Gecarcinus quadratus, known as the red land crab, whitespot crab, Halloween crab, moon crab, mouthless crab or harlequin land crab, is a colourful land crab

Land crab

Gecarcinidae is a family of true crabs that are adapted for terrestrial existence, commonly known as land crabs. Similar to all other crabs, land crabs possess a series of gills. In addition, the part of the carapace covering the gills is inflated and equipped with blood vessels. These organs...

 from the family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Gecarcinidae.

Distribution

G. quadratus is found in mangrove

Mangrove

Mangroves are various kinds of trees up to medium height and shrubs that grow in saline coastal sediment habitats in the tropics and subtropics – mainly between latitudes N and S...

, sand dunes and rainforest along the Pacific coast from Mexico

Mexico

The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 south to Peru

Peru

Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

, and around the Gulf of Mexico

Gulf of Mexico

The Gulf of Mexico is a partially landlocked ocean basin largely surrounded by the North American continent and the island of Cuba. It is bounded on the northeast, north and northwest by the Gulf Coast of the United States, on the southwest and south by Mexico, and on the southeast by Cuba. In...

. The taxonomy

Taxonomy

Taxonomy is the science of identifying and naming species, and arranging them into a classification. The field of taxonomy, sometimes referred to as "biological taxonomy", revolves around the description and use of taxonomic units, known as taxa...

 in relation to the Atlantic Gecarcinus lateralis

Gecarcinus lateralis

Gecarcinus lateralis, also known by the common names blackback land crab, Bermuda land crab, red land crab and moon crab Gecarcinus lateralis, also known by the common names blackback land crab, Bermuda land crab, red land crab (leading to easy confusion with Gecarcoidea natalis) and moon crab...

is disputed, with many considering it and G. quadratus to be conspecific.

Description and behaviour

The carapace of G. quadratus may reach a length of 5 centimetres (2 in). It has a pair of largely purple claw

Claw

A claw is a curved, pointed appendage, found at the end of a toe or finger in most mammals, birds, and some reptiles. However, the word "claw" is also often used in reference to an invertebrate. Somewhat similar fine hooked structures are found in arthropods such as beetles and spiders, at the end...

s, reddish-orange legs, and an almost entirely blackish carapace

Carapace

A carapace is a dorsal section of the exoskeleton or shell in a number of animal groups, including arthropods such as crustaceans and arachnids, as well as vertebrates such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron.-Crustaceans:In crustaceans, the...

 with a pair of yellow, orange or maroon spots behind the eyes, and an additional pair of whitish spots on the central-lower carapace.

This nocturnal crab digs burrow

Burrow

A burrow is a hole or tunnel dug into the ground by an animal to create a space suitable for habitation, temporary refuge, or as a byproduct of locomotion. Burrows provide a form of shelter against predation and exposure to the elements, so the burrowing way of life is quite popular among the...

s – sometimes as long as 1.5 metres (4.9 ft) – in the coastal rainforest

Rainforest

Rainforests are forests characterized by high rainfall, with definitions based on a minimum normal annual rainfall of 1750-2000 mm...

s of Central America

Central America

Central America is the central geographic region of the Americas. It is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. When considered part of the unified continental model, it is considered a subcontinent...

, and is common along the coasts of Costa Rica

Costa Rica

Costa Rica , officially the Republic of Costa Rica is a multilingual, multiethnic and multicultural country in Central America, bordered by Nicaragua to the north, Panama to the southeast, the Pacific Ocean to the west and the Caribbean Sea to the east....

, Panama

Panama

Panama , officially the Republic of Panama , is the southernmost country of Central America. Situated on the isthmus connecting North and South America, it is bordered by Costa Rica to the northwest, Colombia to the southeast, the Caribbean Sea to the north and the Pacific Ocean to the south. The...

, and Nicaragua

Nicaragua

Nicaragua is the largest country in the Central American American isthmus, bordered by Honduras to the north and Costa Rica to the south. The country is situated between 11 and 14 degrees north of the Equator in the Northern Hemisphere, which places it entirely within the tropics. The Pacific Ocean...

. It lives in the forest at least some of its adult life, but needs to return to the ocean to breed. It is largely herbivorous

Herbivore

Herbivores are organisms that are anatomically and physiologically adapted to eat plant-based foods. Herbivory is a form of consumption in which an organism principally eats autotrophs such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in...

, and consumes leaf litter and seedling

Seedling

thumb|Monocot and dicot seedlingsA seedling is a young plant sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle , the hypocotyl , and the cotyledons...

s.