Coccoloba uvifera
Encyclopedia
Coccoloba uvifera is a species of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

 in the buckwheat
Buckwheat
Buckwheat refers to a variety of plants in the dicot family Polygonaceae: the Eurasian genus Fagopyrum, the North American genus Eriogonum, and the Northern Hemisphere genus Fallopia. Either of the latter two may be referred to as "wild buckwheat"...

 family, Polygonaceae
Polygonaceae
Polygonaceae is a family of flowering plants known informally as the "knotweed family" or "smartweed family"— "buckwheat family" in the United States. The name is based on the genus Polygonum and was first used by Antoine Laurent de Jussieu in 1789 in his book, Genera Plantarum. The name refers...

, that is native to coast
Coast
A coastline or seashore is the area where land meets the sea or ocean. A precise line that can be called a coastline cannot be determined due to the dynamic nature of tides. The term "coastal zone" can be used instead, which is a spatial zone where interaction of the sea and land processes occurs...

al beach
Beach
A beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...

es throughout tropical America
Americas
The Americas, or America , are lands in the Western hemisphere, also known as the New World. In English, the plural form the Americas is often used to refer to the landmasses of North America and South America with their associated islands and regions, while the singular form America is primarily...

 and the Caribbean
Caribbean
The Caribbean is a crescent-shaped group of islands more than 2,000 miles long separating the Gulf of Mexico and the Caribbean Sea, to the west and south, from the Atlantic Ocean, to the east and north...

, including southern Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

, the Bahamas
The Bahamas
The Bahamas , officially the Commonwealth of the Bahamas, is a nation consisting of 29 islands, 661 cays, and 2,387 islets . It is located in the Atlantic Ocean north of Cuba and Hispaniola , northwest of the Turks and Caicos Islands, and southeast of the United States...

, Barbados
Barbados
Barbados is an island country in the Lesser Antilles. It is in length and as much as in width, amounting to . It is situated in the western area of the North Atlantic and 100 kilometres east of the Windward Islands and the Caribbean Sea; therein, it is about east of the islands of Saint...

 and Bermuda
Bermuda
Bermuda is a British overseas territory in the North Atlantic Ocean. Located off the east coast of the United States, its nearest landmass is Cape Hatteras, North Carolina, about to the west-northwest. It is about south of Halifax, Nova Scotia, Canada, and northeast of Miami, Florida...

. Common names include Seagrape and Baygrape.

Description

Coccoloba uvifera is a sprawling evergreen
Evergreen
In botany, an evergreen plant is a plant that has leaves in all seasons. This contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.There are many different kinds of evergreen plants, both trees and shrubs...

 shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

 or small tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 that reaches a maximum height of 8 m (26.2 ft), but most specimens are little more than 2 m (6.6 ft) tall. It has large, round, leathery leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 (up to 25 cm (9.8 in) in diameter) with a primary vein that has a red color extending from the base, and the entire leaf turns red as it ages. The bark is smooth and yellowish.

In late summer it bears green fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

, about 2 cm (0.78740157480315 in) diameter, in large grape
Grape
A grape is a non-climacteric fruit, specifically a berry, that grows on the perennial and deciduous woody vines of the genus Vitis. Grapes can be eaten raw or they can be used for making jam, juice, jelly, vinegar, wine, grape seed extracts, raisins, molasses and grape seed oil. Grapes are also...

-like clusters. The fruit gradually ripen to a purplish color. Each contains a large pit that constitutes most of the volume of the fruit.

Cultivation and propagation

Capable of surviving down to approximately 2°C, the tree is unable to survive frosthttp://plants.usda.gov/java/charProfile?symbol=COUV. The leaves turn reddish before falling. Its seeds have to be planted immediately for they can't withstand being stored for future planting unlike most plants.

Coccoloba uvifera is wind resistant http://books.google.com/books?id=WxW4Scq6kU8C&pg=PA318&lpg=PA318&dq=%22sea+grape%22+%22male%22+%22female%22+%22distinguish%22&source=bl&ots=8eD4FlbdnQ&sig=Wy-HBqMlf4rH7m505E-lz8FeD68&hl=en&ei=sYSqSp-bHcnglAfzls3aBg&sa=X&oi=book_result&ct=result&resnum=1#v=onepage&q=%22sea%20grape%22%20%22male%22%20%22female%22%20%22distinguish%22&f=false, moderately tolerant of shade, and highly tolerant
Halophyte
A halophyte is a plant that grows where it is affected by salinity in the root area or by salt spray, such as in saline semi-deserts, mangrove swamps, marshes and sloughs, and seashores. An example of a halophyte is the salt marsh grass Spartina alterniflora . Relatively few plant species are...

 of salt
Sodium chloride
Sodium chloride, also known as salt, common salt, table salt or halite, is an inorganic compound with the formula NaCl. Sodium chloride is the salt most responsible for the salinity of the ocean and of the extracellular fluid of many multicellular organisms...

, so it is often planted to stabilize beach edges; it is also planted as an ornamental shrub
Ornamental plant
Ornamental plants are plants that are grown for decorative purposes in gardens and landscape design projects, as house plants, for cut flowers and specimen display...

. The fruit is very tasty and can be used for jam or eaten right off the tree.

Coccoloba uvifera is a dioecious
Dioecious
Dioecy is the property of a group of biological organisms that have males and females, but not members that have organs of both sexes at the same time. I.e., those whose individual members can usually produce only one type of gamete; each individual organism is thus distinctly female or male...

 species; that is, male and female flowers are borne on separate plants, and cross-pollination is necessary for fruit to develop. Honey bee
Honey bee
Honey bees are a subset of bees in the genus Apis, primarily distinguished by the production and storage of honey and the construction of perennial, colonial nests out of wax. Honey bees are the only extant members of the tribe Apini, all in the genus Apis...

s and other insects help pollinate these plants http://books.google.com/books?id=BNZV6k6LNMsC&pg=PT127&lpg=PT127&dq=pollinate+sea+grapes&source=bl&ots=qqzv3PufeM&sig=0fX0GxnlV5rG7YDa05SFTClrPhU&hl=en&ei=TnyaSrvDAZzLjAeLramfBQ&sa=X&oi=book_result&ct=result&resnum=8#v=onepage&q=pollinate%20sea%20grapes&f=false; male and female plants can be distinguished by the appearance of their flowers, as males usually show dead flower stalks http://m.naplesnews.com/news/2008/feb/08/delnor-wiggins-pass-state-park-dune-plants-are-tou/.
  • Hardiness: USDA zone 9B - 11
  • Propagation: seeds and cuttings
  • Culture: partial shade/full sun, drought tolerance

Classification

The first botanical names of the plant were assigned in 1696 by Hans Sloane, who called it Prunus
Prunus
Prunus is a genus of trees and shrubs, which includes the plums, cherries, peaches, apricots and almonds. There are around 430 species spread throughout the northern temperate regions of the globe. Many members of the genus are widely cultivated for fruit and ornament.-Botany:Members of the genus...

 maritima racemosa
, "maritime grape-cluster Prunus", and Leonard Plukenet, who named it Uvifera littorea, "grape-bearer of the shore", both of which names reflect the European concept of "sea-grape", expressed in a number of languages by the explorers of the times. The natives viewed it as a large mulberry.

The first edition of Linnaeus's Species Plantarum
Species Plantarum
Species Plantarum was first published in 1753, as a two-volume work by Carl Linnaeus. Its prime importance is perhaps that it is the primary starting point of plant nomenclature as it exists today. This means that the first names to be considered validly published in botany are those that appear...

(1753), based on Plukenet, assigned the plant to Polygonum uvifera and noted flores non vidi, "I have not seen the flowers." Subsequently Patrick Browne, The Civil and Natural History of Jamaica (1756) devised Coccoloba, "red-leaf", for it. Relying on Browne, Linnaeus' second edition (1762), changed the classification to Coccolobus uvifera, citing all the other names.
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