4 to 2 m high...

B. ashbyi
Banksia ashbyi
The Ashby's Banksia is a species of shrub in the plant genus Banksia. It occurs in heath and spinifex country along the coast of Western Australia between Geraldton and Exmouth.-Description:...

B. benthamiana
Banksia benthamiana
Bentham's Banksia is a species of shrub or tree in the plant genus Banksia. It occurs in scattered populations in the Wheatbelt region of Western Australia, one near Mullewa and the other near Dalwallinu.-Description:...

B. audax
Banksia audax
Banksia audax is a species of shrub in the plant genus Banksia. It occurs over a large area in the central south of Western Australia.-Description:...

B. lullfitzii
Banksia lullfitzii
Banksia lullfitzii is a species of shrub in the plant genus Banksia. A many-branched, spreading bush with golden-orange flowers, it occurs in scattered populations over a large area of the eastern goldfields of Western Australia...

B. elderiana
Banksia elderiana
The Swordfish Banksia , commonly known as the swordfish banksia or palm banksia, is a species of shrub in the plant genus Banksia. It occurs in two disjunct areas in the Goldfields-Esperance region of Western Australia...

B. laevigata
Banksia laevigata
Banksia laevigata is a species of shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland from Southern Cross south to the Fitzgerald River National Park. It is composed of two closely related subspecies, B. laevigata subsp. laevigata and B. laevigata...

B. laevigata subsp. laevigata
Banksia laevigata subsp. laevigata
The Tennis Ball Banksia is a subspecies of small woody shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland. It and the closely related B. laevigata subsp...

B. laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea
Banksia laevigata subsp. fuscolutea is a subspecies of Banksia laevigata. It is native to the Southwest Botanical Province of Western Australia....

B. elegans
Banksia elegans
The Elegant Banksia is a species of shrub or small tree in the plant genus Banksia. It occurs only over a 65 square kilometre area north and west of Eneabba, Western Australia.-Ecology:...

B. lindleyana
Banksia lindleyana
Banksia lindleyana, commonly known as the Porcupine Banksia, is a species of woody shrub in the genus Banksia of the family Proteaceae. It generally grows as a small shrub to 1 m high with long narrow serrated leaves, and bright yellow oval or round inflorescences...

B. ser. Tetragonae
Banksia ser. Tetragonae
Banksia ser. Tetragonae is a taxonomic series in the genus Banksia. It consists of three closely related species of erect shrub with pendulous inflorescences in section Banksia. These are B. lemanniana , B. caleyi and B. aculeata ....

 (3 species)
B. ser. Bauerinae (1 species)
B. ser. Quercinae
Banksia ser. Quercinae
Banksia ser. Quercinae is avalid botanic name for a series of Banksia. First published by Carl Meissner in 1856, the name has had three circumscriptions.-According to Meissner:...

 (2 species)
B. sect. Coccinea (1 species)
B. sect. Oncostylis
Banksia sect. Oncostylis
Banksia sect. Oncostylis is one of four sections of subgenus Banksia subg. Banksia. It contains those Banksia species with hooked pistils. All of the species in Oncostylis also exhibit a top-down sequence of flower anthesis, except for Banksia nutans which is bottom-up.Banksia sect...

 (4 series, 22 species, 4 subspecies, 11 varieties)
B. subg. Isostylis
Banksia subg. Isostylis
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser...

 (3 species)

In 2002 B. rosserae
Banksia rosserae
Banksia rosserae is a recently described species of Banksia. Endemic to inland Western Australia, it is the only Banksia species to occur solely within the arid zone.-Description:...

 was published; it was tentatively placed in B. ser. Cyrtostylis between B. elderiana
Banksia elderiana
The Swordfish Banksia , commonly known as the swordfish banksia or palm banksia, is a species of shrub in the plant genus Banksia. It occurs in two disjunct areas in the Goldfields-Esperance region of Western Australia...

 and B. laevigata
Banksia laevigata
Banksia laevigata is a species of shrub in the plant genus Banksia. It occurs in Western Australia's semi-arid shrubland from Southern Cross south to the Fitzgerald River National Park. It is composed of two closely related subspecies, B. laevigata subsp. laevigata and B. laevigata...

.

Recent developments

Since 1998, Austin Mast
Austin Mast
Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently an associate professor within the Department of Biological Science at Florida State University , and has been director of FSU's since August 2003.One of his...

 has been publishing results of ongoing cladistic analyses of DNA sequence
DNA sequence
The sequence or primary structure of a nucleic acid is the composition of atoms that make up the nucleic acid and the chemical bonds that bond those atoms. Because nucleic acids, such as DNA and RNA, are unbranched polymers, this specification is equivalent to specifying the sequence of...

 data for the subtribe Banksiinae. His analyses suggest a phylogeny that is rather different to previous taxonomic arrangements. With respect to B. ser. Cyrtostylis, Mast's results accord closely with the arrangement of Thiele and Ladiges, inferring a polytomous clade corresponding exactly with Thiele and Ladiges' series. This clade is however, not particularly close to the clade that corresponds to B. ser. Ochraceae.

Early in 2007 Mast and Thiele initiated a rearrangement of Banksia by transferring Dryandra into it, and publishing B. subg. Spathulatae
Banksia subg. Spathulatae
Banksia subg. Spathulatae is a valid botanic name for a subgenus of Banksia. It was published in 2007 by Austin Mast and Kevin Thiele, and defined as containing all those Banksia species having spathulate cotyledons...

 for the species having spoon-shaped cotyledon
Cotyledon
A cotyledon , is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaves of a seedling. The number of cotyledons present is one characteristic used by botanists to classify the flowering plants...

s; in this way they also redefined the autonym
Autonym (botany)
In botanical nomenclature, autonyms are automatically created names, as regulated by the International Code of Botanical Nomenclature . Autonyms are cited without an author. Relevant provisions are in articles 6.8, 22.1-3 and 26.1-3....

 B. subg. Banksia
Banksia subg. Banksia
Banksia subg. Banksia is a valid botanic name for a subgenus of Banksia. As an autonym, it necessarily contains the type species of Banksia, B. serrata . Within this constraint, however, there have been various circumscriptions.-Banksia verae:B. subg...

. All members of B. ser. Cyrtostylis fall within Mast and Thiele's B. subg. Banksia, but nothing further has been published. Mast and Thiele have foreshadowed publishing a full arrangement once DNA sampling of Dryandra is complete.

External links

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