Endemic (ecology)
Encyclopedia
Endemism is the ecological state of being unique to a defined geographic location, such as an island, nation or other defined zone, or habitat
type; organisms that are indigenous
to a place are not endemic to it if they are also found elsewhere. For example, all species of lemur
are endemic to the island of Madagascar
; none are native elsewhere. The extreme opposite of endemism is cosmopolitan distribution
.
Physical, climatic, and biological factors can contribute to endemism. The Orange-breasted Sunbird
is exclusively found in the Fynbos
vegetation zone of southwestern South Africa
. Political factors can play a part if a species is protected, or actively hunted, in one jurisdiction
but not another.
There are two subcategories of endemism - paleoendemism and neoendemism. Paleoendemism refers to a species that was formerly widespread but is now restricted to a smaller area. Neoendemism refers to a species that has recently arisen such as a species that has diverged and become reproductively isolated, or one that has formed following hybridization and is now classified as a separate species. This is a common process in plants, especially those which exhibit polyploidy
.
Endemic types or species are especially likely to develop on biologically isolated areas such as islands because of their geographical isolation. This includes remote island groups, such as Hawaii
, the Galápagos Islands
, and Socotra
, and biologically isolated but not island areas such as the highlands of Ethiopia
, or large bodies of water like Lake Baikal
.
Endemics can easily become endangered or extinct
if their restricted habitat changes, particularly but not only due to the actions of man, including the introduction of new organisms
. There were millions of both Bermuda Petrel
s and "Bermuda cedars
" (actually junipers) in Bermuda
when it was settled at the start of the seventeenth century. By the end of the century the petrels were thought to be extinct. Cedars, already ravaged by centuries of shipbuilding, were driven nearly to extinction in the twentieth century by the introduction of a parasite. Bermuda petrels and cedars, although not actually extinct, are very rare today, as are other species endemic to Bermuda.
Habitat (ecology)
A habitat is an ecological or environmental area that is inhabited by a particular species of animal, plant or other type of organism...
type; organisms that are indigenous
Indigenous (ecology)
In biogeography, a species is defined as native to a given region or ecosystem if its presence in that region is the result of only natural processes, with no human intervention. Every natural organism has its own natural range of distribution in which it is regarded as native...
to a place are not endemic to it if they are also found elsewhere. For example, all species of lemur
Lemur
Lemurs are a clade of strepsirrhine primates endemic to the island of Madagascar. They are named after the lemures of Roman mythology due to the ghostly vocalizations, reflective eyes, and the nocturnal habits of some species...
are endemic to the island of Madagascar
Madagascar
The Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...
; none are native elsewhere. The extreme opposite of endemism is cosmopolitan distribution
Cosmopolitan distribution
In biogeography, a taxon is said to have a cosmopolitan distribution if its range extends across all or most of the world in appropriate habitats. For instance, the killer whale has a cosmopolitan distribution, extending over most of the world's oceans. Other examples include humans, the lichen...
.
Physical, climatic, and biological factors can contribute to endemism. The Orange-breasted Sunbird
Orange-breasted Sunbird
The Orange-breasted Sunbird, Anthobaphes violacea, is the only member of the bird genus Anthobaphes although it is sometimes placed in the genus Nectarinia...
is exclusively found in the Fynbos
Fynbos
Fynbos is the natural shrubland or heathland vegetation occurring in a small belt of the Western Cape of South Africa, mainly in winter rainfall coastal and mountainous areas with a Mediterranean climate...
vegetation zone of southwestern South Africa
South Africa
The Republic of South Africa is a country in southern Africa. Located at the southern tip of Africa, it is divided into nine provinces, with of coastline on the Atlantic and Indian oceans...
. Political factors can play a part if a species is protected, or actively hunted, in one jurisdiction
Jurisdiction
Jurisdiction is the practical authority granted to a formally constituted legal body or to a political leader to deal with and make pronouncements on legal matters and, by implication, to administer justice within a defined area of responsibility...
but not another.
There are two subcategories of endemism - paleoendemism and neoendemism. Paleoendemism refers to a species that was formerly widespread but is now restricted to a smaller area. Neoendemism refers to a species that has recently arisen such as a species that has diverged and become reproductively isolated, or one that has formed following hybridization and is now classified as a separate species. This is a common process in plants, especially those which exhibit polyploidy
Polyploidy
Polyploid is a term used to describe cells and organisms containing more than two paired sets of chromosomes. Most eukaryotic species are diploid, meaning they have two sets of chromosomes — one set inherited from each parent. However polyploidy is found in some organisms and is especially common...
.
Endemic types or species are especially likely to develop on biologically isolated areas such as islands because of their geographical isolation. This includes remote island groups, such as Hawaii
Endemism in the Hawaiian Islands
Located some 2,400 miles from the nearest continental shore, the Hawaiian Islands are the most isolated group of islands on the planet...
, the Galápagos Islands
Galápagos Islands
The Galápagos Islands are an archipelago of volcanic islands distributed around the equator in the Pacific Ocean, west of continental Ecuador, of which they are a part.The Galápagos Islands and its surrounding waters form an Ecuadorian province, a national park, and a...
, and Socotra
Socotra
Socotra , also spelt Soqotra, is a small archipelago of four islands in the Indian Ocean. The largest island, also called Socotra, is about 95% of the landmass of the archipelago. It lies some east of the Horn of Africa and south of the Arabian Peninsula. The island is very isolated and through...
, and biologically isolated but not island areas such as the highlands of Ethiopia
Ethiopia
Ethiopia , officially known as the Federal Democratic Republic of Ethiopia, is a country located in the Horn of Africa. It is the second-most populous nation in Africa, with over 82 million inhabitants, and the tenth-largest by area, occupying 1,100,000 km2...
, or large bodies of water like Lake Baikal
Lake Baikal
Lake Baikal is the world's oldest at 30 million years old and deepest lake with an average depth of 744.4 metres.Located in the south of the Russian region of Siberia, between Irkutsk Oblast to the northwest and the Buryat Republic to the southeast, it is the most voluminous freshwater lake in the...
.
Endemics can easily become endangered or extinct
Extinct species
This page features extinct species, organisms that have become extinct.* List of extinct animals* List of extinct plants...
if their restricted habitat changes, particularly but not only due to the actions of man, including the introduction of new organisms
Introduced species
An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...
. There were millions of both Bermuda Petrel
Bermuda Petrel
The Bermuda Petrel, Pterodroma cahow, is a gadfly petrel. Commonly known in Bermuda as the Cahow, a name derived from its eerie cries, this nocturnal ground-nesting seabird is the national bird of Bermuda, and a symbol of hope for nature conservation. It was thought extinct for 330 years...
s and "Bermuda cedars
Juniperus bermudiana
Juniperus bermudiana is a species of juniper endemic to Bermuda. This species is most commonly known as Bermuda cedar although, like most "cedars" it is not a true cedar ; a more botanically accurate name would be Bermuda juniper, but this term is extremely rare.It is an evergreen tree growing up...
" (actually junipers) in Bermuda
Bermuda
Bermuda is a British overseas territory in the North Atlantic Ocean. Located off the east coast of the United States, its nearest landmass is Cape Hatteras, North Carolina, about to the west-northwest. It is about south of Halifax, Nova Scotia, Canada, and northeast of Miami, Florida...
when it was settled at the start of the seventeenth century. By the end of the century the petrels were thought to be extinct. Cedars, already ravaged by centuries of shipbuilding, were driven nearly to extinction in the twentieth century by the introduction of a parasite. Bermuda petrels and cedars, although not actually extinct, are very rare today, as are other species endemic to Bermuda.
Ecoregions with high endemism
According to the World Wildlife Fund, the following ecoregions have the highest percentage of endemic plants:- FynbosFynbosFynbos is the natural shrubland or heathland vegetation occurring in a small belt of the Western Cape of South Africa, mainly in winter rainfall coastal and mountainous areas with a Mediterranean climate...
(South AfricaSouth AfricaThe Republic of South Africa is a country in southern Africa. Located at the southern tip of Africa, it is divided into nine provinces, with of coastline on the Atlantic and Indian oceans...
) - Hawaiian tropical dry forestsHawaiian tropical dry forestsThe Hawaiian tropical dry forests are a tropical dry broadleaf forest ecoregion in the Hawaiian Islands. They cover an area of on the leeward side of the main islands and the summits of Niihau and Kahoolawe. These forests are either seasonal or sclerophyllous. Annual rainfall is less than and...
(United StatesUnited StatesThe United States of America is a federal constitutional republic comprising fifty states and a federal district...
) - Hawaiian tropical rainforests (United StatesUnited StatesThe United States of America is a federal constitutional republic comprising fifty states and a federal district...
) - Kwongan heathlandsKwonganKwongan is a type of heathland found on the coastal plains of Western Australia. The name is derived from the language of the Noongar people. Kwongan comprises floristically-rich heath with dense thickets of sclerophyllous shrubs and isolated small trees...
(AustraliaAustraliaAustralia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...
) - Madagascar dry deciduous forestsMadagascar dry deciduous forestsThe Madagascar dry deciduous forests represent a tropical dry forest ecoregion generally situated in the western part of Madagascar. The area has high numbers of endemic plant and animal species but has suffered large-scale clearance for agriculture...
(MadagascarMadagascarThe Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...
) - Madagascar lowland forestsMadagascar lowland forestsThe Madagascar lowland forests are a tropical moist broadleaf forest ecoregion, found on the eastern coast of the island of Madagascar.-Setting:...
(MadagascarMadagascarThe Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...
) - New Caledonia dry forests (New CaledoniaNew CaledoniaNew Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...
) - New Caledonia rain forestsNew Caledonia rain forestsThe New Caledonia rain forests are a terrestrial ecoregion, located in New Caledonia in the South Pacific. It is a tropical moist broadleaf forest ecoregion, part of the Australasia ecozone.-Setting:...
(New CaledoniaNew CaledoniaNew Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...
) - Sierra Madre de Oaxaca pine-oak forestsSierra Madre de Oaxaca pine-oak forestsThe Sierra Madre de Oaxaca pine-oak forests is an ecoregion of southern Mexico, in the Tropical and subtropical coniferous forests biome. It occupies the Sierra Madre de Oaxaca, which lies mostly within the state of Oaxaca, extending north into Puebla and Veracruz states...
(MexicoMexicoThe United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...
) - Sierra Madre del Sur pine-oak forestsSierra Madre del Sur pine-oak forestsThe Sierra Madre del Sur pine-oak forests is a subtropical coniferous forest ecoregion of the Sierra Madre del Sur mountain range of southern Mexico.-Setting:...
(GuatemalaGuatemalaGuatemala is a country in Central America bordered by Mexico to the north and west, the Pacific Ocean to the southwest, Belize to the northeast, the Caribbean to the east, and Honduras and El Salvador to the southeast...
) - LuzonLuzonLuzon is the largest island in the Philippines. It is located in the northernmost region of the archipelago, and is also the name for one of the three primary island groups in the country centered on the Island of Luzon...
montaneMontaneIn biogeography, montane is the highland area located below the subalpine zone. Montane regions generally have cooler temperatures and often have higher rainfall than the adjacent lowland regions, and are frequently home to distinct communities of plants and animals.The term "montane" means "of the...
rainforests (PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
) - Luzon rainforests (PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
) - Luzon tropical pine forestsLuzon tropical pine forestsThe Luzon tropical pine forests are a tropical coniferous forest ecoregion of the Philippines in the western Pacific Ocean. These pine forests are home to a large number of the island's endemic plants and animals.-Location and description:...
(PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
) - Mindanao montane rain forests (PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
) - Mindanao-Eastern Visayas rain forests (PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
) - Palawan rain forests (PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
)
Threats to highly endemistic regions
Some of the principal threats to these special ecosystems are:- Large scale loggingLoggingLogging is the cutting, skidding, on-site processing, and loading of trees or logs onto trucks.In forestry, the term logging is sometimes used in a narrow sense concerning the logistics of moving wood from the stump to somewhere outside the forest, usually a sawmill or a lumber yard...
operations - Slash-and-burn techniques sometimes a part of shifting cultivationShifting cultivationShifting cultivation is an agricultural system in which plots of land are cultivated temporarily, then abandoned. This system often involves clearing of a piece of land followed by several years of wood harvesting or farming, until the soil loses fertility...
- Destruction of habital or vegetation leads to endangering of the endemic species