Clavariadelphus ligula
Encyclopedia
Clavariadelphus ligula, commonly known as the strap coral, is an inedible species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of fungi in the Gomphaceae
Gomphaceae
The Gomphaceae are a diverse family of fungi belonging in what is classically known as the Phallales or cladistically as the gomphoid-phalloid clade. The family has 13 genera and 287 species....

 family. It produces club-shaped fruit bodies
Sporocarp (fungi)
In fungi, the sporocarp is a multicellular structure on which spore-producing structures, such as basidia or asci, are borne...

 with spongy flesh that grow in groups on the forest floor. It is found in Asia, Europe, and North America.

Taxonomy

The species was first described by the German naturalist Jacob Christian Schaeffer in 1774 as Clavaria ligula. It was first placed in Clavariadelphus
Clavariadelphus
Clavariadelphus is a genus of fungi in the Clavariadelphaceae family. The genus has a widespread distribution in temperate areas, and contains 19 species....

by Marinus Anton Donk in 1933. Clavaria ophioglossoides, described by August Batsch
August Batsch
Dr August Johann Georg Karl Batsch was a German naturalist. He was a recognised authority on mushrooms, and also described new species of ferns, bryophytes, and seed plants.- Life and career :...

 in 1783, is considered a synonym.

The fungus is commonly
Common name
A common name of a taxon or organism is a name in general use within a community; it is often contrasted with the scientific name for the same organism...

 known as the "strap coral". The specific epithet ligula is derived from the Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 word for "shoestring".

Description

The club portion of the fruit body is pale yellow, whitish sharp, straight, and stiff hairs at its base. The surface is dull, and does not have hairs. It is smooth at first then later becomes somewhat wrinkled. It is club-shaped to spoon-shaped, and up to 12 cm (4.7 in) tall by 2 cm (0.78740157480315 in) wide at the thickest part. The stipe
Stipe (mycology)
thumb|150px|right|Diagram of a [[basidiomycete]] stipe with an [[annulus |annulus]] and [[volva |volva]]In mycology a stipe refers to the stem or stalk-like feature supporting the cap of a mushroom. Like all tissues of the mushroom other than the hymenium, the stipe is composed of sterile hyphal...

 is not distinct except for the hairs at the base. The flesh
Trama (mycology)
In mycology trama is a term for the inner, fleshy portion of a mushroom's basidiocarp, or fruit body. It is distinct from the outer layer of tissue, known as the pileipellis or cuticle, and from the spore-bearing tissue layer known as the hymenium....

 is whitish, and does not change color with bruising. It is somewhat spongy in the upper part, but firm below. The flesh has no odor, and its taste is slightly bitter. The surface tissue turns green upon the application of a solution of ferric sulphate, and yellow with a dilute solution of potassium hydroxide
Potassium hydroxide
Potassium hydroxide is an inorganic compound with the formula KOH, commonly called caustic potash.Along with sodium hydroxide , this colorless solid is a prototypical strong base. It has many industrial and niche applications. Most applications exploit its reactivity toward acids and its corrosive...

.

The spore
Spore
In biology, a spore is a reproductive structure that is adapted for dispersal and surviving for extended periods of time in unfavorable conditions. Spores form part of the life cycles of many bacteria, plants, algae, fungi and some protozoa. According to scientist Dr...

s are pale yellowish orange ("light buff") in print
Spore print
thumb|300px|right|Making a spore print of the mushroom Volvariella volvacea shown in composite: mushroom cap laid on white and dark paper; cap removed after 24 hours showing pinkish-tan spore print...

. Additional features may be discerned using light microscopy: they are smooth, narrowly ellipsoid, and measure 8–15 by 3–6 μm
Micrometre
A micrometer , is by definition 1×10-6 of a meter .In plain English, it means one-millionth of a meter . Its unit symbol in the International System of Units is μm...

. The hypha
Hypha
A hypha is a long, branching filamentous structure of a fungus, and also of unrelated Actinobacteria. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium; yeasts are unicellular fungi that do not grow as hyphae.-Structure:A hypha consists of one or...

e are monomitic, and clamp connection
Clamp connection
A clamp connection is a structure formed by growing hyphal cells of certain fungi. It is created to ensure each septum, or segment of hypha separated by crossed walls, receives a set of differing nuclei, which are obtained through mating of hyphae of differing sexual types...

s are present. Cystidia
Cystidium
A cystidium is a relatively large cell found on the hymenium of a basidiomycete , often between clusters of basidia. Since cystidia have highly varied and distinct shapes that are often unique to a particular species or genus, they are a useful micromorphological characteristic in the...

 are absent.

Similar species

Clavariadelphus sachalinensis is macroscopically identical from C. ligula, and can be distinguished only by its larger spores, measuring 16–24 by 4–6 μm
Micrometre
A micrometer , is by definition 1×10-6 of a meter .In plain English, it means one-millionth of a meter . Its unit symbol in the International System of Units is μm...

, and longer basidia. However, intermediate forms are often found, and they may represent the same species. Clavaria flavipes is another similar species, but in addition to being smaller and more pale yellow than Clavariadelphus ligula, it also has broader spores.

Habitat and distribution

The fruit bodies of Clavariadelphus ligula grow gregariously (closely scattered over small areas) on the ground, in forest duff
Forest floor
The forest floor, also called detritus, duff and the O horizon, is one of the most distinctive features of a forest ecosystem. It mainly consists of shed vegetative parts, such as leaves, branches, bark, and stems, existing in various stages of decomposition above the soil surface...

. They are widespread and common in coniferous forests, fruiting in the summer and fall.

The species has been collected in the Czech Republic, in the Magadan
Magadan
Magadan is a port town on the Sea of Okhotsk and gateway to the Kolyma region. It is the administrative center of Magadan Oblast , in the Russian Far East. Founded in 1929 on the site of an earlier settlement from the 1920s, it was granted the status of town in 1939...

 region of the Russian Far East
Russian Far East
Russian Far East is a term that refers to the Russian part of the Far East, i.e., extreme east parts of Russia, between Lake Baikal in Eastern Siberia and the Pacific Ocean...

 as well as the arctic zone of the Urals. It has also been collected in the conifer-dominated forests in Kashmir Valley in India. In North America, the distribution extends north to Canada.
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