Bicolor cat
Encyclopedia
A bicolor cat
has white
fur
combined with fur of some other color
, for example black or tabby. There are various patterns of bicolor cat. These range from Van pattern
(color on the crown of the head and the tail only) through to solid color with a throat locket. The tuxedo pattern is one of many possible bicolor patterns.
Where there is low-to-medium grade white spotting limited to the face, paws, throat and chest of an otherwise black cat, they are known in the United States as a tuxedo cat or Billicat. High grade bicolor results in Van-pattern cats. There are many patterns between, such as "cap-and-saddle", "mask-and-mantle" and "magpie" (more randomly splashed). Bicolors are found in many breeds, as well as being common in domestic longhair cats and domestic shorthair cats.
Solid color bicolor cats occur because there is a white spotting gene present along with a recessive allele of the agouti gene
, which evens out the usual striped pattern of the colors of the coat. In contrast, tabby cats have an agouti gene that produces striping of the coat. The Abyssinian
has agouti (ticked tabby) fur, giving the appearance of even color with color-banded hairs.
White spotting can also occur with any of the tabby patterns, resulting in tabby-and-white bicolors. Colorpoint (Himalayan pattern) cats can have bicolor points, although this variation is not recognized for showing. The body markings of bicolor colorpoints become clearer with age, as the body fur of colorpoint cats darkens as the cats grow older and the white patches become more visible.
A low-grade spotting black-and-white bicolor cat is often known as a "tuxedo cat" or a "Billicat". To be considered a tuxedo cat, its black coloring should be solid throughout, with white limited to the paws, belly, chest, throat, face, and possibly the chin: it should appear as if the cat were wearing a tuxedo
.
Another type of black-and-white bicolor cat is nicknamed "cow cat" or "moo cat" (for a perceived resemblance to Holstein cattle
) and includes the magpie, cap-and-saddle and mask-and-mantle patterns. A cow cat does not have the solid black "jacket" of the tuxedo cat. Instead, it has big black patches over a mostly white body, often with a black mask over the head. "Black Mask Cats" are so called because they look like they are wearing a black mask over their head. The Turkish Van
(white and red) is one good example of a bicolor breed. Van pattern is known to animal geneticists as the Seychelles
(Seychellois) Pattern and is classified into 3 variants:
This is high grade white spotting of types 9, 8 and 7 on the bicolor chart above.
, American Shorthair
, Manx
, British Shorthair
, Turkish Angora
, and Bombay
.
In contrast, other common breeds of cat have specific coat patterns specified in their breed standards. These cats are therefore never seen in the bicolor pattern. Cats with such specific coat patterns such as the Russian Blue
which has a coat of one solid color.
to normal coat color as it prevents the migration of melanocyte
s into the developing hair follicles. The genetics of this pattern are not as well understood in cats but at least some of the genes involved in melanocyte migration and survival may play a role similar as in other animals.
Three genotypes possible with the S (white spotting) gene, with capital S standing for a wild-type copy and lower-case s standing for the mutant.
The lack of tabby striping in bicolor cats is controlled by the agouti protein
, which inhibits the production of melanin
and thus prevents the formation of dark hair colors. In agouti cats the gene is turned on and off as the hair grows, producing hairs with alternating stripes yellow and black. In domestic cats, inactivation of the agouti gene by a deletion mutation causes all-black coat color.
and black
coat
. They are called, "tuxedo" cats, because the animal appears to be wearing the type of black tie
formal wear
commonly known in the United States
and Canada
as a tuxedo
. Most tuxedo cats are also black mask cats, a common name for felines who, due to their facial coloration, look as if they are wearing a black
mask
over their eyes, and often over their entire head. To be considered a true tuxedo cat, the feline's coloring should consist of a solid black
coat
, with white
fur
limited to the paws, belly, chest, throat, and often the chin, although many tuxedo cats appear to sport goatee
s, due to the black
coloration of their mandible
-- that is, the lower jaw
and chin
. Bicolor may also appear in the skin color. Paw
pads may be black or pink, often matching the coat in that area; if the color boundary crosses the underside of the paw, the pads on either side may be different colors or even bicolored. Bicoloring of the nose and mouth are also common.
, the tuxedo cat is sometimes known as the "Jellicle cat", after the fictional tribe of black and white cats described by T. S. Eliot
in Old Possum's Book of Practical Cats
, which was first published in 1939. In its derivative musical, Cats
, the tuxedo cat is exemplified by the character of the magical Mr. Mistoffelees
, who is portrayed as a stage magician wearing a lacy ruff
and bow tie
. The musical differed from the book in that the characters included cats with many different coat colors, rather than just bicolor cats, but it retains the repeated assertion that "Jellicle cats are black and white." Cats with these markings also played a starring role in the drawings illustrating The Unadulterated Cat
, a book written by Terry Pratchett
, with cartoons by Joliffe Gray.
Other well known cartoon bicolor cats include Felix the Cat
, Tom Cat
from Tom and Jerry
, Jess from Postman Pat
, Kitty Softpaws from the Shrek
spin-off Puss in Boots
, Figaro
and Sylvester
. A bicolor cat named Mittens is the main deuteragonist in the 2008 Disney animated film "Bolt".
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...
has white
White
White is a color, the perception of which is evoked by light that stimulates all three types of color sensitive cone cells in the human eye in nearly equal amounts and with high brightness compared to the surroundings. A white visual stimulation will be void of hue and grayness.White light can be...
fur
Fur
Fur is a synonym for hair, used more in reference to non-human animals, usually mammals; particularly those with extensives body hair coverage. The term is sometimes used to refer to the body hair of an animal as a complete coat, also known as the "pelage". Fur is also used to refer to animal...
combined with fur of some other color
Color
Color or colour is the visual perceptual property corresponding in humans to the categories called red, green, blue and others. Color derives from the spectrum of light interacting in the eye with the spectral sensitivities of the light receptors...
, for example black or tabby. There are various patterns of bicolor cat. These range from Van pattern
Turkish Van
The Turkish Van is a recognized cat breed that is known for its unusual love of water and swimming. They were created from the cats native to the Lake Van area of Turkey. The cats of this type are named in Turkish Van Kedisi , in Armenian vana katou or vana gadou and in Kurdish...
(color on the crown of the head and the tail only) through to solid color with a throat locket. The tuxedo pattern is one of many possible bicolor patterns.
Where there is low-to-medium grade white spotting limited to the face, paws, throat and chest of an otherwise black cat, they are known in the United States as a tuxedo cat or Billicat. High grade bicolor results in Van-pattern cats. There are many patterns between, such as "cap-and-saddle", "mask-and-mantle" and "magpie" (more randomly splashed). Bicolors are found in many breeds, as well as being common in domestic longhair cats and domestic shorthair cats.
Solid color bicolor cats occur because there is a white spotting gene present along with a recessive allele of the agouti gene
Agouti gene
The Agouti gene is responsible for determining whether a mammal's coat is banded or of a solid color . The chief product of the Agouti gene is Agouti signalling peptide , but there are a number of alternative splice products....
, which evens out the usual striped pattern of the colors of the coat. In contrast, tabby cats have an agouti gene that produces striping of the coat. The Abyssinian
Abyssinian (cat)
The Abyssinian is a breed of domesticated cat with a distinctive ticked coat. There are many stories about its origins, often revolving around Ethiopia, but the actual origins are uncertain...
has agouti (ticked tabby) fur, giving the appearance of even color with color-banded hairs.
White spotting can also occur with any of the tabby patterns, resulting in tabby-and-white bicolors. Colorpoint (Himalayan pattern) cats can have bicolor points, although this variation is not recognized for showing. The body markings of bicolor colorpoints become clearer with age, as the body fur of colorpoint cats darkens as the cats grow older and the white patches become more visible.
Grades 1 - 10 of bicolor in cats
There are 10 identifiable grades of bicolor in cats, plus several patterns with their own names. The cat labelled "bicolor" is the preferred pattern in show-quality bicolor purebred cats.A low-grade spotting black-and-white bicolor cat is often known as a "tuxedo cat" or a "Billicat". To be considered a tuxedo cat, its black coloring should be solid throughout, with white limited to the paws, belly, chest, throat, face, and possibly the chin: it should appear as if the cat were wearing a tuxedo
Black tie
Black tie is a dress code for evening events and social functions. For a man, the main component is a usually black jacket, known as a dinner jacket or tuxedo...
.
Another type of black-and-white bicolor cat is nicknamed "cow cat" or "moo cat" (for a perceived resemblance to Holstein cattle
Holstein (cattle)
Holstein cattle is a breed of cattle known today as the world's highest production dairy animal. Originating in Europe, Holsteins were bred in what is now the Netherlands and more specifically in the two northern provinces of North Holland and Friesland...
) and includes the magpie, cap-and-saddle and mask-and-mantle patterns. A cow cat does not have the solid black "jacket" of the tuxedo cat. Instead, it has big black patches over a mostly white body, often with a black mask over the head. "Black Mask Cats" are so called because they look like they are wearing a black mask over their head. The Turkish Van
Turkish Van
The Turkish Van is a recognized cat breed that is known for its unusual love of water and swimming. They were created from the cats native to the Lake Van area of Turkey. The cats of this type are named in Turkish Van Kedisi , in Armenian vana katou or vana gadou and in Kurdish...
(white and red) is one good example of a bicolor breed. Van pattern is known to animal geneticists as the Seychelles
Seychelles
Seychelles , officially the Republic of Seychelles , is an island country spanning an archipelago of 115 islands in the Indian Ocean, some east of mainland Africa, northeast of the island of Madagascar....
(Seychellois) Pattern and is classified into 3 variants:
- Seychellois Neuvieme - white with colored tail and head splashes (classic Van Pattern)
- Seychellois Huitieme - white with colored tail and head splashes plus additional splashes of color on the legs
- Seychellois Septieme - white with splashes of color on the legs and body in addition to those on the head and the colored tail.
This is high grade white spotting of types 9, 8 and 7 on the bicolor chart above.
Frequency in different breeds
This coloration is not restricted to a specific breed of cat, as it can be found in many different types of pure-breed as well as mixed-breed domestic cats. However, some breeds of cats are especially noted for having bicolor coats in their breed standards. These include the Turkish VanTurkish Van
The Turkish Van is a recognized cat breed that is known for its unusual love of water and swimming. They were created from the cats native to the Lake Van area of Turkey. The cats of this type are named in Turkish Van Kedisi , in Armenian vana katou or vana gadou and in Kurdish...
, American Shorthair
American Shorthair
The American Shorthair is a breed of cat believed to be descended from European domestic cats brought to North America by early settlers to protect valuable cargo from mice and rats...
, Manx
Manx (cat)
The Manx cat , formerly often spelled Manks, is a breed of domestic cat originating on the Isle of Man, with a naturally occurring mutation that shortens the tail...
, British Shorthair
British Shorthair
The British Shorthair is a domesticated cat whose features make it a popular breed in cat shows. It has been the most popular breed of cat registered by the UK's Governing Council of the Cat Fancy since 2001, when it overtook the Persian breed....
, Turkish Angora
Turkish Angora
The Turkish Angora is a breed of domestic cat. Turkish Angoras are one of the ancient, naturally occurring cat breeds, having originated in central Turkey, in the Ankara region...
, and Bombay
Bombay (cat)
The term Bombay cat is used to refer to two different cat breeds, known as the British Bombay and the American Bombay.- General information :* Body: Medium sized, Muscular* Build: Heavy...
.
In contrast, other common breeds of cat have specific coat patterns specified in their breed standards. These cats are therefore never seen in the bicolor pattern. Cats with such specific coat patterns such as the Russian Blue
Russian Blue
The Russian Blue is a cat breed that has a silver-blue coat. These cats are known to be highly intelligent and playful but tend to be timid around strangers...
which has a coat of one solid color.
Genetics of coat patterns
The basic colors and patterns of cat fur are defined by fewer than ten genes. Cats with white color in their coats are thought to have a mutant white-spotting gene that prevents the formation of coat color in patches over the cat's body. This gene has been investigated in several species, particularly mice, and is co-dominantDominance relationship
Dominance in genetics is a relationship between two variant forms of a single gene, in which one allele masks the effect of the other in influencing some trait. In the simplest case, if a gene exists in two allelic forms , three combinations of alleles are possible: AA, AB, and BB...
to normal coat color as it prevents the migration of melanocyte
Melanocyte
-External links: - "Eye: fovea, RPE" - "Integument: pigmented skin"...
s into the developing hair follicles. The genetics of this pattern are not as well understood in cats but at least some of the genes involved in melanocyte migration and survival may play a role similar as in other animals.
Three genotypes possible with the S (white spotting) gene, with capital S standing for a wild-type copy and lower-case s standing for the mutant.
- SS (two dominant alleles) results in high grades of white spotting (sometimes resulting in a solid-looking white cat or a white cat with just a few color hairs)
- Ss (one dominant, one recessive allele) results in medium grades of white spotting
- ss (two recessive alleles) results in solid color or low grades of white spotting (sometimes as little as a few white hairs)
The lack of tabby striping in bicolor cats is controlled by the agouti protein
Agouti signalling peptide
Agouti signalling peptide, a product of the Agouti gene, is a peptide consisting of 131 amino acids. Its discovery was published in 1994 in the scientific journal Nature where its functional properties were described...
, which inhibits the production of melanin
Melanin
Melanin is a pigment that is ubiquitous in nature, being found in most organisms . In animals melanin pigments are derivatives of the amino acid tyrosine. The most common form of biological melanin is eumelanin, a brown-black polymer of dihydroxyindole carboxylic acids, and their reduced forms...
and thus prevents the formation of dark hair colors. In agouti cats the gene is turned on and off as the hair grows, producing hairs with alternating stripes yellow and black. In domestic cats, inactivation of the agouti gene by a deletion mutation causes all-black coat color.
Tuxedo cats
A tuxedo cat is a bicolor cat with a whiteWhite
White is a color, the perception of which is evoked by light that stimulates all three types of color sensitive cone cells in the human eye in nearly equal amounts and with high brightness compared to the surroundings. A white visual stimulation will be void of hue and grayness.White light can be...
and black
Black
Black is the color of objects that do not emit or reflect light in any part of the visible spectrum; they absorb all such frequencies of light...
coat
Coat (animal)
Coat, or the nature and quality of a show mammal's pelage, is important to the animal fancy in the judging of the animal, particularly at conformation dog shows, cat shows and horse shows...
. They are called, "tuxedo" cats, because the animal appears to be wearing the type of black tie
Black tie
Black tie is a dress code for evening events and social functions. For a man, the main component is a usually black jacket, known as a dinner jacket or tuxedo...
formal wear
Formal wear
Formal wear and formal dress are the general terms for clothing suitable for formal social events, such as a wedding, formal garden party or dinner, débutante cotillion, dance, or race...
commonly known in the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...
and Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...
as a tuxedo
Black tie
Black tie is a dress code for evening events and social functions. For a man, the main component is a usually black jacket, known as a dinner jacket or tuxedo...
. Most tuxedo cats are also black mask cats, a common name for felines who, due to their facial coloration, look as if they are wearing a black
Black
Black is the color of objects that do not emit or reflect light in any part of the visible spectrum; they absorb all such frequencies of light...
mask
Mask
A mask is an article normally worn on the face, typically for protection, disguise, performance or entertainment. Masks have been used since antiquity for both ceremonial and practical purposes...
over their eyes, and often over their entire head. To be considered a true tuxedo cat, the feline's coloring should consist of a solid black
Black
Black is the color of objects that do not emit or reflect light in any part of the visible spectrum; they absorb all such frequencies of light...
coat
Coat (animal)
Coat, or the nature and quality of a show mammal's pelage, is important to the animal fancy in the judging of the animal, particularly at conformation dog shows, cat shows and horse shows...
, with white
White
White is a color, the perception of which is evoked by light that stimulates all three types of color sensitive cone cells in the human eye in nearly equal amounts and with high brightness compared to the surroundings. A white visual stimulation will be void of hue and grayness.White light can be...
fur
Fur
Fur is a synonym for hair, used more in reference to non-human animals, usually mammals; particularly those with extensives body hair coverage. The term is sometimes used to refer to the body hair of an animal as a complete coat, also known as the "pelage". Fur is also used to refer to animal...
limited to the paws, belly, chest, throat, and often the chin, although many tuxedo cats appear to sport goatee
Goatee
Goatee refers to a style of facial hair incorporating hair on a man’s chin. The exact nature of the style has varied according to time and culture.Traditionally, goatee refers solely to a beard formed by a tuft of hair on the chin...
s, due to the black
Black
Black is the color of objects that do not emit or reflect light in any part of the visible spectrum; they absorb all such frequencies of light...
coloration of their mandible
Mandible
The mandible pronunciation or inferior maxillary bone forms the lower jaw and holds the lower teeth in place...
-- that is, the lower jaw
Jaw
The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term jaws is also broadly applied to the whole of the structures constituting the vault of the mouth and serving to open and close it and is part of the body plan of...
and chin
Chin
In the human anatomy, the chin is the lowermost part of the face.It is formed by the lower front of the mandible.People show a wide variety of chin structures. See Cleft chin....
. Bicolor may also appear in the skin color. Paw
Paw
A paw is the soft foot of a mammal, generally a quadruped, that has claws or nails. A hard foot is called a hoof. Paws are used to pad feet for walking and increase friction.- Common characteristics :...
pads may be black or pink, often matching the coat in that area; if the color boundary crosses the underside of the paw, the pads on either side may be different colors or even bicolored. Bicoloring of the nose and mouth are also common.
In popular culture
In the United KingdomUnited Kingdom
The United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...
, the tuxedo cat is sometimes known as the "Jellicle cat", after the fictional tribe of black and white cats described by T. S. Eliot
T. S. Eliot
Thomas Stearns "T. S." Eliot OM was a playwright, literary critic, and arguably the most important English-language poet of the 20th century. Although he was born an American he moved to the United Kingdom in 1914 and was naturalised as a British subject in 1927 at age 39.The poem that made his...
in Old Possum's Book of Practical Cats
Old Possum's Book of Practical Cats
Old Possum's Book of Practical Cats is a collection of whimsical poems by T. S. Eliot about feline psychology and sociology, published by Faber and Faber. It is the basis for the record-setting musical Cats....
, which was first published in 1939. In its derivative musical, Cats
Cats (musical)
Cats is a musical composed by Andrew Lloyd Webber, based on Old Possum's Book of Practical Cats by T. S. Eliot...
, the tuxedo cat is exemplified by the character of the magical Mr. Mistoffelees
Mr. Mistoffelees
Mr. Mistoffelees is a character in T. S. Eliot's poetry book Old Possum's Book of Practical Cats and its stage adaptation, Andrew Lloyd Webber's popular musical Cats. He is an important primary character, and one of the most popular of the show...
, who is portrayed as a stage magician wearing a lacy ruff
Ruff (clothing)
A ruff is an item of clothing worn in Western Europe from the mid-sixteenth century to the mid-seventeenth century.The ruff, which was worn by men, women and children, evolved from the small fabric ruffle at the drawstring neck of the shirt or chemise...
and bow tie
Bow tie
The bow tie is a type of men's necktie. It consists of a ribbon of fabric tied around the collar in a symmetrical manner such that the two opposite ends form loops. Ready-tied bow ties are available, in which the distinctive bow is sewn into shape and the band around the neck incorporates a clip....
. The musical differed from the book in that the characters included cats with many different coat colors, rather than just bicolor cats, but it retains the repeated assertion that "Jellicle cats are black and white." Cats with these markings also played a starring role in the drawings illustrating The Unadulterated Cat
The Unadulterated Cat
The Unadulterated Cat by Terry Pratchett, illustrated by Gray Jolliffe, is a book written to promote what Pratchett terms the 'Real Cat', a cat who urinates in the flowerbeds, rips up the furniture, and eats frogs, mice and sundry other small animals...
, a book written by Terry Pratchett
Terry Pratchett
Sir Terence David John "Terry" Pratchett, OBE is an English novelist, known for his frequently comical work in the fantasy genre. He is best known for his popular and long-running Discworld series of comic fantasy novels...
, with cartoons by Joliffe Gray.
Other well known cartoon bicolor cats include Felix the Cat
Felix the Cat
Felix the Cat is a cartoon character created in the silent film era. His black body, white eyes, and giant grin, coupled with the surrealism of the situations in which his cartoons place him, combine to make Felix one of the most recognized cartoon characters in film history...
, Tom Cat
Tom Cat
Thomas "Tom" Cat is a fictional character and half of the academy-award winning Tom and Jerry cartoon cat-and-mouse duo. He is a blue/grey anthropomorphic cat who first appeared in the 1940 animated short Puss Gets the Boot...
from Tom and Jerry
Tom and Jerry
Tom and Jerry are the cat and mouse cartoon characters that were evolved starting in 1939.Tom and Jerry also may refer to:Cartoon works featuring the cat and mouse so named:* The Tom and Jerry Show...
, Jess from Postman Pat
Postman Pat
Postman Pat is a British stop-motion animated children's television series first produced by Woodland Animations. It is aimed at pre-school children, and concerns the adventures of Pat Clifton, a postman in the fictional village of Greendale .Postman Pat's first 13-episode season was screened on...
, Kitty Softpaws from the Shrek
Shrek
Shrek is a 2001 American computer-animated fantasy comedy film directed by Andrew Adamson and Vicky Jenson, featuring the voices of Mike Myers, Eddie Murphy, Cameron Diaz, and John Lithgow. Loosely based on William Steig's 1990 fairy tale picture book Shrek!...
spin-off Puss in Boots
Puss in Boots (2011 film)
Puss in Boots is a 2011 computer-animated adventure Western film produced by DreamWorks Animation, directed by Chris Miller , executive produced by Guillermo del Toro, and written by Brian Lynch, with screenplay by Tom Wheeler. It stars Antonio Banderas, Salma Hayek, Zach Galifianakis, Billy Bob...
, Figaro
Figaro (Disney)
Figaro is a fictional character who first appeared in Disney's Pinocchio.-History:He is probably best known as the pet cat of Mister Geppetto and Pinocchio too. Figaro has also starred independently in a number of Disney shorts, as the pet of Minnie Mouse, which was a common theme for Disney...
and Sylvester
Sylvester (Looney Tunes)
Sylvester J. Pussycat, Sr., Sylvester the Cat or simply Sylvester, is a fictional character, a three-time Academy Award-winning anthropomorphic Tuxedo cat in the Looney Tunes and Merrie Melodies repertory, often chasing Tweety Bird, Speedy Gonzales, or Hippety Hopper...
. A bicolor cat named Mittens is the main deuteragonist in the 2008 Disney animated film "Bolt".
See also
- CatCatThe cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...
- Domestic longhair cat
- Domestic shorthaired catDomestic shorthaired catA domestic shorthaired cat is a cat fancy term used to describe a shorthaired cat that does not have a pedigree or belong to a recognized cat breed...
- FelidaeFelidaeFelidae is the biological family of the cats; a member of this family is called a felid. Felids are the strictest carnivores of the thirteen terrestrial families in the order Carnivora, although the three families of marine mammals comprising the superfamily pinnipedia are as carnivorous as the...
- Fictional cats
- Felix the CatFelix the CatFelix the Cat is a cartoon character created in the silent film era. His black body, white eyes, and giant grin, coupled with the surrealism of the situations in which his cartoons place him, combine to make Felix one of the most recognized cartoon characters in film history...
- Sylvester the Cat
- SocksSocks (cat)Socks was the pet cat of U.S. President Bill Clinton's family during his presidency. An adopted stray cat, he was the only pet of the Clintons during the early years of the administration, and his likeness hosted the children's version of the White House website...
- Tom CatTom CatThomas "Tom" Cat is a fictional character and half of the academy-award winning Tom and Jerry cartoon cat-and-mouse duo. He is a blue/grey anthropomorphic cat who first appeared in the 1940 animated short Puss Gets the Boot...
- Kat KongKat KongKat Kong is a children's picture book created by Dav Pilkey that parodies King Kong with a housecat. Harcourt Brace & Company published this title in 1993...