Juglandaceae
Encyclopedia
The Juglandaceae, also known as the Walnut Family, is a family of tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

s, or sometimes shrubs, in the order Fagales
Fagales
The Fagales are an order of flowering plants, including some of the best known trees. The order name is derived from genus Fagus, Beeches. They belong among the rosid group of dicotyledons...

. Various members of this family are native to the Americas
Americas
The Americas, or America , are lands in the Western hemisphere, also known as the New World. In English, the plural form the Americas is often used to refer to the landmasses of North America and South America with their associated islands and regions, while the singular form America is primarily...

, Eurasia
Eurasia
Eurasia is a continent or supercontinent comprising the traditional continents of Europe and Asia ; covering about 52,990,000 km2 or about 10.6% of the Earth's surface located primarily in the eastern and northern hemispheres...

, and Southeast Asia
Southeast Asia
Southeast Asia, South-East Asia, South East Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India, west of New Guinea and north of Australia. The region lies on the intersection of geological plates, with heavy seismic...

.
Members of the walnut family have large aromatic leaves, that are usually alternate, but opposite in Alfaroa
Alfaroa
Alfaroa is a genus of evergreen trees in the Juglandaceae family of the Fagales, growing in montaine and submontaine tropical rain forests in Central America. The wood is characterized by solid pith, pink heartwood, and vessels with scalariform perforations, as well as simple...

and Oreomunnia. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are pinnately compound, or ternate, and usually 20–100 cm long.

The trees are wind-pollinated
Anemophily
Anemophily or wind pollination is a form of pollination whereby pollen is distributed by wind. Anemophilous plants may be either gymnosperms or angiosperms ....

, and the flowers usually arranged in catkin
Catkin
A catkin or ament is a slim, cylindrical flower cluster, with inconspicuous or no petals, usually wind-pollinated but sometimes insect pollinated . They contain many, usually unisexual flowers, arranged closely along a central stem which is often drooping...

s.


There are eight genera in the family, including the commercially important nut
Nut (fruit)
A nut is a hard-shelled fruit of some plants having an indehiscent seed. While a wide variety of dried seeds and fruits are called nuts in English, only a certain number of them are considered by biologists to be true nuts...

-producing trees walnut
Walnut
Juglans is a plant genus of the family Juglandaceae, the seeds of which are known as walnuts. They are deciduous trees, 10–40 meters tall , with pinnate leaves 200–900 millimetres long , with 5–25 leaflets; the shoots have chambered pith, a character shared with the wingnuts , but not the hickories...

 (Juglans), pecan
Pecan
The pecan , Carya illinoinensis, is a species of hickory, native to south-central North America, in Mexico from Coahuila south to Jalisco and Veracruz, in the United States from southern Iowa, Illinois, Missouri, and Indiana east to western Kentucky, southwestern Ohio, North Carolina, South...

 (Carya illinoinensis), and hickory
Hickory
Trees in the genus Carya are commonly known as hickory, derived from the Powhatan language of Virginia. The genus includes 17–19 species of deciduous trees with pinnately compound leaves and big nuts...

 (Carya). The Persian walnut, Juglans regia, is one of the major nut crops of the world. Walnut, hickory, and gaulin are also valuable timber trees.

Systematics

The known living genera are grouped into subfamilies, tribes, and subtribes as follows:

Subfamily Engelhardioideae
  • Alfaroa
    Alfaroa
    Alfaroa is a genus of evergreen trees in the Juglandaceae family of the Fagales, growing in montaine and submontaine tropical rain forests in Central America. The wood is characterized by solid pith, pink heartwood, and vessels with scalariform perforations, as well as simple...

    Standl. – gaulin
  • Engelhardia
    Engelhardia
    Engelhardia is a genus of seven species of trees in the family Juglandaceae, native to southeast Asia from northern India east to Taiwan, Indonesia and the Philippines...

    Lesch. ex Blume – cheo
  • Oreomunnea
    Oreomunnea
    Oreomunnea is a genus of two species of flowering plants in the family Juglandaceae, native to southern Mexico and Central America, where they occur in montane rainforest....

    Oerst.

Subfamily Juglandoideae
Juglandoideae
Jugandoideae is the walnut subfamily....



 Tribe Platycaryeae
  • Platycarya
    Platycarya
    Platycarya is a genus of flowering plants in the family Juglandaceae, usually treated as comprising a single species Platycarya strobilacea, though one to two additional species are accepted by some authors. It is native to eastern Asia in China, Korea, and Japan.It is a deciduous tree growing to...

    Siebold & Zucc.

 Tribe Juglandeae

  Subtribe Caryinae
Caryinae
Caryinae is a subtribe of the tribe Juglandeae subfamily in the Juglandaceae.The Latin description of this subtribe is as follows:Medulla solida; pollenis grana 3-pora; bracteolae pis- tillatae 2 , ad apicem ovarii connatae; positio carpellorum transversa ; stigmata com- missuralia; papillae...

  • Carya
    Hickory
    Trees in the genus Carya are commonly known as hickory, derived from the Powhatan language of Virginia. The genus includes 17–19 species of deciduous trees with pinnately compound leaves and big nuts...

    Nutt. – hickory and pecan
  • Annamocarya
    Annamocarya
    Annamocarya is a genus of flowering plants in the family Juglandaceae, containing only one species, Annamocarya sinensis, native to southwestern China and northern Vietnam...

    A.Chev.

  Subtribe Juglandinae
  • Cyclocarya
    Cyclocarya
    Cyclocarya is a genus of flowering plants in the family Juglandaceae, comprising a single species Cyclocarya paliurus , formerly treated in the genus Pterocarya as Pterocarya paliurus. It is native to eastern and central China.It is a deciduous tree growing to 30 m tall...

    Iljinsk – wheel wingnut
  • Juglans L. – walnut
  • Pterocarya Kunth – wingnut

Incertae sedis
  • Rhoiptelea Diels & Hand.-Mazz.

The only member of the genus Alfaropsis I.A.Iljinsk., Alfaropsis roxburghiana (Wall.) I.A.Iljinsk. is a synonym for Engelhardia roxburghiana Wall. (or perhaps vice-versa).

The only member of the genus Annamocarya
Annamocarya
Annamocarya is a genus of flowering plants in the family Juglandaceae, containing only one species, Annamocarya sinensis, native to southwestern China and northern Vietnam...

A.Chev., Annamocarya sinensis (Dode) J.-F.Leroy, may actually be a member of Carya.

Tryma

Some fruits are borderline and difficult to categorize. Hickory nuts (Carya) and Walnuts (Juglans) grow within an outer husk; these fruits are technically drupe
Drupe
In botany, a drupe is a fruit in which an outer fleshy part surrounds a shell of hardened endocarp with a seed inside. These fruits develop from a single carpel, and mostly from flowers with superior ovaries...

s or drupaceous nuts, and thus not true botanical nuts. Tryma is a specialized term for such nut-like drupes.
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